204 lines
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204 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.151.50538" ID-GBIF-Dataset="a62873a2-c4e1-4c6d-9baa-5c5fb94e7348" ID-PMC="PMC7329918" ID-Pensoft-Pub="1314-2003-151-107" ID-Pensoft-UUID="42D1EB2DF0545744B92BFCCCB395C45B" ID-PubMed="32655282" ModsDocID="1314-2003-151-107" checkinTime="1593036331120" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Cuta-Alarcon, Lia E., Bravo-Pedraza, William J. & Soreng, Robert J." docDate="2020" docId="16EBC6BFA25759909B0529CDFEA38C7C" docLanguage="en" docName="PhytoKeys 151: 107-160" docOrigin="PhytoKeys 151" docSource="http://dx.doi.org/10.3897/phytokeys.151.50538" docTitle="Agrostis L. Sp. Pl. 1: 61. 1753" docType="treatment" docVersion="4" id="42D1EB2DF0545744B92BFCCCB395C45B" lastPageNumber="107" masterDocId="42D1EB2DF0545744B92BFCCCB395C45B" masterDocTitle="Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species" masterLastPageNumber="160" masterPageNumber="107" pageNumber="107" updateTime="1668139837814" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sylvester, Steven P.</mods:namePart>
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<mods:affiliation>College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China & Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5577-8782</mods:nameIdentifier>
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<mods:nameIdentifier type="email">steven_sylvester@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cuta-Alarcon, Lia E.</mods:namePart>
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<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bravo-Pedraza, William J.</mods:namePart>
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<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soreng, Robert J.</mods:namePart>
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<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>151</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>107</mods:start>
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<mods:end>160</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.151.50538</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.151.50538</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-151-107</mods:identifier>
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<mods:identifier type="Pensoft-UUID">42D1EB2DF0545744B92BFCCCB395C45B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164730843" LSID="urn:lsid:plazi:treatment:16EBC6BFA25759909B0529CDFEA38C7C" httpUri="http://treatment.plazi.org/id/16EBC6BFA25759909B0529CDFEA38C7C" lastPageNumber="107" pageId="0" pageNumber="107">
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<subSubSection pageId="0" pageNumber="107" type="nomenclature">
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName LSID="16EBC6BF-A257-5990-9B05-29CDFEA38C7C" authority="L. Sp. Pl. 1: 61. 1753" authorityName="L. Sp. Pl. 1: 61." authorityYear="1753" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">Agrostis L. Sp. Pl. 1: 61. 1753</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="reference_group">
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<paragraph pageId="0" pageNumber="107">
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=
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Vilfa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vilfa" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">Vilfa</taxonomicName>
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Adans., Fam. Pl. 2: 495. 1763. Type:
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Vilfa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vilfa stolonifera" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="stolonifera">Vilfa stolonifera</taxonomicName>
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(L.) P. Beauv. (lectotype, designated by
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<bibRefCitation author="Hitchcock, AS" journalOrPublisher="Department of Agriculture Bureau of Plant Industry Bulletin" pageId="0" pageNumber="107" refId="B13" refString="Hitchcock, AS, 1920. The genera of grasses of the United States. United States Department of Agriculture: Bulletin (1915-23) 772: 1-302." title="The genera of grasses of the United States. United States Department of Agriculture: Bulletin (1915 - 23) 772: 1 - 302." year="1920">Hitchcock 1920</bibRefCitation>
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: 127).
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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=
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Trichodium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Trichodium" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">Trichodium</taxonomicName>
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Michx., Fl. Bor.-Amer. (Michaux) 1: 41. 1803. Type:
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Trichodium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Trichodium laxiflorum" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="laxiflorum">Trichodium laxiflorum</taxonomicName>
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Michx. (lectotype, designated by
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<bibRefCitation author="Hitchcock, AS" journalOrPublisher="Department of Agriculture Bureau of Plant Industry Bulletin" pageId="0" pageNumber="107" refId="B13" refString="Hitchcock, AS, 1920. The genera of grasses of the United States. United States Department of Agriculture: Bulletin (1915-23) 772: 1-302." title="The genera of grasses of the United States. United States Department of Agriculture: Bulletin (1915 - 23) 772: 1 - 302." year="1920">Hitchcock 1920</bibRefCitation>
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: 127). Many other heterotypic synonyms.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="materials_examined">
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<paragraph pageId="0" pageNumber="107">Type.</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis stolonifera" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="stolonifera">
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<emphasis italics="true" pageId="0" pageNumber="107">Agrostis stolonifera</emphasis>
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</taxonomicName>
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L. (lectotype, designated by
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<bibRefCitation author="Hitchcock, AS" journalOrPublisher="Department of Agriculture Bureau of Plant Industry Bulletin" pageId="0" pageNumber="107" refId="B13" refString="Hitchcock, AS, 1920. The genera of grasses of the United States. United States Department of Agriculture: Bulletin (1915-23) 772: 1-302." title="The genera of grasses of the United States. United States Department of Agriculture: Bulletin (1915 - 23) 772: 1 - 302." year="1920">Hitchcock 1920</bibRefCitation>
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: 125)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="description">
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<paragraph pageId="0" pageNumber="107">Description.</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<emphasis bold="true" pageId="0" pageNumber="107">Annuals or perennials. Leaves</emphasis>
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basal or cauline;
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<emphasis bold="true" pageId="0" pageNumber="107">ligules</emphasis>
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membranous to scarious.
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<emphasis bold="true" pageId="0" pageNumber="107">Inflorescence</emphasis>
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a panicle, lax and open to contracted and spikelike.
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<emphasis bold="true" pageId="0" pageNumber="107">Spikelets</emphasis>
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1-flowered, disarticulating above the glumes, laterally compressed;
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<emphasis bold="true" pageId="0" pageNumber="107">glumes</emphasis>
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as long as the spikelet, equal or subequal, persisting on the plant after the florets have fallen, usually 1-veined, rarely 3-veined;
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<emphasis bold="true" pageId="0" pageNumber="107">floret</emphasis>
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usually notably shorter than the glumes or reaching to
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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the length of the glumes, exceptionally longer;
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<emphasis bold="true" pageId="0" pageNumber="107">lemmas</emphasis>
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membranaceous or hyaline, generally thinner than the glumes, dorsally rounded, 3- or 5-veined, veins not or distinctly evident;
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<emphasis bold="true" pageId="0" pageNumber="107">paleas</emphasis>
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often absent or noticeably shorter than the lemma, sometimes reaching to
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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the length of the lemma, hyaline and slightly to notably thinner than the lemmas, keels usually obscure, rarely distinct, glabrous, usually smooth, rarely scaberulous;
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<emphasis bold="true" pageId="0" pageNumber="107">calluses</emphasis>
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rounded, glabrous or pubescent and usually with 2 lateral tufts of short hairs;
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<emphasis bold="true" pageId="0" pageNumber="107">rachilla</emphasis>
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prolongation absent.
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<emphasis bold="true" pageId="0" pageNumber="107">Flowers</emphasis>
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perfect;
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<emphasis bold="true" pageId="0" pageNumber="107">anthers</emphasis>
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3 in number, 0.3-1.8 mm long.
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<emphasis bold="true" pageId="0" pageNumber="107">Caryopses</emphasis>
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hard (in species from Colombia) or sometimes with liquid endosperm.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="notes">
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<paragraph pageId="0" pageNumber="107">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="107">
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In Colombian
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<normalizedToken originalValue="páramos">paramos</normalizedToken>
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, taxa of
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="107">Agrostis</emphasis>
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</taxonomicName>
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can be most easily confused with those of
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="107">Calamagrostis</emphasis>
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</taxonomicName>
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s.l. (i.e.,
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<taxonomicName authorityName="Grisebach" authorityYear="1874" class="Liliopsida" family="Poaceae" genus="Cinnagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cinnagrostis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="107">Cinnagrostis</emphasis>
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</taxonomicName>
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Griseb.,
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Deschampsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deschampsia" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="107">Deschampsia</emphasis>
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</taxonomicName>
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P. Beauv.,
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<taxonomicName authorityName="P.M.Peterson, R.J.Soreng, K.Romaschenko & P.Barberá" authorityYear="2019" class="Liliopsida" family="Poaceae" genus="Paramochloa" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Paramochloa" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="107">Paramochloa</emphasis>
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</taxonomicName>
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P.M. Peterson, Soreng, Romasch. &
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<normalizedToken originalValue="Barberá">Barbera</normalizedToken>
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,
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Peyritschia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Peyritschia" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="107">Peyritschia</emphasis>
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</taxonomicName>
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E. Fourn.;
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<bibRefCitation author="Peterson, PM" journalOrPublisher="Phytoneuron" pageId="0" pageNumber="107" pagination="1 - 23" refId="B25" refString="Peterson, PM, Soreng, RJ, Romaschenko, K, Barbera, P, Quintanar, A, Aedo, C, 2019. New combinations and new names in American Cinnagrostis, Peyritschia, and Deschampsia, and three new genera: Greeneochloa, Laegaardia, and Paramochloa (Poeae, Poaceae). Phytoneuron 39: 1 - 23" title="New combinations and new names in American Cinnagrostis, Peyritschia, and Deschampsia, and three new genera: Greeneochloa, Laegaardia, and Paramochloa (Poeae, Poaceae)." volume="39" year="2019">Peterson et al. 2019</bibRefCitation>
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;
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<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.122.33032" author="Sylvester, SP" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="107" pagination="29 - 78" refId="B35" refString="Sylvester, SP, Soreng, RJ, Bravo-Pedraza, WJ, Cuta-Alarcon, LE, Giraldo-Canas, D, Aguilar-Cano, J, Peterson, PM, 2019a. Paramo Calamagrostis s.l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety, and five new records for Colombia. PhytoKeys 122: 29 - 78, DOI: https://doi.org/10.3897/phytokeys.122.33032" title="Paramo Calamagrostis s. l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety, and five new records for Colombia." url="https://doi.org/10.3897/phytokeys.122.33032" volume="122" year="2019 a">Sylvester et al. 2019a</bibRefCitation>
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),
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Podagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Podagrostis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="107">Podagrostis</emphasis>
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</taxonomicName>
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,
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Polypogon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Polypogon" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="107">Polypogon</emphasis>
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</taxonomicName>
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Desf., and
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Sporobolus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sporobolus" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="107">Sporobolus</emphasis>
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</taxonomicName>
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R. Br. The genera previously circumscribed as
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="107">Calamagrostis</emphasis>
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</taxonomicName>
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s.l. (
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<bibRefCitation author="Peterson, PM" journalOrPublisher="Phytoneuron" pageId="0" pageNumber="107" pagination="1 - 23" refId="B25" refString="Peterson, PM, Soreng, RJ, Romaschenko, K, Barbera, P, Quintanar, A, Aedo, C, 2019. New combinations and new names in American Cinnagrostis, Peyritschia, and Deschampsia, and three new genera: Greeneochloa, Laegaardia, and Paramochloa (Poeae, Poaceae). Phytoneuron 39: 1 - 23" title="New combinations and new names in American Cinnagrostis, Peyritschia, and Deschampsia, and three new genera: Greeneochloa, Laegaardia, and Paramochloa (Poeae, Poaceae)." volume="39" year="2019">Peterson et al. 2019</bibRefCitation>
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;
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<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.122.33032" author="Sylvester, SP" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="107" pagination="29 - 78" refId="B35" refString="Sylvester, SP, Soreng, RJ, Bravo-Pedraza, WJ, Cuta-Alarcon, LE, Giraldo-Canas, D, Aguilar-Cano, J, Peterson, PM, 2019a. Paramo Calamagrostis s.l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety, and five new records for Colombia. PhytoKeys 122: 29 - 78, DOI: https://doi.org/10.3897/phytokeys.122.33032" title="Paramo Calamagrostis s. l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety, and five new records for Colombia." url="https://doi.org/10.3897/phytokeys.122.33032" volume="122" year="2019 a">Sylvester et al. 2019a</bibRefCitation>
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) can usually be differentiated by a combination of a prolonged hairy rachilla emerging from the base of the floret, a well-developed palea, a hairy callus, an awn present and inserted dorsally on the lemma, and an upper glume with well-developed lateral veins, although certain species are missing some of these characteristics (see
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<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.122.33032" author="Sylvester, SP" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="107" pagination="29 - 78" refId="B35" refString="Sylvester, SP, Soreng, RJ, Bravo-Pedraza, WJ, Cuta-Alarcon, LE, Giraldo-Canas, D, Aguilar-Cano, J, Peterson, PM, 2019a. Paramo Calamagrostis s.l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety, and five new records for Colombia. PhytoKeys 122: 29 - 78, DOI: https://doi.org/10.3897/phytokeys.122.33032" title="Paramo Calamagrostis s. l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety, and five new records for Colombia." url="https://doi.org/10.3897/phytokeys.122.33032" volume="122" year="2019 a">Sylvester et al. 2019a</bibRefCitation>
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).
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Polypogon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Polypogon" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="107">Polypogon</emphasis>
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</taxonomicName>
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is principally differentiated by spikelets that disarticulate below the glumes, with the grain, lemma, palea, glumes and part of the pedicel falling together. The glumes are also often awned in
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<emphasis italics="true" pageId="0" pageNumber="107">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Polypogon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Polypogon" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">Polypogon</taxonomicName>
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.
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Sporobolus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sporobolus" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">Sporobolus</taxonomicName>
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</emphasis>
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is principally differentiated by its ligule in the form of a line of hairs, its well-developed paleas with the same consistency as the lemma, and the lemmas being 1(-3) veined.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |