442 lines
52 KiB
XML
442 lines
52 KiB
XML
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<mods:title id="01D1401615B9DA2AD4C28C63EDD8650F">A new species, new genus and new records of heterostigmatic mites (Acari: Heterostigmata) phoretic on scarab beetles of the subfamily Orphninae (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="3EBFE10B7418DC6B60B4A17DCDCBB7AC">Khaustov, Alexander A.</mods:namePart>
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<mods:namePart id="71F63DA2117ABF775B79FF56A9290831">Frolov, Andrey V.</mods:namePart>
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<mods:title id="EBD7D4682D3834A9C66156E499B412E4">Zootaxa</mods:title>
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<mods:date id="10226AD08D5EB20AE4870EE9DAD3D370">2018</mods:date>
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<treatment id="7E2487A4A5490E6759AEFF3CB37B18F9" ID-DOI="http://doi.org/10.5281/zenodo.5992844" ID-GBIF-Taxon="154496621" ID-Zenodo-Dep="5992844" LSID="urn:lsid:plazi:treatment:7E2487A4A5490E6759AEFF3CB37B18F9" httpUri="http://treatment.plazi.org/id/7E2487A4A5490E6759AEFF3CB37B18F9" lastPageId="9" lastPageNumber="189" pageId="7" pageNumber="188">
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<subSubSection id="BE976539A5490E6959AEFF3CB17D18BC" box="[151,820,152,178]" pageId="7" pageNumber="188" type="nomenclature">
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<paragraph id="F63236B2A5490E6959AEFF3CB17D18BC" blockId="7.[151,820,152,211]" box="[151,820,152,178]" pageId="7" pageNumber="188">
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<heading id="AD7A81DEA5490E6959AEFF3CB17D18BC" bold="true" box="[151,820,152,178]" fontSize="11" level="1" pageId="7" pageNumber="188" reason="1">
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<emphasis id="C4F9EAA0A5490E6959AEFF3CB17D18BC" bold="true" box="[151,820,152,178]" pageId="7" pageNumber="188">
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<taxonomicName id="318D4D31A5490E6959AEFF3CB0EF18BC" authority="(Mahunka, 1975)" authorityName="Khaustov & Frolov" authorityYear="2018" baseAuthorityName="Mahunka" baseAuthorityYear="1975" box="[151,678,152,178]" class="Insecta" family="Trogidae" genus="Troxodania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="problematica" status="comb. nov.">
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<emphasis id="C4F9EAA0A5490E6959AEFF3CB38018BC" bold="true" box="[151,457,152,178]" italics="true" pageId="7" pageNumber="188">Troxodania problematica</emphasis>
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(
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<bibRefCitation id="921C4B43A5490E6958E1FF3CB0D718BC" author="Mahunka, S." box="[472,670,152,178]" pageId="7" pageNumber="188" pagination="39 - 72" refId="ref7808" refString="Mahunka, S. (1975) Auf Insecten lebende Milben (Acari: Acarida und Tarsonemida) aus Africa V. Acta Zoologica Academiae Scientiarum Hungaricae, 21, 39 - 72." type="journal article" year="1975">Mahunka, 1975</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="DFCA57DBA5490E695B97FF3CB17D18BC" box="[686,820,152,178]" pageId="7" pageNumber="188" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="BE976539A5490E6959AEFF1EB35018DD" box="[151,281,186,211]" pageId="7" pageNumber="188" type="description">
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<paragraph id="F63236B2A5490E6959AEFF1EB35018DD" blockId="7.[151,820,152,211]" box="[151,281,186,211]" pageId="7" pageNumber="188">
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(
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<figureCitation id="6EB62A37A5490E6959A6FF1EB35918DD" box="[159,272,186,211]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="7.[151,250,1407,1429]" captionStartId-1="8.[151,250,1116,1138]" captionStartId-2="9.[151,250,1341,1363]" captionStartId-3="10.[151,250,1591,1613]" captionStartId-4="11.[151,250,975,997]" captionTargetBox-0="[167,1418,472,1363]" captionTargetBox-1="[237,1350,313,1046]" captionTargetBox-2="[151,1408,248,1303]" captionTargetBox-3="[167,1418,373,1558]" captionTargetBox-4="[151,1435,193,954]" captionTargetId-0="figure@7.[151,1435,402,1385]" captionTargetId-1="figure@8.[151,1436,292,1111]" captionTargetId-2="figure@9.[151,1435,219,1319]" captionTargetId-3="figure@10.[151,1435,345,1570]" captionTargetId-4="figure@11.[151,1435,193,954]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionText-0="FIGURE 6. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." captionText-1="FIGURE 7. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view." captionText-2="FIGURE 8. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—left leg I in dorsal view, B—left leg II in dorsal view." captionText-3="FIGURE 9. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—left leg III in dorsal view, B—left leg IV in dorsal view." captionText-4="FIGURE 10. DIC micrographs of Troxodania problematica (Mahunka, 1975) comb. nov., female: A—prodorsum, Banterior part of posterior sternal plate, C—claw on tibiotarsus I." httpUri-0="https://zenodo.org/record/2607908/files/figure.png" httpUri-1="https://zenodo.org/record/2607910/files/figure.png" httpUri-2="https://zenodo.org/record/2607912/files/figure.png" httpUri-3="https://zenodo.org/record/2607914/files/figure.png" httpUri-4="https://zenodo.org/record/2607916/files/figure.png" pageId="7" pageNumber="188">Figs 6–10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="BE976539A5490E6959AEFEA6B14A1916" box="[151,771,258,280]" pageId="7" pageNumber="188" type="reference_group">
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<paragraph id="F63236B2A5490E6959AEFEA6B14A1916" blockId="7.[151,771,258,280]" box="[151,771,258,280]" pageId="7" pageNumber="188">
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<taxonomicName id="318D4D31A5490E6959AEFEA6B0361916" authority="Mahunka, 1975, p. 69" authorityName="Mahunka" authorityPageNumber="69" authorityYear="1975" box="[151,639,258,280]" class="Arachnida" family="Pygmephoridae" genus="Bakerdania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="problematica">
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<emphasis id="C4F9EAA0A5490E6959AEFEA6B3D11916" box="[151,408,258,280]" italics="true" pageId="7" pageNumber="188">Bakerdania problematica</emphasis>
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<bibRefCitation id="921C4B43A5490E6958A6FEA6B0091916" author="Mahunka, S." box="[415,576,258,280]" pageId="7" pageNumber="188" pagination="39 - 72" refId="ref7808" refString="Mahunka, S. (1975) Auf Insecten lebende Milben (Acari: Acarida und Tarsonemida) aus Africa V. Acta Zoologica Academiae Scientiarum Hungaricae, 21, 39 - 72." type="journal article" year="1975">Mahunka, 1975</bibRefCitation>
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, p. 69
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</taxonomicName>
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, Figs 20A-F.
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</paragraph>
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</subSubSection>
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<subSubSection id="BE976539A5490E6659AEFEE1B06C1F33" lastPageId="8" lastPageNumber="189" pageId="7" pageNumber="188" type="description">
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<paragraph id="F63236B2A5490E6959AEFEE1B61C1950" blockId="7.[151,1109,325,350]" box="[151,1109,325,350]" pageId="7" pageNumber="188">
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<emphasis id="C4F9EAA0A5490E6959AEFEE1B3741950" bold="true" box="[151,317,325,350]" pageId="7" pageNumber="188">Redescription</emphasis>
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. FEMALE (
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<figureCitation id="6EB62A37A5490E6958F3FEE1B0761950" box="[458,575,325,350]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="7.[151,250,1407,1429]" captionStartId-1="8.[151,250,1116,1138]" captionStartId-2="9.[151,250,1341,1363]" captionStartId-3="10.[151,250,1591,1613]" captionStartId-4="11.[151,250,975,997]" captionTargetBox-0="[167,1418,472,1363]" captionTargetBox-1="[237,1350,313,1046]" captionTargetBox-2="[151,1408,248,1303]" captionTargetBox-3="[167,1418,373,1558]" captionTargetBox-4="[151,1435,193,954]" captionTargetId-0="figure@7.[151,1435,402,1385]" captionTargetId-1="figure@8.[151,1436,292,1111]" captionTargetId-2="figure@9.[151,1435,219,1319]" captionTargetId-3="figure@10.[151,1435,345,1570]" captionTargetId-4="figure@11.[151,1435,193,954]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionText-0="FIGURE 6. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." captionText-1="FIGURE 7. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view." captionText-2="FIGURE 8. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—left leg I in dorsal view, B—left leg II in dorsal view." captionText-3="FIGURE 9. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—left leg III in dorsal view, B—left leg IV in dorsal view." captionText-4="FIGURE 10. DIC micrographs of Troxodania problematica (Mahunka, 1975) comb. nov., female: A—prodorsum, Banterior part of posterior sternal plate, C—claw on tibiotarsus I." httpUri-0="https://zenodo.org/record/2607908/files/figure.png" httpUri-1="https://zenodo.org/record/2607910/files/figure.png" httpUri-2="https://zenodo.org/record/2607912/files/figure.png" httpUri-3="https://zenodo.org/record/2607914/files/figure.png" httpUri-4="https://zenodo.org/record/2607916/files/figure.png" pageId="7" pageNumber="188">Figs 6–10</figureCitation>
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). Length of idiosoma 245–255, width 160–165.
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</paragraph>
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<caption id="A2F2663AA5490E6959AEFADBB36A1DBD" httpUri="https://zenodo.org/record/2607908/files/figure.png" pageId="7" pageNumber="188" startId="7.[151,250,1407,1429]" targetBox="[167,1418,472,1363]" targetPageId="7">
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<paragraph id="F63236B2A5490E6959AEFADBB36A1DBD" blockId="7.[151,1437,1407,1459]" pageId="7" pageNumber="188">
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<emphasis id="C4F9EAA0A5490E6959AEFADBB3471D9A" bold="true" box="[151,270,1407,1429]" pageId="7" pageNumber="188">FIGURE 6</emphasis>
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.
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<taxonomicName id="318D4D31A5490E695822FADBB09D1D9B" authority="(Mahunka, 1975)" authorityName="Khaustov & Frolov" authorityYear="2018" baseAuthorityName="Mahunka" baseAuthorityYear="1975" box="[283,724,1407,1429]" class="Insecta" family="Trogidae" genus="Troxodania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="188" phylum="Arthropoda" rank="species" species="problematica" status="comb. nov.">
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<emphasis id="C4F9EAA0A5490E695822FADBB0551D9B" box="[283,540,1407,1429]" italics="true" pageId="7" pageNumber="188">Troxodania problematica</emphasis>
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(Mahunka, 1975)
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</taxonomicName>
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<emphasis id="C4F9EAA0A5490E695BE2FADBB1191D9A" bold="true" box="[731,848,1407,1428]" pageId="7" pageNumber="188">
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<taxonomicNameLabel id="DFCA57DBA5490E695BE2FADBB1191D9A" box="[731,848,1407,1428]" pageId="7" pageNumber="188" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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, female: A—dorsum of the body, B—venter of the body. Legs omitted.
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</paragraph>
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</caption>
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<paragraph id="F63236B2A5490E6959FEFA44B0B81EFB" blockId="7.[151,1437,1504,2007]" pageId="7" pageNumber="188">
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<emphasis id="C4F9EAA0A5490E6959FEFA44B31F1DF7" box="[199,342,1504,1529]" italics="true" pageId="7" pageNumber="188">Gnathosoma</emphasis>
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(
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<figureCitation id="6EB62A37A5490E69585CFA44B3E31DF7" box="[357,426,1504,1529]" captionStart="FIGURE 7" captionStartId="8.[151,250,1116,1138]" captionTargetBox="[237,1350,313,1046]" captionTargetId="figure@8.[151,1436,292,1111]" captionTargetPageId="8" captionText="FIGURE 7. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view." httpUri="https://zenodo.org/record/2607910/files/figure.png" pageId="7" pageNumber="188">Fig. 7</figureCitation>
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). Gnathosomal capsule almost round in outline, length 23–25, width 24–27. Dorsum with two pairs of distinctly barbed and pointed cheliceral setae
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<emphasis id="C4F9EAA0A5490E695A7AF9A0B1221E13" box="[835,875,1540,1565]" italics="true" pageId="7" pageNumber="188">cha</emphasis>
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12–13 and
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<emphasis id="C4F9EAA0A5490E695AC3F9A0B66A1E13" box="[1018,1059,1540,1565]" italics="true" pageId="7" pageNumber="188">chb</emphasis>
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17–18. Dorsal median apodeme present, weakly developed. Postpalpal setae (
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<emphasis id="C4F9EAA0A5490E695BADF98DB0F91E4E" box="[660,688,1577,1600]" italics="true" pageId="7" pageNumber="188">pp</emphasis>
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) needle-like and blunt-ended, situated anterolaterally to setae
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<emphasis id="C4F9EAA0A5490E695C55F98CB7DD1E4F" box="[1388,1428,1576,1601]" italics="true" pageId="7" pageNumber="188">cha</emphasis>
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. Ventral gnathosoma with one pair of smooth, pointed subcapitular setae
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<emphasis id="C4F9EAA0A5490E695AFBF9E9B19F1E6A" box="[962,982,1613,1636]" italics="true" pageId="7" pageNumber="188">m</emphasis>
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9–10 and round pits situated posteriorly to setae
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<emphasis id="C4F9EAA0A5490E6959C0F9D5B3441E86" box="[249,269,1649,1672]" italics="true" pageId="7" pageNumber="188">m</emphasis>
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. Palps with smooth, pointed setae
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<emphasis id="C4F9EAA0A5490E695B9DF9D4B0991E87" box="[676,720,1648,1673]" italics="true" pageId="7" pageNumber="188">dFe</emphasis>
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and
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<emphasis id="C4F9EAA0A5490E695A35F9D4B1731E87" box="[780,826,1648,1673]" italics="true" pageId="7" pageNumber="188">dGe</emphasis>
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dorsolaterally; setae
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<emphasis id="C4F9EAA0A5490E695D0AF9D4B62B1E87" box="[1075,1122,1648,1673]" italics="true" pageId="7" pageNumber="188">dGe</emphasis>
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distinctly longer than
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<emphasis id="C4F9EAA0A5490E695C50F9D4B7DC1E87" box="[1385,1429,1648,1673]" italics="true" pageId="7" pageNumber="188">dFe</emphasis>
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. Palps ventrally each with mushroom-like
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<emphasis id="C4F9EAA0A5490E695B57F931B0DB1EA2" box="[622,658,1685,1708]" italics="true" pageId="7" pageNumber="188">ass</emphasis>
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and small finger-shaped solenidion laterally to
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<emphasis id="C4F9EAA0A5490E695D94F931B6981EA2" box="[1197,1233,1685,1708]" italics="true" pageId="7" pageNumber="188">ass</emphasis>
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. Palps terminated with small tibial claw and tiny eupathidium-like seta; palpal tibiotarsus usually completely covered by anterior part of femorogenu. Pharyngeal pumps not clearly visible.
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</paragraph>
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<paragraph id="F63236B2A5490E6659FEF8A4B3F018F0" blockId="7.[151,1437,1504,2007]" lastBlockId="8.[151,1436,151,258]" lastPageId="8" lastPageNumber="189" pageId="7" pageNumber="188">
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<emphasis id="C4F9EAA0A5490E6959FEF8A4B3DB1F17" box="[199,402,1792,1817]" italics="true" pageId="7" pageNumber="188">Idiosomal dorsum</emphasis>
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(
|
||
<figureCitation id="6EB62A37A5490E69589BF8A4B0481F17" box="[418,513,1792,1817]" captionStart="FIGURE 6" captionStartId="7.[151,250,1407,1429]" captionTargetBox="[167,1418,472,1363]" captionTargetId="figure@7.[151,1435,402,1385]" captionTargetPageId="7" captionText="FIGURE 6. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." httpUri="https://zenodo.org/record/2607908/files/figure.png" pageId="7" pageNumber="188">Figs 6A</figureCitation>
|
||
,
|
||
<figureCitation id="6EB62A37A5490E695B35F8A5B0091F17" box="[524,576,1793,1817]" captionStart="FIGURE 10" captionStartId="11.[151,250,975,997]" captionTargetBox="[151,1435,193,954]" captionTargetId="figure@11.[151,1435,193,954]" captionTargetPageId="11" captionText="FIGURE 10. DIC micrographs of Troxodania problematica (Mahunka, 1975) comb. nov., female: A—prodorsum, Banterior part of posterior sternal plate, C—claw on tibiotarsus I." httpUri="https://zenodo.org/record/2607916/files/figure.png" pageId="7" pageNumber="188">10A</figureCitation>
|
||
). Only posterior part of prodorsum covered by anterior part of tergite C. All dorsal shields with numerous tiny, barely discernable dimples (
|
||
<figureCitation id="6EB62A37A5490E695A5CF880B1861F33" box="[869,975,1828,1853]" captionStart="FIGURE 10" captionStartId="11.[151,250,975,997]" captionTargetBox="[151,1435,193,954]" captionTargetId="figure@11.[151,1435,193,954]" captionTargetPageId="11" captionText="FIGURE 10. DIC micrographs of Troxodania problematica (Mahunka, 1975) comb. nov., female: A—prodorsum, Banterior part of posterior sternal plate, C—claw on tibiotarsus I." httpUri="https://zenodo.org/record/2607916/files/figure.png" pageId="7" pageNumber="188">Fig. 10A</figureCitation>
|
||
). Prodorsal shield with two pair of setae (
|
||
<date id="82331072A5490E695999F8EDB2FD1F6B" box="[160,180,1865,1893]" pageId="7" pageNumber="188">
|
||
<emphasis id="C4F9EAA0A5490E695999F8EDB2E51F6E" box="[160,172,1865,1888]" italics="true" pageId="7" pageNumber="188">v</emphasis>
|
||
<subScript id="6A0934F7A5490E695995F8F3B2FD1F6B" attach="left" box="[172,180,1879,1893]" fontSize="6" pageId="7" pageNumber="188">2</subScript>
|
||
</date>
|
||
,
|
||
<emphasis id="C4F9EAA0A5490E6959FFF8EDB2941F6E" box="[198,221,1865,1888]" italics="true" pageId="7" pageNumber="188">sc</emphasis>
|
||
<subScript id="6A0934F7A5490E6959E4F8F3B2AC1F6B" attach="left" box="[221,229,1879,1893]" fontSize="6" pageId="7" pageNumber="188">2</subScript>
|
||
), one pair of weakly barbed capitate trichobothria and one pair of round stigmata (
|
||
<figureCitation id="6EB62A37A5490E695D8FF8ECB76B1F6F" box="[1206,1314,1864,1889]" captionStart="FIGURE 10" captionStartId="11.[151,250,975,997]" captionTargetBox="[151,1435,193,954]" captionTargetId="figure@11.[151,1435,193,954]" captionTargetPageId="11" captionText="FIGURE 10. DIC micrographs of Troxodania problematica (Mahunka, 1975) comb. nov., female: A—prodorsum, Banterior part of posterior sternal plate, C—claw on tibiotarsus I." httpUri="https://zenodo.org/record/2607916/files/figure.png" pageId="7" pageNumber="188">Fig. 10A</figureCitation>
|
||
). Stigmata associated with well developed tracheal trunks branched posteriorly. All dorsal setae barbed. Setae
|
||
<emphasis id="C4F9EAA0A5490E695C08F8D4B7741F89" box="[1329,1341,1904,1927]" italics="true" pageId="7" pageNumber="188">c</emphasis>
|
||
<subScript id="6A0934F7A5490E695C07F8DAB70F1F82" attach="left" box="[1342,1350,1918,1932]" fontSize="6" pageId="7" pageNumber="188">2</subScript>
|
||
and
|
||
<emphasis id="C4F9EAA0A5490E695CBFF8CBB7DD1F86" box="[1414,1428,1903,1928]" italics="true" pageId="7" pageNumber="188">h</emphasis>
|
||
<subScript id="6A0934F7A5490E695CADF8DAB7D51F82" attach="left" box="[1428,1436,1918,1932]" fontSize="6" pageId="7" pageNumber="188">2</subScript>
|
||
pointed, other dorsal setae moderately blunt-ended. Cupules
|
||
<emphasis id="C4F9EAA0A5490E695A64F833B13A1FA0" box="[861,883,1943,1966]" italics="true" pageId="7" pageNumber="188">ia</emphasis>
|
||
on tergite D and
|
||
<emphasis id="C4F9EAA0A5490E695D72F831B6281FA0" box="[1099,1121,1941,1966]" italics="true" pageId="7" pageNumber="188">ih</emphasis>
|
||
on tergite H small, round. Posterior margin of tergite H with subtriangular median projection. Setae
|
||
<emphasis id="C4F9EAA0A5490E695AF2F81EB1901FDD" box="[971,985,1978,2003]" italics="true" pageId="7" pageNumber="188">h</emphasis>
|
||
<subScript id="6A0934F7A5490E695AE0F86DB1A81FD9" attach="left" box="[985,993,1993,2007]" fontSize="6" pageId="7" pageNumber="188">2</subScript>
|
||
situated on the same transverse level as
|
||
<emphasis id="C4F9EAA0A5460E6659AEFF33B2EC18BE" box="[151,165,151,176]" italics="true" pageId="8" pageNumber="189">h</emphasis>
|
||
<subScript id="6A0934F7A5460E66599CFF02B2E418BA" attach="left" box="[165,173,166,180]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
. Lengths of dorsal setae:
|
||
<emphasis id="C4F9EAA0A5460E6658F5FF3DB39118BE" box="[460,472,153,176]" italics="true" pageId="8" pageNumber="189">v</emphasis>
|
||
<date id="82331072A5460E6658E1FF02B07918BE" box="[472,560,152,180]" pageId="8" pageNumber="189" value="2013-02-14">
|
||
<subScript id="6A0934F7A5460E6658E1FF02B3A918BA" attach="left" box="[472,480,166,180]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
13–14
|
||
</date>
|
||
,
|
||
<emphasis id="C4F9EAA0A5460E665B02FF3DB01B18BE" box="[571,594,153,176]" italics="true" pageId="8" pageNumber="189">sc</emphasis>
|
||
<subScript id="6A0934F7A5460E665B6BFF02B01318BA" attach="left" box="[594,602,166,180]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
55–60,
|
||
<emphasis id="C4F9EAA0A5460E665B8DFF3DB08918BE" box="[692,704,153,176]" italics="true" pageId="8" pageNumber="189">c</emphasis>
|
||
<subScript id="6A0934F7A5460E665BF8FF02B08018BA" attach="left" box="[705,713,166,180]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
55–61,
|
||
<emphasis id="C4F9EAA0A5460E665A19FF3DB16518BE" box="[800,812,153,176]" italics="true" pageId="8" pageNumber="189">c</emphasis>
|
||
<subScript id="6A0934F7A5460E665A15FF02B17D18BA" attach="left" box="[812,820,166,180]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
70–72,
|
||
<emphasis id="C4F9EAA0A5460E665AB6FF33B1D418BE" box="[911,925,151,176]" italics="true" pageId="8" pageNumber="189">d</emphasis>
|
||
66–69,
|
||
<emphasis id="C4F9EAA0A5460E665AC1FF3DB64D18BE" box="[1016,1028,153,176]" italics="true" pageId="8" pageNumber="189">e</emphasis>
|
||
31–33,
|
||
<emphasis id="C4F9EAA0A5460E665D66FF3CB62E18BE" box="[1119,1127,152,176]" italics="true" pageId="8" pageNumber="189">f</emphasis>
|
||
84–91,
|
||
<emphasis id="C4F9EAA0A5460E665DFBFF33B69918BE" box="[1218,1232,151,176]" italics="true" pageId="8" pageNumber="189">h</emphasis>
|
||
<subScript id="6A0934F7A5460E665DE9FF02B69118BA" attach="left" box="[1232,1240,166,180]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
73–74,
|
||
<emphasis id="C4F9EAA0A5460E665C0BFF33B70918BE" box="[1330,1344,151,176]" italics="true" pageId="8" pageNumber="189">h</emphasis>
|
||
<subScript id="6A0934F7A5460E665C79FF02B70118BA" attach="left" box="[1344,1352,166,180]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
73–74. Distances between setae:
|
||
<date id="82331072A5460E665883FF64B05F18D9" box="[442,534,190,219]" pageId="8" pageNumber="189" value="1959-05-02" valueMax="1959-05-02" valueMin="1959-05-02">
|
||
<emphasis id="C4F9EAA0A5460E665883FF64B38F18D9" box="[442,454,192,215]" italics="true" pageId="8" pageNumber="189">v</emphasis>
|
||
<subScript id="6A0934F7A5460E6658FEFF69B38618D5" attach="right" box="[455,463,205,219]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
<emphasis id="C4F9EAA0A5460E6658F7FF64B3A018D9" box="[462,489,192,215]" italics="true" pageId="8" pageNumber="189">–v</emphasis>
|
||
<subScript id="6A0934F7A5460E6658D0FF69B3B818D5" attach="left" box="[489,497,205,219]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
59
|
||
</date>
|
||
–62,
|
||
<emphasis id="C4F9EAA0A5460E665B76FF64B02F18D9" box="[591,614,192,215]" italics="true" pageId="8" pageNumber="189">sc</emphasis>
|
||
<subScript id="6A0934F7A5460E665B5FFF69B02718D5" attach="left" box="[614,622,205,219]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
<emphasis id="C4F9EAA0A5460E665B57FF64B0DB18D9" box="[622,658,192,215]" italics="true" pageId="8" pageNumber="189">–sc</emphasis>
|
||
<subScript id="6A0934F7A5460E665BAAFF69B0D218D5" attach="left" box="[659,667,205,219]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
51–52,
|
||
<emphasis id="C4F9EAA0A5460E665BC0FF64B14C18D9" box="[761,773,192,215]" italics="true" pageId="8" pageNumber="189">c</emphasis>
|
||
<subScript id="6A0934F7A5460E665A3CFF69B14418D5" attach="left" box="[773,781,205,219]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
–
|
||
<emphasis id="C4F9EAA0A5460E665A22FF64B16E18D9" box="[795,807,192,215]" italics="true" pageId="8" pageNumber="189">c</emphasis>
|
||
<subScript id="6A0934F7A5460E665A1EFF69B16618D5" attach="left" box="[807,815,205,219]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
69–70,
|
||
<emphasis id="C4F9EAA0A5460E665AB4FF64B1D018D9" box="[909,921,192,215]" italics="true" pageId="8" pageNumber="189">c</emphasis>
|
||
<subScript id="6A0934F7A5460E665AA3FF69B1EB18D5" attach="right" box="[922,930,205,219]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
–
|
||
<emphasis id="C4F9EAA0A5460E665A96FF64B1F218D9" box="[943,955,192,215]" italics="true" pageId="8" pageNumber="189">c</emphasis>
|
||
<date id="82331072A5460E665A85FF69B65C18D9" box="[956,1045,191,219]" pageId="8" pageNumber="189" value="1928-02-30">
|
||
<subScript id="6A0934F7A5460E665A85FF69B18D18D5" attach="left" box="[956,964,205,219]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
28–30
|
||
</date>
|
||
,
|
||
<emphasis id="C4F9EAA0A5460E665D18FF1AB60218D9" box="[1057,1099,190,215]" italics="true" pageId="8" pageNumber="189">d–d</emphasis>
|
||
72–78,
|
||
<emphasis id="C4F9EAA0A5460E665D93FF64B68518D9" box="[1194,1228,191,215]" italics="true" pageId="8" pageNumber="189">e–f</emphasis>
|
||
10–11,
|
||
<emphasis id="C4F9EAA0A5460E665C10FF1BB70F18D9" box="[1321,1350,191,215]" italics="true" pageId="8" pageNumber="189">f–f</emphasis>
|
||
86–90,
|
||
<emphasis id="C4F9EAA0A5460E6659AEFF41B2EC18F0" box="[151,165,229,254]" italics="true" pageId="8" pageNumber="189">h</emphasis>
|
||
<subScript id="6A0934F7A5460E66599CFF50B2E4190C" attach="left" box="[165,173,244,258]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
–
|
||
<emphasis id="C4F9EAA0A5460E665982FF41B28018F0" box="[187,201,229,254]" italics="true" pageId="8" pageNumber="189">h</emphasis>
|
||
<subScript id="6A0934F7A5460E6659F1FF50B299190C" attach="left" box="[200,208,244,258]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
40–45,
|
||
<emphasis id="C4F9EAA0A5460E665812FF41B37018F0" box="[299,313,229,254]" italics="true" pageId="8" pageNumber="189">h</emphasis>
|
||
<subScript id="6A0934F7A5460E665800FF50B308190C" attach="left" box="[313,321,244,258]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
–
|
||
<emphasis id="C4F9EAA0A5460E665876FF41B31418F0" box="[335,349,229,254]" italics="true" pageId="8" pageNumber="189">h</emphasis>
|
||
<date id="82331072A5460E665864FF50B3F018F0" box="[349,441,229,258]" pageId="8" pageNumber="189" value="2016-02-17">
|
||
<subScript id="6A0934F7A5460E665864FF50B32C190C" attach="left" box="[349,357,244,258]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
16–17.
|
||
</date>
|
||
</paragraph>
|
||
<caption id="A2F2663AA5460E6659AEFBF8B37D1C9E" httpUri="https://zenodo.org/record/2607910/files/figure.png" pageId="8" pageNumber="189" startId="8.[151,250,1116,1138]" targetBox="[237,1350,313,1046]" targetPageId="8">
|
||
<paragraph id="F63236B2A5460E6659AEFBF8B37D1C9E" blockId="8.[151,1436,1116,1168]" pageId="8" pageNumber="189">
|
||
<emphasis id="C4F9EAA0A5460E6659AEFBF8B3441C7F" bold="true" box="[151,269,1116,1138]" pageId="8" pageNumber="189">FIGURE 7</emphasis>
|
||
.
|
||
<taxonomicName id="318D4D31A5460E665823FBF8B09B1C7C" authority="(Mahunka, 1975)" authorityName="Khaustov & Frolov" authorityYear="2018" baseAuthorityName="Mahunka" baseAuthorityYear="1975" box="[282,722,1116,1138]" class="Insecta" family="Trogidae" genus="Troxodania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="189" phylum="Arthropoda" rank="species" species="problematica" status="comb. nov.">
|
||
<emphasis id="C4F9EAA0A5460E665823FBF8B0521C7C" box="[282,539,1116,1138]" italics="true" pageId="8" pageNumber="189">Troxodania problematica</emphasis>
|
||
(Mahunka, 1975)
|
||
</taxonomicName>
|
||
<emphasis id="C4F9EAA0A5460E665BE0FBF8B1041C7F" bold="true" box="[729,845,1116,1137]" pageId="8" pageNumber="189">
|
||
<taxonomicNameLabel id="DFCA57DBA5460E665BE0FBF8B1041C7F" box="[729,845,1116,1137]" pageId="8" pageNumber="189" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F63236B2A5460E6659FEFB13B7DC1D8A" blockId="8.[151,1437,1207,2033]" pageId="8" pageNumber="189">
|
||
<emphasis id="C4F9EAA0A5460E6659FEFB13B3CC1CDE" box="[199,389,1207,1232]" italics="true" pageId="8" pageNumber="189">Idiosomal venter</emphasis>
|
||
(
|
||
<figureCitation id="6EB62A37A5460E6658AFFB13B3BA1CDE" box="[406,499,1207,1232]" captionStart="FIGURE 6" captionStartId="7.[151,250,1407,1429]" captionTargetBox="[167,1418,472,1363]" captionTargetId="figure@7.[151,1435,402,1385]" captionTargetPageId="7" captionText="FIGURE 6. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." httpUri="https://zenodo.org/record/2607908/files/figure.png" pageId="8" pageNumber="189">Figs 6B</figureCitation>
|
||
,
|
||
<figureCitation id="6EB62A37A5460E665B39FB1CB07B1CDE" box="[512,562,1208,1232]" captionStart="FIGURE 10" captionStartId="11.[151,250,975,997]" captionTargetBox="[151,1435,193,954]" captionTargetId="figure@11.[151,1435,193,954]" captionTargetPageId="11" captionText="FIGURE 10. DIC micrographs of Troxodania problematica (Mahunka, 1975) comb. nov., female: A—prodorsum, Banterior part of posterior sternal plate, C—claw on tibiotarsus I." httpUri="https://zenodo.org/record/2607916/files/figure.png" pageId="8" pageNumber="189">10B</figureCitation>
|
||
). All ventral plates with numerous tiny dimples (
|
||
<figureCitation id="6EB62A37A5460E665D5AFB13B6851CDE" box="[1123,1228,1207,1232]" captionStart="FIGURE 10" captionStartId="11.[151,250,975,997]" captionTargetBox="[151,1435,193,954]" captionTargetId="figure@11.[151,1435,193,954]" captionTargetPageId="11" captionText="FIGURE 10. DIC micrographs of Troxodania problematica (Mahunka, 1975) comb. nov., female: A—prodorsum, Banterior part of posterior sternal plate, C—claw on tibiotarsus I." httpUri="https://zenodo.org/record/2607916/files/figure.png" pageId="8" pageNumber="189">Fig. 10B</figureCitation>
|
||
). All ventral setae barbed and pointed. Ap1 well developed and joined with appr; ap2 vestigial; apsej well developed, thick; ap3 very weak, linear, joined with appo; ap4 well developed, exceeding far beyond bases of setae 3
|
||
<emphasis id="C4F9EAA0A5460E665DA0FB5BB6EE1D16" box="[1177,1191,1279,1304]" italics="true" pageId="8" pageNumber="189">b</emphasis>
|
||
. Apodemes 5 absent. Ags and pgs small, subrectangular; median genital sclerite (mgs) small, oval. Posterior margin of posterior sternal plate almost straight. Posterior margin of aggenital plate rounded. Lengths of ventral setae: 1
|
||
<emphasis id="C4F9EAA0A5460E665D98FAEDB6E61D6E" box="[1185,1199,1353,1376]" italics="true" pageId="8" pageNumber="189">a</emphasis>
|
||
40–50, 1
|
||
<emphasis id="C4F9EAA0A5460E665C21FAE3B76F1D6E" box="[1304,1318,1351,1376]" italics="true" pageId="8" pageNumber="189">b</emphasis>
|
||
32–37, 2
|
||
<emphasis id="C4F9EAA0A5460E665CB7FAEDB7D51D6E" box="[1422,1436,1353,1376]" italics="true" pageId="8" pageNumber="189">a</emphasis>
|
||
43–46, 2
|
||
<emphasis id="C4F9EAA0A5460E6659C0FACFB34E1D8A" box="[249,263,1387,1412]" italics="true" pageId="8" pageNumber="189">b</emphasis>
|
||
24–30, 3
|
||
<emphasis id="C4F9EAA0A5460E665849FAC9B3371D8A" box="[368,382,1389,1412]" italics="true" pageId="8" pageNumber="189">a</emphasis>
|
||
36–38, 3
|
||
<emphasis id="C4F9EAA0A5460E6658DEFACFB3BC1D8A" box="[487,501,1387,1412]" italics="true" pageId="8" pageNumber="189">b</emphasis>
|
||
36–37, 3
|
||
<emphasis id="C4F9EAA0A5460E665B67FAC9B0231D8A" box="[606,618,1389,1412]" italics="true" pageId="8" pageNumber="189">c</emphasis>
|
||
32–34, 4
|
||
<emphasis id="C4F9EAA0A5460E665BEAFAC9B0A81D8A" box="[723,737,1389,1412]" italics="true" pageId="8" pageNumber="189">a</emphasis>
|
||
32–34, 4
|
||
<emphasis id="C4F9EAA0A5460E665A73FACFB1111D8A" box="[842,856,1387,1412]" italics="true" pageId="8" pageNumber="189">b</emphasis>
|
||
44–45, 4
|
||
<emphasis id="C4F9EAA0A5460E665AF8FAC9B1841D8A" box="[961,973,1389,1412]" italics="true" pageId="8" pageNumber="189">c</emphasis>
|
||
38–40,
|
||
<emphasis id="C4F9EAA0A5460E665D11FAC9B6091D8A" box="[1064,1088,1389,1412]" italics="true" pageId="8" pageNumber="189">ps</emphasis>
|
||
<date id="82331072A5460E665D78FADEB6D01D8A" box="[1089,1177,1388,1416]" pageId="8" pageNumber="189" value="1930-01-31">
|
||
<subScript id="6A0934F7A5460E665D78FADEB6001D86" attach="left" box="[1089,1097,1402,1416]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
30–31
|
||
</date>
|
||
,
|
||
<emphasis id="C4F9EAA0A5460E665D9DFAC9B6F41D8A" box="[1188,1213,1389,1412]" italics="true" pageId="8" pageNumber="189">ps</emphasis>
|
||
<date id="82331072A5460E665D84FADEB75D1D8A" box="[1213,1300,1388,1416]" pageId="8" pageNumber="189" value="1932-02-28">
|
||
<subScript id="6A0934F7A5460E665D84FADEB68C1D86" attach="left" box="[1213,1221,1402,1416]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
28–32
|
||
</date>
|
||
,
|
||
<emphasis id="C4F9EAA0A5460E665C19FAC9B7701D8A" box="[1312,1337,1389,1412]" italics="true" pageId="8" pageNumber="189">ps</emphasis>
|
||
<date id="82331072A5460E665C00FADEB7DC1D8A" box="[1337,1429,1387,1416]" pageId="8" pageNumber="189" value="1925-03-28">
|
||
<subScript id="6A0934F7A5460E665C00FADEB7081D86" attach="left" box="[1337,1345,1402,1416]" fontSize="6" pageId="8" pageNumber="189">3</subScript>
|
||
25–28.
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="F63236B2A5460E6659FEFA30B1A11F17" blockId="8.[151,1437,1207,2033]" pageId="8" pageNumber="189">
|
||
<emphasis id="C4F9EAA0A5460E6659FEFA30B2B51DA5" box="[199,252,1428,1451]" italics="true" pageId="8" pageNumber="189">Legs</emphasis>
|
||
(
|
||
<figureCitation id="6EB62A37A5460E665832FA36B31A1DA5" box="[267,339,1426,1451]" captionStart="FIGURE 8" captionStartId="9.[151,250,1341,1363]" captionTargetBox="[151,1408,248,1303]" captionTargetId="figure@9.[151,1435,219,1319]" captionTargetPageId="9" captionText="FIGURE 8. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—left leg I in dorsal view, B—left leg II in dorsal view." httpUri="https://zenodo.org/record/2607912/files/figure.png" pageId="8" pageNumber="189">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="6EB62A37A5460E665866FA37B3261DA5" box="[351,367,1427,1451]" captionStart="FIGURE 9" captionStartId="10.[151,250,1591,1613]" captionTargetBox="[167,1418,373,1558]" captionTargetId="figure@10.[151,1435,345,1570]" captionTargetPageId="10" captionText="FIGURE 9. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—left leg III in dorsal view, B—left leg IV in dorsal view." httpUri="https://zenodo.org/record/2607914/files/figure.png" pageId="8" pageNumber="189">9</figureCitation>
|
||
,
|
||
<figureCitation id="6EB62A37A5460E665845FA37B3E71DA5" box="[380,430,1427,1451]" captionStart="FIGURE 10" captionStartId="11.[151,250,975,997]" captionTargetBox="[151,1435,193,954]" captionTargetId="figure@11.[151,1435,193,954]" captionTargetPageId="11" captionText="FIGURE 10. DIC micrographs of Troxodania problematica (Mahunka, 1975) comb. nov., female: A—prodorsum, Banterior part of posterior sternal plate, C—claw on tibiotarsus I." httpUri="https://zenodo.org/record/2607916/files/figure.png" pageId="8" pageNumber="189">10C</figureCitation>
|
||
). Leg I (
|
||
<figureCitation id="6EB62A37A5460E665B35FA36B0201DA5" box="[524,617,1426,1451]" captionStart="FIGURE 8" captionStartId="9.[151,250,1341,1363]" captionTargetBox="[151,1408,248,1303]" captionTargetId="figure@9.[151,1435,219,1319]" captionTargetPageId="9" captionText="FIGURE 8. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—left leg I in dorsal view, B—left leg II in dorsal view." httpUri="https://zenodo.org/record/2607912/files/figure.png" pageId="8" pageNumber="189">Figs 8A</figureCitation>
|
||
,
|
||
<figureCitation id="6EB62A37A5460E665B4CFA37B0EF1DA5" box="[629,678,1427,1451]" captionStart="FIGURE 10" captionStartId="11.[151,250,975,997]" captionTargetBox="[151,1435,193,954]" captionTargetId="figure@11.[151,1435,193,954]" captionTargetPageId="11" captionText="FIGURE 10. DIC micrographs of Troxodania problematica (Mahunka, 1975) comb. nov., female: A—prodorsum, Banterior part of posterior sternal plate, C—claw on tibiotarsus I." httpUri="https://zenodo.org/record/2607916/files/figure.png" pageId="8" pageNumber="189">10C</figureCitation>
|
||
) distinctly thicker and slightly shorter than leg II. Setal formula:
|
||
<date id="82331072A5460E665CB9FA37B2A61DC1" pageId="8" pageNumber="189" value="2016-03-01" valueMax="2016-04-01" valueMin="2016-03-01">1– 3–4–16</date>
|
||
(4). Tibiotarsus strongly enlarged, with massive blunt-ended claw (
|
||
<figureCitation id="6EB62A37A5460E665AE1FA12B6761DC1" box="[984,1087,1462,1487]" captionStart="FIGURE 10" captionStartId="11.[151,250,975,997]" captionTargetBox="[151,1435,193,954]" captionTargetId="figure@11.[151,1435,193,954]" captionTargetPageId="11" captionText="FIGURE 10. DIC micrographs of Troxodania problematica (Mahunka, 1975) comb. nov., female: A—prodorsum, Banterior part of posterior sternal plate, C—claw on tibiotarsus I." httpUri="https://zenodo.org/record/2607916/files/figure.png" pageId="8" pageNumber="189">Fig. 10C</figureCitation>
|
||
). Seta
|
||
<emphasis id="C4F9EAA0A5460E665DB0FA13B6E41DC0" box="[1161,1197,1463,1486]" italics="true" pageId="8" pageNumber="189">tc”</emphasis>
|
||
situated on short and wide pinnaculum. Solenidion
|
||
<emphasis id="C4F9EAA0A5460E6658D1FA78B3B51DFD" box="[488,508,1500,1523]" italics="true" pageId="8" pageNumber="189">ω</emphasis>
|
||
<subScript id="6A0934F7A5460E6658C5FA4DB04D1DF9" attach="left" box="[508,516,1513,1527]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
thumb-like, other solenidia weakly clavate. Lengths of solenidia
|
||
<emphasis id="C4F9EAA0A5460E665DDAFA78B6BE1DFD" box="[1251,1271,1500,1523]" italics="true" pageId="8" pageNumber="189">ω</emphasis>
|
||
<subScript id="6A0934F7A5460E665DCEFA4DB6B61DF9" attach="left" box="[1271,1279,1513,1527]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
6,
|
||
<emphasis id="C4F9EAA0A5460E665C1BFA78B77F1DFD" box="[1314,1334,1500,1523]" italics="true" pageId="8" pageNumber="189">ω</emphasis>
|
||
<subScript id="6A0934F7A5460E665C0FFA4DB7771DF9" attach="left" box="[1334,1342,1513,1527]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
5,
|
||
<emphasis id="C4F9EAA0A5460E665C58FA78B7391DFD" box="[1377,1392,1500,1523]" italics="true" pageId="8" pageNumber="189">φ</emphasis>
|
||
<subScript id="6A0934F7A5460E665C48FA4DB7301DF9" attach="left" box="[1393,1401,1513,1527]" fontSize="6" pageId="8" pageNumber="189">1</subScript>
|
||
6– 7,
|
||
<emphasis id="C4F9EAA0A5460E66598FF9A6B28C1E17" box="[182,197,1538,1561]" italics="true" pageId="8" pageNumber="189">φ</emphasis>
|
||
<subScript id="6A0934F7A5460E6659FFF9B4B2871E10" attach="left" box="[198,206,1552,1566]" fontSize="6" pageId="8" pageNumber="189">2</subScript>
|
||
4–5. Seta
|
||
<emphasis id="C4F9EAA0A5460E665877F9A5B3151E14" box="[334,348,1537,1562]" italics="true" pageId="8" pageNumber="189">d</emphasis>
|
||
of femur hook-like; seta
|
||
<emphasis id="C4F9EAA0A5460E665BB0F9A5B0D21E14" box="[649,667,1537,1562]" italics="true" pageId="8" pageNumber="189">l’</emphasis>
|
||
of femur blunt-ended and smooth, other setae (except eupathidia) pointed and barbed. Leg II (
|
||
<figureCitation id="6EB62A37A5460E6658E7F98CB0711E4E" box="[478,568,1576,1601]" captionStart="FIGURE 8" captionStartId="9.[151,250,1341,1363]" captionTargetBox="[151,1408,248,1303]" captionTargetId="figure@9.[151,1435,219,1319]" captionTargetPageId="9" captionText="FIGURE 8. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—left leg I in dorsal view, B—left leg II in dorsal view." httpUri="https://zenodo.org/record/2607912/files/figure.png" pageId="8" pageNumber="189">Fig. 8B</figureCitation>
|
||
). Setal formula: 1–3–3–4(1)–6(1). Tarsus with thickened padded claws and flipper-like empodium. Solenidia
|
||
<emphasis id="C4F9EAA0A5460E665B2BF9E9B06F1E6A" box="[530,550,1613,1636]" italics="true" pageId="8" pageNumber="189">ω</emphasis>
|
||
6–7 clavate and
|
||
<emphasis id="C4F9EAA0A5460E665BDDF9E9B0BA1E6A" box="[740,755,1613,1636]" italics="true" pageId="8" pageNumber="189">φ</emphasis>
|
||
3–4 weakly clavate. All setae barbed. Seta
|
||
<emphasis id="C4F9EAA0A5460E665DE5F9E8B6A31E6B" box="[1244,1258,1612,1637]" italics="true" pageId="8" pageNumber="189">d</emphasis>
|
||
of femur bluntended, other setae pointed. Leg III (
|
||
<figureCitation id="6EB62A37A5460E665B1DF9D4B0341E86" box="[548,637,1648,1673]" captionStart="FIGURE 9" captionStartId="10.[151,250,1591,1613]" captionTargetBox="[167,1418,373,1558]" captionTargetId="figure@10.[151,1435,345,1570]" captionTargetPageId="10" captionText="FIGURE 9. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—left leg III in dorsal view, B—left leg IV in dorsal view." httpUri="https://zenodo.org/record/2607914/files/figure.png" pageId="8" pageNumber="189">Fig. 9A</figureCitation>
|
||
). Femur divided into basi- and telofemur. Setal formula: 1–2–2–4(1)–6. Claws and empodium of same shape as on tarsus II. Solenidia
|
||
<emphasis id="C4F9EAA0A5460E665A52F931B1331EA2" box="[875,890,1685,1708]" italics="true" pageId="8" pageNumber="189">φ</emphasis>
|
||
3–4 weakly clavate All setae barbed. Seta
|
||
<emphasis id="C4F9EAA0A5460E665C54F930B7321EA3" box="[1389,1403,1684,1709]" italics="true" pageId="8" pageNumber="189">d</emphasis>
|
||
of femur blunt-ended, other setae pointed. Leg IV (
|
||
<figureCitation id="6EB62A37A5460E665B8AF91CB14C1EDE" box="[691,773,1720,1745]" captionStart="FIGURE 9" captionStartId="10.[151,250,1591,1613]" captionTargetBox="[167,1418,373,1558]" captionTargetId="figure@10.[151,1435,345,1570]" captionTargetPageId="10" captionText="FIGURE 9. Troxodania problematica (Mahunka, 1975) comb. nov., female: A—left leg III in dorsal view, B—left leg IV in dorsal view." httpUri="https://zenodo.org/record/2607914/files/figure.png" pageId="8" pageNumber="189">Fig 9B</figureCitation>
|
||
). Femur divided into basi- and telofemur. Setal formula: 1– 2–1–4(1)–6. Claws simple, hooked, empodium smaller than those on tarsi II and III. Solenidion
|
||
<emphasis id="C4F9EAA0A5460E665DC5F979B7421EFA" box="[1276,1291,1757,1780]" italics="true" pageId="8" pageNumber="189">φ</emphasis>
|
||
2–3 weakly clavate. All setae barbed. Setae
|
||
<emphasis id="C4F9EAA0A5460E6658C5F8A4B0431F17" box="[508,522,1792,1817]" italics="true" pageId="8" pageNumber="189">d</emphasis>
|
||
of femur blunt-ended, other setae pointed.
|
||
</paragraph>
|
||
<paragraph id="F63236B2A5460E6659FEF881B06C1F33" blockId="8.[151,1437,1207,2033]" box="[199,549,1828,1853]" pageId="8" pageNumber="189">MALE and LARVA unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BE976539A5460E6659FEF8ECB0CA1FA7" pageId="8" pageNumber="189" type="materials_examined">
|
||
<paragraph id="F63236B2A5460E6659FEF8ECB0CA1FA7" blockId="8.[151,1437,1207,2033]" pageId="8" pageNumber="189">
|
||
<emphasis id="C4F9EAA0A5460E6659FEF8ECB3E61F6F" bold="true" box="[199,431,1864,1889]" pageId="8" pageNumber="189">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="46E53CEFA5460E6658FDF8ECB7421F6E" ID-GBIF-Occurrence="2028142429" box="[452,1291,1864,1889]" collectingDate="1983-02-07" country="Sri Lanka" location="SRI LANKA" pageId="8" pageNumber="189" specimenCount="3" specimenCount-female="3">
|
||
<collectingCountry id="8E9A7622A5460E6658FDF8ECB02A1F6F" box="[452,611,1864,1889]" name="Sri Lanka" pageId="8" pageNumber="189">SRI LANKA</collectingCountry>
|
||
:
|
||
<specimenCount id="E08BFD3BA5460E665B4AF8ECB0AC1F6F" box="[627,741,1864,1889]" pageId="8" pageNumber="189" type="female">3 females</specimenCount>
|
||
<collectedFrom id="4591754FA5460E665BCAF8EDB6DD1F6E" box="[755,1172,1864,1889]" pageId="8" pageNumber="189">
|
||
on
|
||
<taxonomicName id="318D4D31A5460E665A25F8ECB6DD1F6E" authority=", Kala Oya" authorityName="Kala Oya" box="[796,1172,1864,1889]" class="Insecta" family="Scarabaeidae" genus="Orphnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="189" phylum="Arthropoda" rank="species" species="mysoriensis">
|
||
<emphasis id="C4F9EAA0A5460E665A25F8ECB6461F6E" box="[796,1039,1864,1889]" italics="true" pageId="8" pageNumber="189">Orphnus mysoriensis</emphasis>
|
||
, Kala Oya
|
||
</taxonomicName>
|
||
</collectedFrom>
|
||
<date id="82331072A5460E665D9BF8ECB7421F6E" box="[1186,1291,1864,1888]" pageId="8" pageNumber="189" value="1983-02-07">
|
||
<collectingDate id="9277E99AA5460E665D9BF8ECB7421F6E" box="[1186,1291,1864,1888]" pageId="8" pageNumber="189" value="1983-02-07">7.II.1983</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="46E53CEFA5460E665C24F8EDB1C11F8B" ID-GBIF-Occurrence="2028142426" country="Burkina Faso" location="BURKINA FASO" pageId="8" pageNumber="189" specimenCount="4" specimenCount-female="4">
|
||
<collectingCountry id="8E9A7622A5460E665C24F8EDB2A81F8B" name="Burkina Faso" pageId="8" pageNumber="189">BURKINA FASO</collectingCountry>
|
||
:
|
||
<specimenCount id="E08BFD3BA5460E6659D4F8C9B3151F8B" box="[237,348,1900,1925]" pageId="8" pageNumber="189" type="female">4 females</specimenCount>
|
||
<collectedFrom id="4591754FA5460E66585CF8C9B11E1F8B" box="[357,855,1900,1925]" pageId="8" pageNumber="189">
|
||
on
|
||
<taxonomicName id="318D4D31A5460E6658B2F8C8B11E1F8B" authority="Laporte de Castelnau" authorityName="Laporte de Castelnau" authorityYear="1832" box="[395,855,1900,1925]" class="Insecta" family="Scarabaeidae" genus="Orphnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="189" phylum="Arthropoda" rank="species" species="macleayi">
|
||
<emphasis id="C4F9EAA0A5460E6658B2F8C8B0121F8B" box="[395,603,1900,1925]" italics="true" pageId="8" pageNumber="189">Orphnus macleayi</emphasis>
|
||
Laporte de Castelnau
|
||
</taxonomicName>
|
||
</collectedFrom>
|
||
and
|
||
</materialsCitation>
|
||
<specimenCount id="E08BFD3BA5460E665AABF8C9B6491F8B" box="[914,1024,1900,1925]" pageId="8" pageNumber="189" type="female">2 females</specimenCount>
|
||
on
|
||
<taxonomicName id="318D4D31A5460E665D16F8C8B37E1FA7" authority="Fairmaire, Ouagadougou" authorityName="Fairmaire, Ouagadougou" authorityYear="1898" class="Insecta" family="Scarabaeidae" genus="Orphnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="189" phylum="Arthropoda" rank="species" species="subfoveatus">
|
||
<emphasis id="C4F9EAA0A5460E665D16F8C8B7561F8B" box="[1071,1311,1900,1925]" italics="true" pageId="8" pageNumber="189">Orphnus subfoveatus</emphasis>
|
||
Fairmaire, Ouagadougou
|
||
</taxonomicName>
|
||
,
|
||
<date id="82331072A5460E66587BF834B3FF1FA7" box="[322,438,1936,1961]" pageId="8" pageNumber="189" value="1927-06-06">6.VI.1927</date>
|
||
, G. Olsoufieff leg.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BE976539A5460E6759FEF810B37B18F9" lastPageId="9" lastPageNumber="190" pageId="8" pageNumber="189" type="discussion">
|
||
<paragraph id="F63236B2A5460E6759FEF810B37B18F9" blockId="8.[151,1437,1207,2033]" lastBlockId="9.[151,1437,151,248]" lastPageId="9" lastPageNumber="190" pageId="8" pageNumber="189">
|
||
<emphasis id="C4F9EAA0A5460E6659FEF810B37D1FC3" bold="true" box="[199,308,1972,1997]" pageId="8" pageNumber="189">Remarks</emphasis>
|
||
. We accommodated this species in the genus
|
||
<taxonomicName id="318D4D31A5460E665A68F810B1981FC3" authorityName="Khaustov and Trach" authorityYear="2014" box="[849,977,1972,1997]" class="Insecta" family="Trogidae" genus="Troxodania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C4F9EAA0A5460E665A68F810B1981FC3" box="[849,977,1972,1997]" italics="true" pageId="8" pageNumber="189">Troxodania</emphasis>
|
||
</taxonomicName>
|
||
because of the presence of the median genital sclerite, small anterior and posterior genital sclerites and characteristic shape (strongly thickened, oval) of tibiotarsus I.
|
||
<taxonomicName id="318D4D31A5470E675810FF33B14418BE" authority="(Mahunka, 1975)" authorityName="Khaustov & Frolov" authorityYear="2018" baseAuthorityName="Mahunka" baseAuthorityYear="1975" box="[297,781,151,176]" class="Insecta" family="Trogidae" genus="Troxodania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="190" phylum="Arthropoda" rank="species" species="problematica" status="comb. nov.">
|
||
<emphasis id="C4F9EAA0A5470E675810FF33B00C18BE" box="[297,581,151,176]" italics="true" pageId="9" pageNumber="190">Troxodania problematica</emphasis>
|
||
(
|
||
<bibRefCitation id="921C4B43A5470E675B6BFF33B14C18BE" author="Mahunka, S." box="[594,773,151,176]" pageId="9" pageNumber="190" pagination="39 - 72" refId="ref7808" refString="Mahunka, S. (1975) Auf Insecten lebende Milben (Acari: Acarida und Tarsonemida) aus Africa V. Acta Zoologica Academiae Scientiarum Hungaricae, 21, 39 - 72." type="journal article" year="1975">Mahunka, 1975</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="C4F9EAA0A5470E675A2AFF33B1DD18A1" bold="true" box="[787,916,151,176]" pageId="9" pageNumber="190">
|
||
<taxonomicNameLabel id="DFCA57DBA5470E675A2AFF33B1DD18A1" box="[787,916,151,176]" pageId="9" pageNumber="190" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was described from
|
||
<collectingCountry id="8E9A7622A5470E675D45FF33B68118BE" box="[1148,1224,151,176]" name="Ghana" pageId="9" pageNumber="190">Ghana</collectingCountry>
|
||
, collected on fruits (
|
||
<bibRefCitation id="921C4B43A5470E6759A7FF18B31818DB" author="Mahunka, S." box="[158,337,188,213]" pageId="9" pageNumber="190" pagination="39 - 72" refId="ref7808" refString="Mahunka, S. (1975) Auf Insecten lebende Milben (Acari: Acarida und Tarsonemida) aus Africa V. Acta Zoologica Academiae Scientiarum Hungaricae, 21, 39 - 72." type="journal article" year="1975">Mahunka 1975</bibRefCitation>
|
||
). Examined specimens from Africa and
|
||
<collectingCountry id="8E9A7622A5470E675A09FF18B1EA18DB" box="[816,931,188,213]" name="Sri Lanka" pageId="9" pageNumber="190">Sri Lanka</collectingCountry>
|
||
fit well with the original description of
|
||
<taxonomicName id="318D4D31A5470E675CBEFF19B36218F6" authorityName="Khaustov & Frolov" authorityYear="2018" baseAuthorityName="Mahunka" baseAuthorityYear="1975" class="Insecta" family="Trogidae" genus="Troxodania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="190" phylum="Arthropoda" rank="species" species="problematica">
|
||
<emphasis id="C4F9EAA0A5470E675CBEFF19B36218F6" italics="true" pageId="9" pageNumber="190">T. problematica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |