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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1128.89751" ID-Pensoft-Pub="1313-2970-1128-129" ID-Pensoft-UUID="9462C7DFB85D5DC780F645BC75150827" ID-ZooBank="7AC1EB83EE914109892AE5714A65EFB9" ModsDocID="1313-2970-1128-129" checkinTime="1668024534632" checkinUser="pensoft" docAuthor="Saxton, Natalie A., Marinov, Milen G. &amp; Bybee, Seth M." docDate="2022" docId="D8738B6ACDCB5BC9AC8E9FE4849ED810" docLanguage="en" docName="ZooKeys 1128: 129-169" docOrigin="ZooKeys 1128" docPubDate="2022-11-09" docSource="http://dx.doi.org/10.3897/zookeys.1128.89751" docTitle="Vanuatubasis santoensis Ober &amp; Staniczek 2009" docType="treatment" docVersion="1" id="9462C7DFB85D5DC780F645BC75150827" lastPageNumber="129" masterDocId="9462C7DFB85D5DC780F645BC75150827" masterDocTitle="Revision of Vanuatubasis Ober &amp; Staniczek, 2009 (Odonata, Coenagrionidae), with description of seven new species" masterLastPageNumber="169" masterPageNumber="129" pageNumber="129" updateTime="1668024534632" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Vanuatubasis Ober &amp; Staniczek, 2009 (Odonata, Coenagrionidae), with description of seven new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Saxton, Natalie A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5993-9782</mods:nameIdentifier>
<mods:affiliation>Research and Collections Division, The Cleveland Museum of Natural History, Cleveland, OH, 44106, USA &amp; Department of Biology, Case Western Reserve University, Cleveland, OH, 44106, USA &amp; Department of Biology and Monte L. Bean Museum Brigham Young University, Provo, UT 84604, USA</mods:affiliation>
<mods:nameIdentifier type="email">nsaxton55@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marinov, Milen G.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3284-2555</mods:nameIdentifier>
<mods:affiliation>Biosecurity Surveillance and Incursion Investigation Plant Health Team, Ministry for Primary Industries, 14 Sir William Pickering Drive, Christchurch 8053, New Zealand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bybee, Seth M.</mods:namePart>
<mods:affiliation>Department of Biology and Monte L. Bean Museum Brigham Young University, Provo, UT 84604, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1128</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>129</mods:start>
<mods:end>169</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1128.89751</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1128.89751</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1128-129</mods:identifier>
<mods:identifier type="ZooBank">7AC1EB83EE914109892AE5714A65EFB9</mods:identifier>
<mods:identifier type="Pensoft-UUID">9462C7DFB85D5DC780F645BC75150827</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:D8738B6ACDCB5BC9AC8E9FE4849ED810" httpUri="http://treatment.plazi.org/id/D8738B6ACDCB5BC9AC8E9FE4849ED810" lastPageNumber="129" pageId="0" pageNumber="129">
<subSubSection pageId="0" pageNumber="129" type="nomenclature">
<paragraph pageId="0" pageNumber="129">
<taxonomicName LSID="D8738B6A-CDCB-5BC9-AC8E-9FE4849ED810" authority="Ober &amp; Staniczek, 2009" authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">Vanuatubasis santoensis Ober &amp; Staniczek, 2009</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="description">
<paragraph pageId="0" pageNumber="129">
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Vanuatubasis santoensis (♂ BYU) A dorsal thorax B lateral terminalia C dorsal terminalia D lateral habitus V. santoensis (♀ BYU) E dorsal thorax F lateral terminalia. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1128.89751.figure10" httpUri="https://binary.pensoft.net/fig/767336" pageId="0" pageNumber="129">Figs 10</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male dorso-posterior and dorsal terminalia of A V. bidens B V. evelynae C V. insularivorum D V. kapularum E V. malekulana F V. nunggoli G V. santoensis." figureDoi="10.3897/zookeys.1128.89751.figure20" httpUri="https://binary.pensoft.net/fig/767346" pageId="0" pageNumber="129">, 20G</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female face and dorsal terminalia of A V. bidens B V. discontinua C V. evelynae D V. insularivorum E V. kapularum F V. malekulana G V. nunggoli H V. rhomboides I V. santoensis J V. xanthochroa." figureDoi="10.3897/zookeys.1128.89751.figure21" httpUri="https://binary.pensoft.net/fig/767347" pageId="0" pageNumber="129">, 21I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="reference_group">
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">Vanuatubasis santoensis</taxonomicName>
Ober &amp; Staniczek (2009): 487-490.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="remarks">
<paragraph pageId="0" pageNumber="129">Remark.</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">
<emphasis italics="true" pageId="0" pageNumber="129">Vanuatubasis santoensis</emphasis>
</taxonomicName>
was known by males only. Here we describe the female.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="materials_examined">
<paragraph pageId="0" pageNumber="129">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">
<typeStatus>Holotype</typeStatus>
</emphasis>
(
<specimenCount type="male">1 ♂</specimenCount>
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">SMNS).</emphasis>
&quot;Vanuatubasis santoensis
<specimenCount>♂</specimenCount>
<materialsCitation collectingDate="2006-11-13" collectingDateMax="2018-06-06" collectingDateMin="2006-11-13" collectorName="A. H. Staniczek, M. Pallmann, S. Bybee, G. Powell" country="VANUATU" county="Penaoru River" determinerName="S. V. Ober" elevation="90" latitude="-14.96105" location="Espiritu Santo" longLatPrecision="1" longitude="166.63316" municipality="Penaoru" specimenCount="28" specimenCount-female="9" specimenCount-male="19" stateProvince="Sanma Province" typeStatus="HOLOTYPE">
<typeStatus>HOLOTYPE</typeStatus>
|
<bibRefCitation DOI="https://doi.org/10.5252/z2009n3a6" author="Ober, S" journalOrPublisher="Zoosystema" pageId="0" pageNumber="129" pagination="485 - 497" refId="B9" refString="Ober, S, Staniczek, A, 2009. A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485 - 497, DOI: https://doi.org/10.5252/z2009n3a6" title="A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu." url="https://doi.org/10.5252/z2009n3a6" volume="31" year="2009">Ober and Staniczek 2009</bibRefCitation>
| Vanuatu,
<collectingRegion country="Vanuatu" name="Sanma">Sanma Province</collectingRegion>
|
<location LSID="urn:lsid:plazi:treatment:D8738B6ACDCB5BC9AC8E9FE4849ED810:A4A51B84D109514573F29CF1772FE0A2" country="VANUATU" county="Penaoru River" latitude="-14.96105" longLatPrecision="1" longitude="166.63316" municipality="Penaoru" name="Espiritu Santo" stateProvince="Sanma Province">Espiritu Santo</location>
, surroundings of |
<collectingMunicipality>Penaoru</collectingMunicipality>
,
<collectingCounty>Penaoru River</collectingCounty>
|leg.
<collectorName>A.H. Staniczek</collectorName>
and
<collectorName>M. Pallmann</collectorName>
,
<collectingDate value="2006-11-13">13.XI.2006</collectingDate>
| det.
<determinerName>S.V. Ober</determinerName>
,
<collectingDate value="2008-07-21">21.VII.2008</collectingDate>
| coll.- |
<bibRefCitation DOI="https://doi.org/10.5252/z2009n3a6" author="Ober, S" journalOrPublisher="Zoosystema" pageId="0" pageNumber="129" pagination="485 - 497" refId="B9" refString="Ober, S, Staniczek, A, 2009. A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485 - 497, DOI: https://doi.org/10.5252/z2009n3a6" title="A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu." url="https://doi.org/10.5252/z2009n3a6" volume="31" year="2009">Ober and Staniczek 2009</bibRefCitation>
|
<geoCoordinate degrees="14.96105" direction="south" orientation="latitude" precision="1" value="-14.96105">14.96105°S</geoCoordinate>
,
<geoCoordinate degrees="166.63316" direction="east" orientation="longitude" precision="1" value="166.63316">166.63316°E</geoCoordinate>
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">90 m</elevation>
</quantity>
| ODO 000242 K.&quot;
<emphasis bold="true" pageId="0" pageNumber="129">
(
<specimenCount type="male">4 ♂♂</specimenCount>
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BYU)</emphasis>
&quot;
<collectingCountry name="Vanuatu">VANUATU</collectingCountry>
: Santo Is: | Wailapa, -15.5781 | 167.0024,
<collectingDate value="2018-06-06">6.vi.2018</collectingDate>
, coll, |
<collectorName>S. Bybee</collectorName>
and
<collectorName>G. Powell</collectorName>
&quot;
<emphasis bold="true" pageId="0" pageNumber="129">
(
<specimenCount type="male">1 ♂</specimenCount>
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BYU)</emphasis>
&quot;
<collectingCountry name="Vanuatu">VANUATU</collectingCountry>
: Santo Is: | Narango, -15.6274 | 166.8535,
<collectingDate value="2018-06-04">4.vi.2018</collectingDate>
, | coll,
<collectorName>S. Bybee</collectorName>
and
<collectorName>G. Powell</collectorName>
&quot;
<emphasis bold="true" pageId="0" pageNumber="129">
(
<specimenCount type="male">2 ♂♂</specimenCount>
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BYU)</emphasis>
&quot;
<collectingCountry name="Vanuatu">VANUATU</collectingCountry>
: Santo Is: | Narango, -15.5538 | 166.9814,
<collectingDate value="2018-06-04">4.vi.2018</collectingDate>
, | coll,
<collectorName>S. Bybee</collectorName>
and
<collectorName>G. Powell</collectorName>
&quot;
<emphasis bold="true" pageId="0" pageNumber="129">
(
<specimenCount type="male">12 ♂♂</specimenCount>
</emphasis>
<specimenCount type="female">
<emphasis bold="true" pageId="0" pageNumber="129">4</emphasis>
♀♀
</specimenCount>
<emphasis bold="true" pageId="0" pageNumber="129">BYU)</emphasis>
&quot;
<collectingCountry name="Vanuatu">VANUATU</collectingCountry>
: Santo Is: | Ipayato, -15.6296| 166.8426,
<collectingDate value="2018-06-04">4 vi.2018</collectingDate>
. | coll.
<collectorName>S. Bybee</collectorName>
and
<collectorName>G. Powell</collectorName>
&quot;
<emphasis bold="true" pageId="0" pageNumber="129">
(
<specimenCount type="female">5 ♀♀</specimenCount>
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BYU)</emphasis>
&quot;
<collectingCountry name="Vanuatu">VANUATU</collectingCountry>
: Santo Is: Felea |
<geoCoordinate degrees="15.3839" direction="south" orientation="latitude" precision="5" value="-15.3839">-15.3839</geoCoordinate>
<geoCoordinate degrees="166.8426" direction="east" orientation="longitude" precision="5" value="166.8426">166.8426</geoCoordinate>
,
<collectingDate value="2018-06-04">4.vi.2018</collectingDate>
, | coll,
<collectorName>S. Bybee</collectorName>
and
<collectorName>G. Powell</collectorName>
&quot;
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="129">
<materialsCitation collectingDate="2006-11-09" collectingDateMax="2006-11-11" collectingDateMin="2006-11-09" collectorName="A. H. Staniczek, M. Pallmann" country="Vanuatu" county="Espiritu Santo" determinerName="S. V. Ober" elevation="139" latitude="-15.2175" location="Pae River" longLatPrecision="5" longitude="166.68706" municipality="Tasmate" specimenCount="7" specimenCount-male="7" stateProvince="Sanma Province" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">
<typeStatus>Paratypes</typeStatus>
</emphasis>
(
<specimenCount type="male">2 ♂♂</specimenCount>
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">SMNS).</emphasis>
same label data as Holotype with the following barcode numbers: ODO 000246 K, ODO 000247 K.
<emphasis bold="true" pageId="0" pageNumber="129">
(
<specimenCount type="male">1 ♂</specimenCount>
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">SMNS).</emphasis>
&quot;
<collectingCountry name="Vanuatu">Vanuatu</collectingCountry>
,
<collectingRegion country="Vanuatu" name="Sanma">Sanma Province</collectingRegion>
|
<collectingCounty>Espiritu Santo</collectingCounty>
, surroundings of
<collectingMunicipality>Tasmate</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:D8738B6ACDCB5BC9AC8E9FE4849ED810:833467CD654822EFF1A36CD7892DC3D8" country="Vanuatu" county="Espiritu Santo" latitude="-15.2175" longLatPrecision="5" longitude="166.68706" municipality="Tasmate" name="Pae River" stateProvince="Sanma Province">
<normalizedToken originalValue="Paé">Pae</normalizedToken>
River
</location>
,
<geoCoordinate degrees="15.2175" direction="south" orientation="latitude" precision="5" value="-15.2175">15.2175°S</geoCoordinate>
,
<geoCoordinate degrees="166.68706" direction="east" orientation="longitude" precision="1" value="166.68706">166.68706°E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.39" unit="m" value="139.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.39" unit="m" value="139.0">139 m</elevation>
</quantity>
,
<collectingDate value="2006-11-11">11.XI.2006</collectingDate>
, leg.
<collectorName>A.H. Staniczek</collectorName>
and
<collectorName>M. Pallmann</collectorName>
&quot;, with the following barcode number: ODO 000245 K.
<emphasis bold="true" pageId="0" pageNumber="129">
(
<specimenCount type="male">2 ♂♂</specimenCount>
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">
SNMS,
<specimenCount type="male">2 ♂♂</specimenCount>
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">MNHN).</emphasis>
&quot;
<collectingCountry name="Vanuatu">Vanuatu</collectingCountry>
,
<collectingRegion country="Vanuatu" name="Sanma">Sanma Province</collectingRegion>
, Espiritu Santo, surroundings of Tasmate,
<location LSID="urn:lsid:plazi:treatment:D8738B6ACDCB5BC9AC8E9FE4849ED810:740EE93AF4D4F70F53B7D30E5BF5B098" country="Vanuatu" county="Espiritu Santo" latitude="-15.2175" longLatPrecision="5" longitude="166.68706" municipality="Tasmate" name="Mamasa River" stateProvince="Sanma Province">Mamasa River</location>
,
<geoCoordinate degrees="15.20976" direction="south" orientation="latitude" precision="1" value="-15.20976">15.20976°S</geoCoordinate>
,
<geoCoordinate degrees="166.67705" direction="east" orientation="longitude" precision="1" value="166.67705">166.67705°E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20 m</elevation>
</quantity>
,
<collectingDate value="2006-11-09">9.XI.2006</collectingDate>
, leg.
<collectorName>A. H. Staniczek</collectorName>
and
<collectorName>M. Pallmann</collectorName>
&quot;, SNMS specimens with the following barcodes: ODO 000243 K, ODO 000244 K
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="description">
<paragraph pageId="0" pageNumber="129">Description of the female.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Head</emphasis>
: Labium overall pale beige; labrum pale green with brown latero-posterior edges and small, slightly indented medially brown spot at posterior edge; anteclypeus, genae, and mandibles (expect for reddish tips) pale green; postclypeus translucent brown with green lateral edges; frons pale green, turning black posteriorly; scapes, pedicels, and flagella pale brown; vertex and rear of head black with line of setae and a bronze shimmer; three pale ocelli with small beige patch apical of the median ocellus; two white post-ocular spots; eyes pale green.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Thorax</emphasis>
: Prothorax dorsally black; laterally green; pronotum black medially with brown edges, latero-posterior corners rounded to acute angles and weakly explanate, mid-line obviously indented across the pronotum, hind lobe with raised ridge that is shorter than the width of the pronotum, is curved outward medially, and extends to a sharp, rounded point that protrudes posteriorly; mesostigmal plate black with green lateral edges and not significantly raised. Pterothorax with black carina; laterally mesepisternum with black stripe reaching the dorsal carina and extending more than 3/4 of the mesepisternum, pale green on latter 1/4 of mesepisternum; mesepimeron overall pale green with hints of yellow; metepisternum overall green with short, brown line located on metapleural suture near the base of the wings, extending ~ 1/6 the
<normalizedToken originalValue="suture’s">suture's</normalizedToken>
length; mesinfraepisternum pale green with apical brown spot; metepimeron green, with beige anterior portion; coxae and trochanters beige, femora dorsally dark brown and ventrally beige with black spines; tibiae pale brown with slightly darker, and smaller, spines than that of the femora; tarsi beige with dark brown edges and small, dense spines; pale brown tarsal claws that darken apically to reddish tips, claws with a small tooth located on the basal 1/4 of their length.
</paragraph>
<caption doi="10.3897/zookeys.1128.89751.figure10" httpUri="https://binary.pensoft.net/fig/767336" pageId="0" pageNumber="129" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 10.</emphasis>
<taxonomicName authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">
<emphasis italics="true" pageId="0" pageNumber="129">Vanuatubasis santoensis</emphasis>
</taxonomicName>
(♂ BYU)
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
dorsal thorax
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
lateral terminalia
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
dorsal terminalia
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
lateral habitus
<taxonomicName lsidName="V. santoensis" pageId="0" pageNumber="129" rank="species" species="santoensis">
<emphasis italics="true" pageId="0" pageNumber="129">V. santoensis</emphasis>
</taxonomicName>
(♀ BYU)
<emphasis bold="true" pageId="0" pageNumber="129">E</emphasis>
dorsal thorax
<emphasis bold="true" pageId="0" pageNumber="129">F</emphasis>
lateral terminalia. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Wings</emphasis>
: Hyaline with dark brown; pterostigma dark brown and rhombus-shaped, darkest on the edges; CuP halfway between antenodals in both wings; arculus just distal of second antenodal crossvein in both wings; discoidal cells unequal with FW dorsal edge being 1/2 as long as HW. Nodal index: 13/2-2/12 in FW and 12/2-2/11 in HW.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Abdomen</emphasis>
: Overall beige yellow with black dorsal stripe from S1-8 and lightening towards the edges, with pale brown setae; S1 and S2 green laterally; S3-S8 yellow laterally; S9 with pale dorsal stripe; S10 pale brown; ovipositor overall beige, with darker serrated ventral edge; stylus with rounded edges, pale brown and lightening apically; gonapophysis reddish brown with slightly serrated dorsal edge. Cerci roughly triangular, dark brown, and narrowing to a rounded apex, dorsal edge straight a ventral edge slightly rounded ventrally.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">Measurements</emphasis>
(mm)
</emphasis>
: total length 36-39 mm, abdomen 29-33 mm, HW 22-24 mm (
<emphasis italics="true" pageId="0" pageNumber="129">n</emphasis>
= 9).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="diagnosis">
<paragraph pageId="0" pageNumber="129">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Male.</emphasis>
<taxonomicName authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">
<emphasis italics="true" pageId="0" pageNumber="129">Vanuatubasis santoensis</emphasis>
</taxonomicName>
can be distinguished from other
<taxonomicName authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Vanuatubasis</emphasis>
</taxonomicName>
by its black pterostigma, pale blue postclypeus without any bars or maculations, dorso-posterior corner of mesostigmal plate not raised in an auricle (or if elevated, only slightly so).
<emphasis bold="true" pageId="0" pageNumber="129">Female.</emphasis>
<taxonomicName authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">
<emphasis italics="true" pageId="0" pageNumber="129">Vanuatubasis santoensis</emphasis>
</taxonomicName>
can be distinguished from other females in this genus, by its dark brown to black pterostigma, uniformly colored postclypeus (sometimes with pale spots), frons with a dark triangle extending from the base of the median ocelli, and the dark area on the mesepisternum reaching the black spot at the posterior end of the mesopleural suture.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="variation">
<paragraph pageId="0" pageNumber="129">Variation.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Male.</emphasis>
Postclypeal maculation more prominent than in some individuals. Pterothorax sometimes with variable short black stripes.
<emphasis bold="true" pageId="0" pageNumber="129">Female.</emphasis>
Thoracic coloring varies from yellow to green, presumably due to the age of the individual.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="distribution">
<paragraph pageId="0" pageNumber="129">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="129">Espiritu Santo, Vanuatu.</paragraph>
</subSubSection>
</treatment>
</document>
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