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<document id="D319F90114E5CB10FC9A6149FE7855B3" ID-CLB-Dataset="4067" ID-DOI="10.11646/phytotaxa.522.4.2" ID-GBIF-Dataset="0e37c39e-2994-495d-9960-0cc7e8df1122" ID-ISSN="1179-3163" ID-Zenodo-Dep="5565838" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634105212392" checkinUser="plazi" docAuthor="He, Meng, He, Zhao-Rong, Zhang, Li-Bing &amp; Zhou, Xin-Mao" docDate="2021" docId="C430543B6D57BC51FF52CBA99681F80C" docLanguage="en" docName="phytotaxa.522.4.2.pdf" docOrigin="Phytotaxa 522 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.522.4.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Selaginella qingchengshanensis Li Bing Zhang &amp; X. M. Zhou 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="287" masterDocId="38092C436D56BC53FFDAC83D922DFFAD" masterDocTitle="Selaginella qingchengshanensis (sect. Heterostachys; Selaginellaceae), a new species from Sichuan, China" masterLastPageNumber="293" masterPageNumber="285" pageNumber="286" updateTime="1699459722089" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="07AF968700DED5C949F530E0238C120D">Selaginella qingchengshanensis (sect. Heterostachys; Selaginellaceae), a new species from Sichuan, China</mods:title>
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<mods:namePart id="E22DA3914FA9AC228EE359E378BF43EC">He, Meng</mods:namePart>
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<mods:affiliation id="AAB89D0D304AA59F70EC312FAB4645AF">School of Ecology and Environmental Science, Yunnan University, Kunming, 650091 Yunnan, China &amp; hemeng 968 @ 163. com; https: // orcid. org / 0000 - 0002 - 7609 - 6251</mods:affiliation>
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<mods:namePart id="53A8E4093468369867E96CAB0A3E7663">He, Zhao-Rong</mods:namePart>
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<mods:affiliation id="6A80B48CDAB12801C49BB6CE48A69572">School of Life Sciences, Yunnan University, East Outer Ring Road, Chenggong District, Kunming, Yunnan 650500, China &amp; zhrhe @ ynu. edu. cn; https: // orcid. org / 0000 - 0001 - 6492 - 3689</mods:affiliation>
<mods:nameIdentifier id="50205E46B0996E36485147DA2515088C" type="email">zhrhe@ynu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="6C66B9FA4E5A8B2F864FCA9ABE6705E3">Zhang, Li-Bing</mods:namePart>
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<mods:affiliation id="89DF0266E70CFFF1629DB152EB01BEAF">Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A; Chengdu Institute of Biology, Chinese Academy of Sciences &amp; Libing. Zhang @ mobot. org; https: // orcid. org / 0000 - 0002 - 4905 - 040 X</mods:affiliation>
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<mods:namePart id="44AEDC93761FC694893FEC2A67BC3451">Zhou, Xin-Mao</mods:namePart>
<mods:nameIdentifier id="EE2A48F0EEDC2DD0835307553DB86392" type="ORCID">0000-0003-3555-7784</mods:nameIdentifier>
<mods:affiliation id="6E74D33BF6423590B692D189E5493419">School of Ecology and Environmental Science, Yunnan University, Kunming, 650091 Yunnan, China &amp; xinmao. zhou @ ynu. edu. cn; http: // orcid. org / 0000 - 0003 - 3555 - 7784</mods:affiliation>
<mods:nameIdentifier id="FD0AA33E5C3D8D67D2A180027D0014E9" type="email">xinmao.zhou@ynu.edu.cn</mods:nameIdentifier>
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<mods:title id="9010DDE7844CC1600DF95AAC468DFD7F">Phytotaxa</mods:title>
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<mods:date id="148ED41E412DF575FF19DB063743248D">2021</mods:date>
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<treatment id="C430543B6D57BC51FF52CBA99681F80C" ID-DOI="http://doi.org/10.5281/zenodo.5565836" ID-GBIF-Taxon="188628601" ID-Zenodo-Dep="5565836" LSID="urn:lsid:plazi:treatment:C430543B6D57BC51FF52CBA99681F80C" httpUri="http://treatment.plazi.org/id/C430543B6D57BC51FF52CBA99681F80C" lastPageId="2" lastPageNumber="287" pageId="1" pageNumber="286">
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<paragraph id="4C26E52D6D57BC52FF52CBA99186FC03" blockId="1.[136,939,916,943]" box="[136,939,916,943]" pageId="1" pageNumber="286">
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<taxonomicName id="8B999EAE6D57BC52FF52CBA99166FC03" ID-CoL="8SKPV" authority="Li Bing Zhang &amp; X.M. Zhou" authorityName="Li Bing Zhang &amp; X. M. Zhou" authorityYear="2021" box="[136,843,916,943]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="286" phylum="Tracheophyta" rank="species" species="qingchengshanensis" status="sp. nov.">
<emphasis id="7EED393F6D57BC52FF52CBA993D6FC02" bold="true" box="[136,507,916,943]" italics="true" pageId="1" pageNumber="286">Selaginella qingchengshanensis</emphasis>
Li Bing Zhang &amp; X.M. Zhou
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,
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<taxonomicNameLabel id="65DE84446D57BC52FC8CCBA89186FC03" box="[854,939,917,942]" pageId="1" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<emphasis id="7EED393F6D57BC52FF52CB8192F7FC7C" bold="true" box="[136,218,956,977]" pageId="1" pageNumber="286">Type:—</emphasis>
<materialsCitation id="FCF1EF706D57BC52FF00CB819705FC5B" ID-GBIF-Occurrence="3383804303" collectingDate="2020-08-23" collectionCode="Z" collectorName="Liang &amp; X. Pu" country="China" county="Dujiangyan City" elevation="960" isTailEnumeration="true" latitude="30.907873" location="Qingchengshan Town" longLatPrecision="1" longitude="103.56846" municipality="Qingcheng Mountain" pageId="1" pageNumber="286" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<collectingCountry id="348EA5BD6D57BC52FF00CB819304FC7C" box="[218,297,956,977]" name="China" pageId="1" pageNumber="286">CHINA</collectingCountry>
.
<collectingRegion id="8E5D2BCF6D57BC52FEEDCB8193A5FC7F" box="[311,392,956,978]" country="China" name="Sichuan" pageId="1" pageNumber="286">Sichuan</collectingRegion>
:
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,
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,
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,
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, elev.
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<elevation id="C7B4021E6D57BC52FB08CB81973BFC7C" box="[1234,1302,956,978]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="1" pageNumber="286" unit="m" value="960.0">960 m</elevation>
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,
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,
<geoCoordinate id="29AD83EA6D57BC52FF67CBDD9315FC58" box="[189,312,992,1013]" degrees="30.907873" direction="north" orientation="latitude" pageId="1" pageNumber="286" precision="1" value="30.907873">30.907873N</geoCoordinate>
,
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<collectingDate id="28633A056D57BC52FE9BCBDD93F6FC58" box="[321,475,992,1013]" pageId="1" pageNumber="286" value="2020-08-23">23 August 2020</collectingDate>
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, in humid place in evergreen broadleaf forest,
<emphasis id="7EED393F6D57BC52FC43CBDD9653FC58" box="[921,1150,992,1013]" italics="true" pageId="1" pageNumber="286">
<collectionCode id="2A887DE86D57BC52FC43CBDD9184FC58" box="[921,937,992,1013]" country="Switzerland" name="Universität Zürich" pageId="1" pageNumber="286" type="Herbarium">Z</collectionCode>
.-
<collectionCode id="2A887DE86D57BC52FC69CBDD91EFFC58" box="[947,962,992,1013]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="1" pageNumber="286" type="Herbarium">L</collectionCode>
<collectorName id="E16C80FB6D57BC52FC12CBDD962DFC58" box="[968,1024,992,1013]" pageId="1" pageNumber="286">Liang</collectorName>
&amp;
<collectorName id="E16C80FB6D57BC52FBC4CBDD9679FC58" box="[1054,1108,992,1013]" pageId="1" pageNumber="286">X. Pu</collectorName>
085
</emphasis>
(
<typeStatus id="93225B8F6D57BC52FB50CBDD96CDFC5B" box="[1162,1248,992,1014]" pageId="1" pageNumber="286" type="holotype">holotype</typeStatus>
<collectionCode id="2A887DE86D57BC52FB3CCBDD9733FC5B" box="[1254,1310,992,1014]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14817" lsid="urn:lsid:biocol.org:col:14817" name="Yunnan University" pageId="1" pageNumber="286" type="Herbarium">PYU</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0483B6A66D57BC52FF52CC3E969FF833" pageId="1" pageNumber="286" type="diagnosis">
<paragraph id="4C26E52D6D57BC52FF52CC3E90B7FB04" blockId="1.[136,1452,956,1194]" pageId="1" pageNumber="286">
<emphasis id="7EED393F6D57BC52FF52CC3E9326FBB4" bold="true" box="[136,267,1027,1049]" pageId="1" pageNumber="286">Diagnosis:—</emphasis>
<taxonomicName id="8B999EAE6D57BC52FED1CC399012FBB4" authorityName="Li Bing Zhang &amp; X. M. Zhou" authorityYear="2021" box="[267,575,1028,1049]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="286" phylum="Tracheophyta" rank="species" species="qingchengshanensis">
<emphasis id="7EED393F6D57BC52FED1CC399012FBB4" box="[267,575,1028,1049]" italics="true" pageId="1" pageNumber="286">Selaginella qingchengshanensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="8B999EAE6D57BC52FD18CC39963FFBB7" authority="Alston (1934: 287)" authorityName="Alston" authorityPageNumber="287" authorityYear="1934" box="[706,1042,1028,1050]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="286" phylum="Tracheophyta" rank="species" species="amblyphylla">
<emphasis id="7EED393F6D57BC52FD18CC39917EFBB4" box="[706,851,1028,1049]" italics="true" pageId="1" pageNumber="286">S. amblyphylla</emphasis>
<bibRefCitation id="280898DC6D57BC52FC83CC39963FFBB7" author="Alston, A. H. G." box="[857,1042,1028,1050]" pageId="1" pageNumber="286" pagination="261 - 304" refId="ref2665" refString="Alston, A. H. G. (1934) An enumeration of the Chinese species of Selaginella. Bulletin of the Fan Memorial Institute of Biology: Botany 5: 261 - 304." type="journal article" year="1934">Alston (1934: 287)</bibRefCitation>
</taxonomicName>
in having plants creeping to suberect and the apex of dorsal leaves long aristate, but the new species has axillary leaves ovate-triangular to broadly ovate (
<figureCitation id="D4A2F9A86D57BC52FB2DCC15976EFB90" box="[1271,1347,1064,1086]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1H</figureCitation>
; vs. ovatelanceolate or triangular-lanceolate in
<taxonomicName id="8B999EAE6D57BC52FDFACC719083FBCC" authorityName="Alston" authorityYear="1934" box="[544,686,1100,1121]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="286" phylum="Tracheophyta" rank="species" species="amblyphylla">
<emphasis id="7EED393F6D57BC52FDFACC719083FBCC" box="[544,686,1100,1121]" italics="true" pageId="1" pageNumber="286">S. amblyphylla</emphasis>
</taxonomicName>
), sterile leaf margins denticulate to slightly ciliolate at base (
<figureCitation id="D4A2F9A86D57BC52FB35CC719762FBCC" box="[1263,1359,1100,1122]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1G–I</figureCitation>
; vs. longciliate at base in
<taxonomicName id="8B999EAE6D57BC52FE87CC4D93C1FB28" authorityName="Alston" authorityYear="1934" box="[349,492,1136,1157]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="286" phylum="Tracheophyta" rank="species" species="amblyphylla">
<emphasis id="7EED393F6D57BC52FE87CC4D93C1FB28" box="[349,492,1136,1157]" italics="true" pageId="1" pageNumber="286">S. amblyphylla</emphasis>
</taxonomicName>
), and arista of dorsal leaves nearly perpendicular to stems and branches in living plants (
<figureCitation id="D4A2F9A86D57BC52FAE6CC4D97A8FB28" box="[1340,1413,1136,1158]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1L</figureCitation>
; vs. parallel to stems and branches in
<taxonomicName id="8B999EAE6D57BC52FE27CCA990A1FB04" authorityName="Alston" authorityYear="1934" box="[509,652,1172,1193]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="286" phylum="Tracheophyta" rank="species" species="amblyphylla">
<emphasis id="7EED393F6D57BC52FE27CCA990A1FB04" box="[509,652,1172,1193]" italics="true" pageId="1" pageNumber="286">S. amblyphylla</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="4C26E52D6D57BC52FF52CCE5969FF833" blockId="1.[136,1452,1240,1950]" pageId="1" pageNumber="286">
<emphasis id="7EED393F6D57BC52FF52CCE592FEFB5F" bold="true" box="[136,211,1240,1266]" pageId="1" pageNumber="286">Plants</emphasis>
terrestrial, creeping to suberect,
<quantity id="8B6148C86D57BC52FD9ECCE59088FB5E" box="[580,677,1240,1267]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="2.0" metricValueMin="0.6" pageId="1" pageNumber="286" unit="cm" value="13.0" valueMax="20.0" valueMin="6.0">6–20 cm</quantity>
(
<figureCitation id="D4A2F9A86D57BC52FD6FCCE59123FB5F" box="[693,782,1240,1266]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1A</figureCitation>
,
<figureCitation id="D4A2F9A86D57BC52FCC0CCE59116FB5F" box="[794,827,1240,1266]" captionStart="FIGURE 3" captionStartId="4.[136,229,1366,1388]" captionTargetBox="[374,1243,198,1334]" captionTargetId="figure-36@4.[330,1258,190,1342]" captionTargetPageId="4" captionText="FIGURE 3. Type of Selaginella qingchengshanensis (Z.-L Liang &amp; X.Pu 085).—A. Dorsal view of strobili.—B.Ventral view of strobili.— C. Portion of branch showing axillary leaf and ventral leaf.—D. Portion of branch showing dorsal leaves.—E. Portion of branch showing ventral leaves." figureDoi="http://doi.org/10.5281/zenodo.5565844" httpUri="https://zenodo.org/record/5565844/files/figure.png" pageId="1" pageNumber="286">3F</figureCitation>
).
<emphasis id="7EED393F6D57BC52FC95CCE591C8FB5F" bold="true" box="[847,997,1240,1266]" pageId="1" pageNumber="286">Rhizophores</emphasis>
restricted to lower part of stem, born on ventral side in axils of main branches, glabrous,
<quantity id="8B6148C86D57BC52FD7ECCC1910CFABA" box="[676,801,1276,1303]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.5" metricValueMin="0.5" pageId="1" pageNumber="286" unit="cm" value="2.5" valueMax="4.5" valueMin="0.5">0.5–4.5 cm</quantity>
long (
<figureCitation id="D4A2F9A86D57BC52FCB2CCC19190FABB" box="[872,957,1276,1302]" captionStart="FIGURE 3" captionStartId="4.[136,229,1366,1388]" captionTargetBox="[374,1243,198,1334]" captionTargetId="figure-36@4.[330,1258,190,1342]" captionTargetPageId="4" captionText="FIGURE 3. Type of Selaginella qingchengshanensis (Z.-L Liang &amp; X.Pu 085).—A. Dorsal view of strobili.—B.Ventral view of strobili.— C. Portion of branch showing axillary leaf and ventral leaf.—D. Portion of branch showing dorsal leaves.—E. Portion of branch showing ventral leaves." figureDoi="http://doi.org/10.5281/zenodo.5565844" httpUri="https://zenodo.org/record/5565844/files/figure.png" pageId="1" pageNumber="286">Fig. 3F</figureCitation>
).
<emphasis id="7EED393F6D57BC52FC0BCCC19677FABB" bold="true" box="[977,1114,1276,1302]" pageId="1" pageNumber="286">Main stems</emphasis>
with one flat vascular bundle, stramineous, branched from near base, 2 to 3 times pinnately branched, branches many, ultimate branches 3.3–5.0 mm wide (
<figureCitation id="D4A2F9A86D57BC52FEDACD799355FAF3" box="[256,376,1348,1374]" captionStart="FIGURE 3" captionStartId="4.[136,229,1366,1388]" captionTargetBox="[374,1243,198,1334]" captionTargetId="figure-36@4.[330,1258,190,1342]" captionTargetPageId="4" captionText="FIGURE 3. Type of Selaginella qingchengshanensis (Z.-L Liang &amp; X.Pu 085).—A. Dorsal view of strobili.—B.Ventral view of strobili.— C. Portion of branch showing axillary leaf and ventral leaf.—D. Portion of branch showing dorsal leaves.—E. Portion of branch showing ventral leaves." figureDoi="http://doi.org/10.5281/zenodo.5565844" httpUri="https://zenodo.org/record/5565844/files/figure.png" pageId="1" pageNumber="286">Fig. 3A–F</figureCitation>
).
<emphasis id="7EED393F6D57BC52FE51CD7993F0FAF3" bold="true" box="[395,477,1348,1374]" pageId="1" pageNumber="286">Leaves</emphasis>
herbaceous.
<emphasis id="7EED393F6D57BC52FDAACD799135FAF3" bold="true" box="[624,792,1348,1374]" pageId="1" pageNumber="286">Ventral leaves</emphasis>
on branches contiguous, slightly ascending, asymmetrical, those on main stems slightly larger than those on branches; oblong or oblong-ovate, 2.3–3.2 ×
<quantity id="8B6148C86D57BC52FB4CCD559733FA2E" box="[1174,1310,1384,1411]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="1" pageNumber="286" unit="mm" value="1.3" valueMax="1.5" valueMin="1.1">1.1–1.5 mm</quantity>
, apex obtuse to round; acroscopic base rounded, slightly enlarged, overlapping stem and branches, margin denticulate to slightly ciliolate; basiscopic base rounded, margin entire or minutely denticulate (
<figureCitation id="D4A2F9A86D57BC52FC6ECD8D9621FA67" box="[948,1036,1456,1482]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1G</figureCitation>
;
<figureCitation id="D4A2F9A86D57BC52FBCDCD8D9674FA66" box="[1047,1113,1456,1483]" captionStart="FIGURE 3" captionStartId="4.[136,229,1366,1388]" captionTargetBox="[374,1243,198,1334]" captionTargetId="figure-36@4.[330,1258,190,1342]" captionTargetPageId="4" captionText="FIGURE 3. Type of Selaginella qingchengshanensis (Z.-L Liang &amp; X.Pu 085).—A. Dorsal view of strobili.—B.Ventral view of strobili.— C. Portion of branch showing axillary leaf and ventral leaf.—D. Portion of branch showing dorsal leaves.—E. Portion of branch showing ventral leaves." figureDoi="http://doi.org/10.5281/zenodo.5565844" httpUri="https://zenodo.org/record/5565844/files/figure.png" pageId="1" pageNumber="286">3C, E</figureCitation>
).
<emphasis id="7EED393F6D57BC52FBB1CD8D9725FA67" bold="true" box="[1131,1288,1456,1482]" pageId="1" pageNumber="286">Dorsal leaves</emphasis>
asymmetrical, those on main stems more sparse than those on branches, ovate or ovate-lanceolate, 1.8–2.9 ×
<quantity id="8B6148C86D57BC52FB69CDE99766FA42" box="[1203,1355,1492,1519]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="9.5" metricValueMin="8.0" pageId="1" pageNumber="286" unit="mm" value="0.875" valueMax="0.95" valueMin="0.8">0.8–0.95 mm</quantity>
, slightly carinate, base round obliquely attenuate, apex aristate (arista as long as the leaves), reflexed, but perpendicular to axis in living plants, margin denticulate to slightly ciliolate (
<figureCitation id="D4A2F9A86D57BC52FD0BCE219130F99B" box="[721,797,1564,1590]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1I</figureCitation>
,
<figureCitation id="D4A2F9A86D57BC52FCF2CE219163F99B" box="[808,846,1564,1590]" captionStart="FIGURE 3" captionStartId="4.[136,229,1366,1388]" captionTargetBox="[374,1243,198,1334]" captionTargetId="figure-36@4.[330,1258,190,1342]" captionTargetPageId="4" captionText="FIGURE 3. Type of Selaginella qingchengshanensis (Z.-L Liang &amp; X.Pu 085).—A. Dorsal view of strobili.—B.Ventral view of strobili.— C. Portion of branch showing axillary leaf and ventral leaf.—D. Portion of branch showing dorsal leaves.—E. Portion of branch showing ventral leaves." figureDoi="http://doi.org/10.5281/zenodo.5565844" httpUri="https://zenodo.org/record/5565844/files/figure.png" pageId="1" pageNumber="286">3D</figureCitation>
).
<emphasis id="7EED393F6D57BC52FCBBCE21963DF99B" bold="true" box="[865,1040,1564,1590]" pageId="1" pageNumber="286">Axillary leaves</emphasis>
symmetrical, those on main stem not obviously larger than those on branches, ovate-triangular or broadly ovate, 1.5–2.2 × 1.0–
<quantity id="8B6148C86D57BC52FBA9CE7D96F5F9F6" box="[1139,1240,1600,1627]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="286" unit="mm" value="1.65">1.65 mm</quantity>
, base exauriculate, margin denticulate to shortly ciliolate, apex obtuse or slightly acute (
<figureCitation id="D4A2F9A86D57BC52FC5BCE5991F5F9D3" box="[897,984,1636,1662]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1H</figureCitation>
,
<figureCitation id="D4A2F9A86D57BC52FC3ECE599625F9D3" box="[996,1032,1636,1662]" captionStart="FIGURE 3" captionStartId="4.[136,229,1366,1388]" captionTargetBox="[374,1243,198,1334]" captionTargetId="figure-36@4.[330,1258,190,1342]" captionTargetPageId="4" captionText="FIGURE 3. Type of Selaginella qingchengshanensis (Z.-L Liang &amp; X.Pu 085).—A. Dorsal view of strobili.—B.Ventral view of strobili.— C. Portion of branch showing axillary leaf and ventral leaf.—D. Portion of branch showing dorsal leaves.—E. Portion of branch showing ventral leaves." figureDoi="http://doi.org/10.5281/zenodo.5565844" httpUri="https://zenodo.org/record/5565844/files/figure.png" pageId="1" pageNumber="286">3C</figureCitation>
).
<emphasis id="7EED393F6D57BC52FBC6CE59965BF9D3" bold="true" box="[1052,1142,1636,1662]" pageId="1" pageNumber="286">Strobili</emphasis>
solitary, terminal, compact, dorsiventrally complanate, resupinate, 3.7–5.6 ×
<quantity id="8B6148C86D57BC52FD64CEB59164F90E" box="[702,841,1672,1699]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.7" metricValueMin="1.4" pageId="1" pageNumber="286" unit="mm" value="2.05" valueMax="2.7" valueMin="1.4">1.4–2.7 mm</quantity>
(
<figureCitation id="D4A2F9A86D57BC52FC81CEB59198F90F" box="[859,949,1672,1698]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1B</figureCitation>
,
<figureCitation id="D4A2F9A86D57BC52FC1FCEB59621F90F" box="[965,1036,1672,1698]" captionStart="FIGURE 3" captionStartId="4.[136,229,1366,1388]" captionTargetBox="[374,1243,198,1334]" captionTargetId="figure-36@4.[330,1258,190,1342]" captionTargetPageId="4" captionText="FIGURE 3. Type of Selaginella qingchengshanensis (Z.-L Liang &amp; X.Pu 085).—A. Dorsal view of strobili.—B.Ventral view of strobili.— C. Portion of branch showing axillary leaf and ventral leaf.—D. Portion of branch showing dorsal leaves.—E. Portion of branch showing ventral leaves." figureDoi="http://doi.org/10.5281/zenodo.5565844" httpUri="https://zenodo.org/record/5565844/files/figure.png" pageId="1" pageNumber="286">3A–B</figureCitation>
); sporophylls dimorphic, not whitemargined; dorsal sporophylls ovate-lanceolate, with sporophyll-pteryx nearly complete and ciliolate, margin ciliolate, apex acute to acuminate (
<figureCitation id="D4A2F9A86D57BC52FE74CEED902BF947" box="[430,518,1744,1770]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1E</figureCitation>
); ventral sporophylls ovate, carinate, margin denticulate to ciliolate, apex aristate (
<figureCitation id="D4A2F9A86D57BC52FF4ACEC992C9F8A3" box="[144,228,1780,1806]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1F</figureCitation>
); megasporophylls distributed at the base of strobili, not intersect with microsporophylls.
<emphasis id="7EED393F6D57BC52FB0BCEC9974DF8A3" bold="true" box="[1233,1376,1780,1806]" pageId="1" pageNumber="286">Megaspores</emphasis>
white, oblate spheroid to subglobose; prominent laesurae extend 2/3 of the distance to the equator; proximal surfaces densely papillate or verrucate; distal surfaces papillate, vermiculate, or verrucate, surfaces densely covered with fine spines (
<figureCitation id="D4A2F9A86D57BC52FF4ACF5D933DF8D7" box="[144,272,1888,1914]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1M–O</figureCitation>
).
<emphasis id="7EED393F6D57BC52FEFBCF5D9398F8D7" bold="true" box="[289,437,1888,1914]" pageId="1" pageNumber="286">Microspores</emphasis>
yellowish orange, spherical tetrahedral; laesurae nearly extend to the equator; both proximal and laesurae surfaces with low fused tuberculate, densely covered with fine spines (
<figureCitation id="D4A2F9A86D57BC52FBE8CFB99688F833" box="[1074,1189,1924,1950]" captionStart="FIGURE 1" captionStartId="2.[136,229,1426,1448]" captionTargetBox="[298,1290,190,1402]" captionTargetId="figure-36@2.[298,1290,190,1402]" captionTargetPageId="2" captionText="FIGURE 1. Selaginella qingchengshanensis.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface." figureDoi="http://doi.org/10.5281/zenodo.5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="1" pageNumber="286">Fig. 1J–K</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="18E6B5A56D54BC51FF52CDAF9689F9B8" ID-DOI="http://doi.org/10.5281/zenodo.5565840" ID-Zenodo-Dep="5565840" httpUri="https://zenodo.org/record/5565840/files/figure.png" pageId="2" pageNumber="287" startId="2.[136,229,1426,1448]" targetBox="[298,1290,190,1402]" targetPageId="2">
<paragraph id="4C26E52D6D54BC51FF52CDAF9689F9B8" blockId="2.[136,1452,1426,1557]" pageId="2" pageNumber="287">
<emphasis id="7EED393F6D54BC51FF52CDAF932CFA0A" bold="true" box="[136,257,1426,1448]" pageId="2" pageNumber="287">FIGURE 1.</emphasis>
<taxonomicName id="8B999EAE6D54BC51FED0CDAE906DFA05" authorityName="Li Bing Zhang &amp; X. M. Zhou" authorityYear="2021" box="[266,576,1427,1448]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="2" pageNumber="287" phylum="Tracheophyta" rank="species" species="qingchengshanensis">
<emphasis id="7EED393F6D54BC51FED0CDAE906DFA05" box="[266,576,1427,1448]" italics="true" pageId="2" pageNumber="287">Selaginella qingchengshanensis</emphasis>
</taxonomicName>
.—A. Habit.—B. Ventral view of strobili on branch.—C. Dorsal view of branches.—D. Ventral view of branches.—E. Dorsal sporophyll.—F. Ventral sporophyll.—G. Ventral leaf.—H. Axillary leaf.—I. Dorsal leaf.—J. Proximal surface of microspore.—K. Distal surface of microspore.—L. Portion of branch, showing the upturned dorsal leaves.—M. Proximal surface of megaspore.—N. Distal surface of megaspore.—O. Detail of megaspore on distal surface.
</paragraph>
</caption>
<subSubSection id="0483B6A66D54BC51FF67CE7E90D5F92F" pageId="2" pageNumber="287" type="etymology">
<paragraph id="4C26E52D6D54BC51FF67CE7E90D5F92F" blockId="2.[136,1452,1603,1953]" pageId="2" pageNumber="287">
<emphasis id="7EED393F6D54BC51FF67CE7E9349F9F0" bold="true" box="[189,356,1603,1629]" pageId="2" pageNumber="287">Etymology:—</emphasis>
The specific epithet refers to the
<typeStatus id="93225B8F6D54BC51FD01CE7E9126F9F0" box="[731,779,1603,1629]" pageId="2" pageNumber="287">type</typeStatus>
locality in Qingcheng Mountain,
<collectingRegion id="8E5D2BCF6D54BC51FB56CE7E9778F9F0" box="[1164,1365,1603,1629]" country="China" name="Sichuan" pageId="2" pageNumber="287">Sichuan Province</collectingRegion>
,
<collectingCountry id="348EA5BD6D54BC51FABBCE7E9785F9F0" box="[1377,1448,1603,1629]" name="China" pageId="2" pageNumber="287">China</collectingCountry>
. Chinese name: Ḧ城山卷柏 (qing cheng shan juan bai).
</paragraph>
</subSubSection>
<subSubSection id="0483B6A66D54BC51FF67CEB69681F80C" pageId="2" pageNumber="287" type="distribution">
<paragraph id="4C26E52D6D54BC51FF67CEB69145F940" blockId="2.[136,1452,1603,1953]" pageId="2" pageNumber="287">
<emphasis id="7EED393F6D54BC51FF67CEB6903CF908" bold="true" box="[189,529,1675,1701]" pageId="2" pageNumber="287">Distribution and habitat:—</emphasis>
Known only from the
<typeStatus id="93225B8F6D54BC51FCFACEB6917DF908" box="[800,848,1675,1701]" pageId="2" pageNumber="287">type</typeStatus>
locality in Qingcheng Mountain, Dujiangyan City,
<collectingRegion id="8E5D2BCF6D54BC51FF52CE929362F964" box="[136,335,1711,1737]" country="China" name="Sichuan" pageId="2" pageNumber="287">Sichuan province</collectingRegion>
,
<collectingCountry id="348EA5BD6D54BC51FE86CE92938FF964" box="[348,418,1711,1737]" name="China" pageId="2" pageNumber="287">China</collectingCountry>
, at
<quantity id="8B6148C86D54BC51FE10CE929076F964" box="[458,603,1711,1738]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.5" metricValueMin="8.5" pageId="2" pageNumber="287" unit="m" value="950.0" valueMax="1050.0" valueMin="850.0">850–1050 m</quantity>
. This species grows in humid places in evergreen broadleaved forests. It is common on the roadside of tourist trails on Qingcheng Mountain.
</paragraph>
<paragraph id="4C26E52D6D54BC51FF67CECA930BF8F4" blockId="2.[136,1452,1603,1953]" pageId="2" pageNumber="287">
<emphasis id="7EED393F6D54BC51FF67CECA93FEF8BC" bold="true" box="[189,467,1783,1809]" pageId="2" pageNumber="287">Conservation status:—</emphasis>
<taxonomicName id="8B999EAE6D54BC51FE0ECECA9113F8BC" authorityName="Li Bing Zhang &amp; X. M. Zhou" authorityYear="2021" box="[468,830,1783,1809]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="2" pageNumber="287" phylum="Tracheophyta" rank="species" species="qingchengshanensis">
<emphasis id="7EED393F6D54BC51FE0ECECA9113F8BC" box="[468,830,1783,1809]" italics="true" pageId="2" pageNumber="287">Selaginella qingchengshanensis</emphasis>
</taxonomicName>
is only known from Qingcheng Mountain, but it has relatively wide distribution there. We consider it to be Near Threatened (NT) based on the IUCN red list criterion (
<bibRefCitation id="280898DC6D54BC51FF4ACF029337F8F4" author="IUCN" box="[144,282,1855,1881]" pageId="2" pageNumber="287" refId="ref2954" refString="IUCN (2017) The IUCN Red List of Threatened Species. Version 2017 - 1. Available at: www. iucnredlist. org (accessed 13 October 2021)" type="book" year="2017">IUCN 2017</bibRefCitation>
).
</paragraph>
<paragraph id="4C26E52D6D54BC51FF67CF5E9681F80C" blockId="2.[136,1452,1603,1953]" pageId="2" pageNumber="287">
<materialsCitation id="FCF1EF706D54BC51FF67CF5E9027F80C" ID-GBIF-Occurrence="3383804302" accessionNumber="DJY04053, DJY06425" collectingDate="2019-08-23" collectionCode="F" collectorName="Qingcheng Mountain, X." country="China" location="Qingcheng Mountain" pageId="2" pageNumber="287" specimenCount="2" stateProvince="Sichuan" typeStatus="paratype">
<emphasis id="7EED393F6D54BC51FF67CF5E90C0F8D0" bold="true" box="[189,749,1891,1917]" pageId="2" pageNumber="287">
Additional specimens examined (
<typeStatus id="93225B8F6D54BC51FD9CCF5E90EFF8D0" box="[582,706,1891,1917]" pageId="2" pageNumber="287" type="paratype">paratypes</typeStatus>
):—
</emphasis>
<collectingCountry id="348EA5BD6D54BC51FD36CF5E9165F8D0" box="[748,840,1891,1917]" name="China" pageId="2" pageNumber="287">CHINA</collectingCountry>
.
<collectingRegion id="8E5D2BCF6D54BC51FC89CF5E9631F8D0" box="[851,1052,1891,1917]" country="China" name="Sichuan" pageId="2" pageNumber="287">Sichuan Province</collectingRegion>
:
<collectorName id="E16C80FB6D54BC51FBFDCF5E9711F8D0" box="[1063,1340,1891,1917]" pageId="2" pageNumber="287">
<location id="4946B3F66D54BC51FBFDCF5E9735F8D0" LSID="urn:lsid:plazi:treatment:C430543B6D57BC51FF52CBA99681F80C:4946B3F66D54BC51FBFDCF5E9735F8D0" box="[1063,1304,1891,1917]" country="China" name="Qingcheng Mountain" pageId="2" pageNumber="287" stateProvince="Sichuan">Qingcheng Mountain</location>
,
<emphasis id="7EED393F6D54BC51FAF9CF599711F8D0" box="[1315,1340,1892,1917]" italics="true" pageId="2" pageNumber="287">X.</emphasis>
</collectorName>
-
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<emphasis id="7EED393F6D54BC51FA8CCF5997AFF8D0" box="[1366,1410,1891,1917]" italics="true" pageId="2" pageNumber="287">. Ga</emphasis>
o
<emphasis id="7EED393F6D54BC51FA4DCF59938BF80C" italics="true" pageId="2" pageNumber="287">et al. DJY04053, DJY06425</emphasis>
(
<collectionCode id="2A887DE86D54BC51FE6FCFBA93D3F80C" box="[437,510,1927,1953]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12796" lsid="urn:lsid:biocol.org:col:12796" name="Chengdu Institute of Biology" pageId="2" pageNumber="287" type="Herbarium">CDBI</collectionCode>
);
</materialsCitation>
<materialsCitation id="FCF1EF706D54BC51FDC8CFBA9685F80C" ID-GBIF-Occurrence="3383804301" accessionNumber="DJY04053, DJY06425" box="[530,1192,1927,1953]" collectingDate="2019-08-23" collectionCode="CDBI, Z, L" collectorName="X. Pu" country="China" pageId="2" pageNumber="287" specimenCount="1" stateProvince="Sichuan Province">
<date id="3827C3ED6D54BC51FDC8CFBA90EBF80C" box="[530,710,1927,1953]" pageId="2" pageNumber="287" value="2019-08-23">
<collectingDate id="28633A056D54BC51FDC8CFBA90EBF80C" box="[530,710,1927,1953]" pageId="2" pageNumber="287" value="2019-08-23">23 August 2019</collectingDate>
</date>
,
<emphasis id="7EED393F6D54BC51FD0BCFB5967EF80C" box="[721,1107,1927,1953]" italics="true" pageId="2" pageNumber="287">
<collectionCode id="2A887DE86D54BC51FD0BCFB590C8F80C" box="[721,741,1928,1953]" country="Switzerland" name="Universität Zürich" pageId="2" pageNumber="287" type="Herbarium">Z</collectionCode>
.-
<collectionCode id="2A887DE86D54BC51FD2BCFB59129F80C" box="[753,772,1928,1953]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="2" pageNumber="287" type="Herbarium">L</collectionCode>
. Liang &amp;
<collectorName id="E16C80FB6D54BC51FCAECFB5919FF80C" box="[884,946,1928,1953]" pageId="2" pageNumber="287">X. Pu</collectorName>
056, 078, 097
</emphasis>
(
<collectionCode id="2A887DE86D54BC51FBB8CFBA968DF80C" box="[1122,1184,1927,1953]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14817" lsid="urn:lsid:biocol.org:col:14817" name="Yunnan University" pageId="2" pageNumber="287" type="Herbarium">PYU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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