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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.47.31130" ID-GBIF-Dataset="c1116709-9015-44c4-b306-4149c3f511ff" ID-PMC="PMC6405736" ID-Pensoft-Pub="1314-4049-47-97" ID-PubMed="30858753" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-4049-47-97" ModsDocOrigin="MycoKeys 47" ModsDocTitle="Phylogenetic and morphological studies in Xylodon (Hymenochaetales, Basidiomycota) with the addition of four new species" checkinTime="1555333420989" checkinUser="pensoft" docAuthor="Riebesehl, Janett, Yurchenko, Eugene, Nakasone, Karen K. &amp; Langer, Ewald" docDate="2019" docId="37852A492D309FE57A837209E8E36A36" docLanguage="en" docName="MycoKeys 47: 97-137" docOrigin="MycoKeys 47" docSource="http://dx.doi.org/10.3897/mycokeys.47.31130" docTitle="Xylodon filicinus Yurchenko &amp; Riebesehl, sp. nov." docType="treatment" docVersion="4" lastPageNumber="97" masterDocId="FFC93C6FD904FFF0FFD5FFDDFFC45F71" masterDocTitle="Phylogenetic and morphological studies in Xylodon (Hymenochaetales, Basidiomycota) with the addition of four new species" masterLastPageNumber="137" masterPageNumber="97" pageNumber="97" updateTime="1668136171551" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Phylogenetic and morphological studies in Xylodon (Hymenochaetales, Basidiomycota) with the addition of four new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Riebesehl, Janett</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yurchenko, Eugene</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nakasone, Karen K.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Langer, Ewald</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>47</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>97</mods:start>
<mods:end>137</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.47.31130</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.47.31130</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-47-97</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="156202048" LSID="urn:lsid:plazi:treatment:37852A492D309FE57A837209E8E36A36" httpUri="http://treatment.plazi.org/id/37852A492D309FE57A837209E8E36A36" lastPageNumber="97" pageId="0" pageNumber="97">
<subSubSection pageId="0" pageNumber="97" type="nomenclature">
<paragraph pageId="0" pageNumber="97">
<taxonomicName LSID="MB827463" authority="Yurchenko &amp; Riebesehl" class="Agaricomycetes" family="Schizoporaceae" genus="Xylodon" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Xylodon filicinus" order="Hymenochaetales" pageId="0" pageNumber="97" phylum="Basidiomycota" rank="species" species="filicinus">Xylodon filicinus Yurchenko &amp; Riebesehl</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="97">sp. nov.</taxonomicNameLabel>
Figs 3b, 5
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="holotype">
<paragraph pageId="0" pageNumber="97">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="97">
TAIWAN, Nantou Co., Xitou (Shitou) Forest Recreation Area, W slope of Phoenix Mt. Range, 1470 m a.s.l.,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="23.666666">23°40'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="120.8">120°48'E</geoCoordinate>
, old-growth sparse broadleaf forest, on dead detached rachis of
<taxonomicName class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cyathea" order="Cyatheales" pageId="0" pageNumber="97" phylum="Tracheophyta" rank="genus">Cyathea</taxonomicName>
sp., leg. E. Yurchenko, 31 Jul 2011 (field No. 38; MSK-F 12869; isotype in KAS).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="description">
<paragraph pageId="0" pageNumber="97">Description.</paragraph>
<paragraph pageId="0" pageNumber="97">
Basidiomata effused, white, 2-4 cm in extent, farinaceous or pruinose, very loose or discontinuous, odontioid, 30-55
<normalizedToken originalValue="μm">μm</normalizedToken>
thick between aculei. Margin thinning out. Aculei conical or subcylindrical, 40-80
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 15-45
<normalizedToken originalValue="μm">μm</normalizedToken>
diam., peg-like, of loose texture, 8-14/mm. Hyphal system monomitic, hyphae colourless, clamped at all septa. Subicular hyphae in a loose tissue, rarely branched, 2-3
<normalizedToken originalValue="μm">μm</normalizedToken>
diam., thin- or slightly thick-walled, loosely encrusted, under the subhymenium with inflations 5-6.5
<normalizedToken originalValue="μm">μm</normalizedToken>
wide. The largest crystals in subiculum 6-8
<normalizedToken originalValue="μm">μm</normalizedToken>
across, aggregated in clusters 15-18
<normalizedToken originalValue="μm">μm</normalizedToken>
diam. Subhymenial hyphae moderately branched, partly short-celled and slightly inflated, 2
<normalizedToken originalValue="3.5(">-3.5(-</normalizedToken>
4)
<normalizedToken originalValue="μm">μm</normalizedToken>
diam., lightly encrusted. Projecting hyphae in aculei richly encrusted, 20-45
<normalizedToken originalValue="×">x</normalizedToken>
5-7
<normalizedToken originalValue="μm">μm</normalizedToken>
in encrusted part, with clamped and simple septa, basally thick-walled, then becoming thin-walled, obtuse, sometimes subacute at apex. Cystidia in hymenium thin-walled, lightly encrusted, of three types: (1) subcylindrical, often slightly tapered to apex, numerous, 20-35
<normalizedToken originalValue="×">x</normalizedToken>
4.5-5.5
<normalizedToken originalValue="μm">μm</normalizedToken>
; (2) capitate, rare, 26-32
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 3-5
<normalizedToken originalValue="μm">μm</normalizedToken>
wide at base, 2.5-3
<normalizedToken originalValue="μm">μm</normalizedToken>
wide at apex; (3) hyphoid to narrowly ventricose, about 30
<normalizedToken originalValue="×">x</normalizedToken>
4.5
<normalizedToken originalValue="μm">μm</normalizedToken>
. Basidioles ellipsoid, ovoid, clavate, 7.5-18
<normalizedToken originalValue="×">x</normalizedToken>
4.5-7.5
<normalizedToken originalValue="μm">μm</normalizedToken>
, more or less encrusted. Basidia utriform, (14
<normalizedToken originalValue=")16">-)16-</normalizedToken>
20
<normalizedToken originalValue="×">x</normalizedToken>
(3.5
<normalizedToken originalValue=")4.5">-)4.5-</normalizedToken>
5.5
<normalizedToken originalValue="μm">μm</normalizedToken>
, thin-walled, smooth or sparsely encrusted, with four sterigmata 2-6.5
<normalizedToken originalValue="×">x</normalizedToken>
1-1.5
<normalizedToken originalValue="μm">μm</normalizedToken>
. Spores globose to subglobose, 4
<normalizedToken originalValue="5(">-5(-</normalizedToken>
5.5)
<normalizedToken originalValue="×">x</normalizedToken>
(3.7
<normalizedToken originalValue=")4">-)4-</normalizedToken>
4.5
<normalizedToken originalValue="μm">μm</normalizedToken>
, holotype L = 4.7
<normalizedToken originalValue="µm">µm</normalizedToken>
, W = 4.1
<normalizedToken originalValue="µm">µm</normalizedToken>
, Q = 1.1-1.2, thin-walled, often with one large oil-like globule, negative in Mz, weakly cyanophilous, with minute apiculus.
</paragraph>
<caption pageId="0" pageNumber="97">
<paragraph pageId="0" pageNumber="97">
Figure 5. Micromorphology of
<taxonomicName class="Agaricomycetes" family="Schizoporaceae" genus="Xylodon" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Xylodon filicinus" order="Hymenochaetales" pageId="0" pageNumber="97" phylum="Basidiomycota" rank="species" species="filicinus">Xylodon filicinus</taxonomicName>
(MSK-F 12369, holotype): A vertical section through basidioma B subicular hyphae C inflations on hyphae in lower subhymenium D crystals from subiculum E portions of hymenium and subhymenium F bundle of encrusted projecting hyphae G separate projecting hyphae H subcylindrical cystidia J capitate cystidia K hyphoid cystidium L basidioles M basidia N basidiospores. Scale bars: 100
<normalizedToken originalValue="μm">μm</normalizedToken>
(A); 10
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<normalizedToken originalValue="BM">B-M</normalizedToken>
); 5
<normalizedToken originalValue="μm">μm</normalizedToken>
(N).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="distribution">
<paragraph pageId="0" pageNumber="97">Distribution and ecology.</paragraph>
<paragraph pageId="0" pageNumber="97">From the lower mountainous belt in Taiwan, on dead fern rachises.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="etymology">
<paragraph pageId="0" pageNumber="97">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="97">from Latin filix fern, refers to the occurrence on dead fern rachises.</paragraph>
<paragraph pageId="0" pageNumber="97">Additional specimen examined. TAIWAN, the same locality and the same substrate as holotype, leg. E. Yurchenko, 31 Jul 2011 (field No. 18; MSK-F 12870; dup. in KAS).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="97" type="remarks">
<paragraph pageId="0" pageNumber="97">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="97">
The distinctive features of this species are the pruinose, minutely odontioid basidiomata, fascicles of richly encrusted projecting hyphae in aculei and the three types of cystidia.
<taxonomicName class="Agaricomycetes" family="Schizoporaceae" genus="Xylodon" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Xylodon filicinus" order="Hymenochaetales" pageId="0" pageNumber="97" phylum="Basidiomycota" rank="species" species="filicinus">Xylodon filicinus</taxonomicName>
is morphologically similar to
<taxonomicName lsidName="X. hyphodontinus" pageId="0" pageNumber="97" rank="species" species="hyphodontinus">X. hyphodontinus</taxonomicName>
, which differs in having projecting hyphae in the aculei that are straighter with more septa, a denser subhymenium composed of short-celled hyphae, short, ventricose cystidioles and spore walls that are slightly thickened at maturity (see Fig. 6).
</paragraph>
<caption pageId="0" pageNumber="97">
<paragraph pageId="0" pageNumber="97">
Figure 6. Micromorphology of
<taxonomicName class="Agaricomycetes" family="Schizoporaceae" genus="Xylodon" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Xylodon hyphodontinus" order="Hymenochaetales" pageId="0" pageNumber="97" phylum="Basidiomycota" rank="species" species="hyphodontinus">Xylodon hyphodontinus</taxonomicName>
. LIP GG-MAR 15-127: A vertical section through basidioma B subicular hyphae C excerpt of tramal hyphae to hymenium and projecting hyphae D bundle of encrusted aculeal hyphae E encrustation on projecting hypha in water F encrustation on projecting hyphae in 3% KOHG naked projecting hyphal end H variously shaped hyphal ends J capitate cystidia K portion of hymenium and subhymenium L basidioles and cystidioles M basidiospores. LIP GG-MAR 12-238: N basidia. Scale bars: 100
<normalizedToken originalValue="μm">μm</normalizedToken>
(A); 10
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<normalizedToken originalValue="BL">B-L</normalizedToken>
, N); 5
<normalizedToken originalValue="μm">μm</normalizedToken>
(M).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>