345 lines
35 KiB
XML
345 lines
35 KiB
XML
<document id="F323B88BE82F767ACD366A684138BF62" ID="10.11646/zootaxa.4232.3.5SLASH10210" ID-CLB-Dataset="36518" ID-DOI="10.11646/zootaxa.4232.3.5" ID-GBIF-Dataset="8980f1c5-4651-44d4-8df4-5583165b9b2f" ID-ISSN="1175-5326" ID-Zenodo-Dep="312401" ID-ZooBank="277F79DF-4639-4F2D-A60C-107F55D5C037" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487320530468" checkinUser="plazi" docAuthor="Vidlička, Ľubomír, Vršanský, Peter, Kúdelová, Tatiana, Kúdela, Matúš, Deharveng, Louis & Hain, Miroslav" docDate="2017" docId="045A87BDFFBFB20AFF2F3AC3212AFB84" docLanguage="en" docName="zootaxa.4232.3.5SLASH10210.pdf" docOrigin="Zootaxa 4232 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helmablatta louisrothi Vidlicka, Vrsansky et Deharveng, sp. n." docType="treatment" docVersion="8" lastPageNumber="370" masterDocId="F863FFC5FFBBB203FFB83A542554F92C" masterDocTitle="New genus and species of cavernicolous cockroach (Blattaria, Nocticolidae) from Vietnam" masterLastPageNumber="375" masterPageNumber="361" pageNumber="365" updateTime="1698437054477" updateUser="ExternalLinkService">
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<mods:titleInfo id="11F2ABD460E9397F224D2CF459C72953">
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<mods:title id="AADE02FC449279945239D5B1371CBD35">New genus and species of cavernicolous cockroach (Blattaria, Nocticolidae) from Vietnam</mods:title>
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</mods:titleInfo>
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<mods:name id="A2F08D85F1B6781677AC38EB37BC75F4" type="personal">
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<mods:roleTerm id="019FB623ED5FA6EC4338C0287EB572DD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6E46EF1E9D885496C3251C209CF74247">Vidlička, Ľubomír</mods:namePart>
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</mods:name>
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<mods:roleTerm id="3DF9C118F14679C45CE817089BCC83C1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F694D9EDDA19E0204371CAEAA55BC7D4">Vršanský, Peter</mods:namePart>
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</mods:name>
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<mods:name id="B70144D8A453E0E3ECA3764A3AF761A8" type="personal">
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<mods:roleTerm id="DCA2C529EA6FFC519DBAFB4B48F8E8FE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="53035B5B42CD8E18A27BB75E77A26FFC">Kúdelová, Tatiana</mods:namePart>
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</mods:name>
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<mods:name id="F9438D61F55F8DF3E5DFEF7EC8609FF8" type="personal">
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<mods:role id="696BF39FB05E2947FBCFEB359A5EAC7E">
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<mods:roleTerm id="B927DBEC49A8B9B215057731CD4883E2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="479F2EC5FBDFEABF394DC3C7D6090521">Kúdela, Matúš</mods:namePart>
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</mods:name>
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<mods:name id="06538271E759E10AE00B333C30875CCC" type="personal">
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</mods:role>
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<mods:namePart id="8D7974394DBD4814BBEBAB562C55C0EA">Deharveng, Louis</mods:namePart>
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</mods:name>
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<mods:name id="9557D12E3D8DC7FF0C4C51DFE43758B3" type="personal">
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<mods:roleTerm id="104BD6AB12E0F71440C6028E968E24BD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="13D75EC0568B1E281BA70B09976F1830">Hain, Miroslav</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1322295364C6A61C8B557C567D36E63E">text</mods:typeOfResource>
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<mods:title id="C267CFACA4A4A077FA72279D660C0C04">Zootaxa</mods:title>
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<mods:part id="B8A137BC1C107AB638F1DD4CF121A036">
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<mods:date id="D349F3571EBD3713170DDEF88CA56B8C">2017</mods:date>
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<mods:detail id="C54616CC458B7805A06744127667557A" type="volume">
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<mods:number id="3C725EDF3A1B08492F33F06C924D7FA1">4232</mods:number>
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</mods:detail>
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<mods:number id="096ABE263181AC2E50260E2B727FBD10">3</mods:number>
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<mods:classification id="95A5023D4A50221EEA9973980BEF302C">journal article</mods:classification>
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<mods:identifier id="1D50DE2642579E32307AC1C21E4CC0A9" type="CLB-Dataset">36518</mods:identifier>
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<mods:identifier id="6442F52D7A203B8D8FC855E7EE2104E4" type="DOI">10.11646/zootaxa.4232.3.5</mods:identifier>
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<mods:identifier id="39A43767BE655427F4E5904D50ED1E9D" type="GBIF-Dataset">8980f1c5-4651-44d4-8df4-5583165b9b2f</mods:identifier>
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<mods:identifier id="F7CDFE7A0FF6F0B43F1FC54698D77D81" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="38B7AA0D321B587BC2AB068F363A87EF" type="Zenodo-Dep">312401</mods:identifier>
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<mods:identifier id="ED59FD774907199B070296B84FC093AB" type="ZooBank">277F79DF-4639-4F2D-A60C-107F55D5C037</mods:identifier>
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</mods:mods>
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<treatment id="045A87BDFFBFB20AFF2F3AC3212AFB84" ID-DOI="http://doi.org/10.5281/zenodo.6029276" ID-GBIF-Taxon="127696874" ID-Zenodo-Dep="6029276" LSID="urn:lsid:plazi:treatment:045A87BDFFBFB20AFF2F3AC3212AFB84" httpUri="http://treatment.plazi.org/id/045A87BDFFBFB20AFF2F3AC3212AFB84" lastPageId="9" lastPageNumber="370" pageId="4" pageNumber="365">
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<subSubSection id="C4E96520FFBFB207FF2F3AC326CFF99E" box="[151,923,151,178]" pageId="4" pageNumber="365" type="nomenclature">
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<paragraph id="8C4C36ABFFBFB207FF2F3AC326CFF99E" blockId="4.[151,923,151,178]" box="[151,923,151,178]" pageId="4" pageNumber="365">
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<heading id="D70481C7FFBFB207FF2F3AC326CFF99E" bold="true" box="[151,923,151,178]" fontSize="11" level="1" pageId="4" pageNumber="365" reason="1">
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||
<emphasis id="BE87EAB9FFBFB207FF2F3AC326CFF99E" bold="true" box="[151,923,151,178]" pageId="4" pageNumber="365">
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||
<taxonomicName id="4BF34D28FFBFB207FF2F3AC32618F99E" authority="Vidlicka, Vrsansky et Deharveng" authorityName="Vidlicka, Vrsansky et Deharveng" box="[151,844,151,178]" class="Insecta" family="Nocticolidae" genus="Helmablatta" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="365" phylum="Arthropoda" rank="species" species="louisrothi" status="sp. nov.">
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<emphasis id="BE87EAB9FFBFB207FF2F3AC324FEF99D" bold="true" box="[151,426,151,177]" italics="true" pageId="4" pageNumber="365">Helmablatta louisrothi</emphasis>
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Vidlička, Vršanský et Deharveng
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</taxonomicName>
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,
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<taxonomicNameLabel id="A5B457C2FFBFB207FCE13ACC26CFF99E" box="[857,923,152,178]" pageId="4" pageNumber="365" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C4E96520FFBFB207FF2F3A8A268EF84F" pageId="4" pageNumber="365" type="materials_examined">
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||
<paragraph id="8C4C36ABFFBFB207FF2F3A8A21F6F812" blockId="4.[151,1437,222,355]" pageId="4" pageNumber="365">
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<materialsCitation id="3C9B3CF6FFBFB207FF2F3A8A275DF813" ID-GBIF-Occurrence="1434656472" collectorName="Louis Deharveng & Anne Bedos" country="Vietnam" county="Lava" elevation="180" latitude="11.250478" location="Tan Phu area" longLatPrecision="1" longitude="107.40399" municipality="Nai" pageId="4" pageNumber="365" specimenCount="2" typeStatus="holotype">
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||
<emphasis id="BE87EAB9FFBFB207FF2F3A8A2469F9DB" bold="true" box="[151,317,222,247]" pageId="4" pageNumber="365">
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||
<typeStatus id="53488809FFBFB207FF2F3A8A2586F9DB" box="[151,210,222,247]" pageId="4" pageNumber="365">Type</typeStatus>
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||
material
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||
</emphasis>
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||
.
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||
<emphasis id="BE87EAB9FFBFB207FEF33A8A24ECF9DB" bold="true" box="[331,440,222,247]" pageId="4" pageNumber="365">
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||
<typeStatus id="53488809FFBFB207FEF33A8A24ECF9DB" box="[331,440,222,247]" pageId="4" pageNumber="365" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
—male (Vn06 253): southeastern region of
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<collectingCountry id="F4E4763BFFBFB207FC183A8A2157F9DB" box="[928,1027,222,247]" name="Vietnam" pageId="4" pageNumber="365">Vietnam</collectingCountry>
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:
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<location id="892C6070FFBFB207FBB53A8B21F0F9DA" LSID="urn:lsid:plazi:treatment:045A87BDFFBFB20AFF2F3AC3212AFB84:892C6070FFBFB207FBB53A8B21F0F9DA" box="[1037,1188,222,247]" country="Vietnam" county="Lava" latitude="11.250478" longLatPrecision="1" longitude="107.40399" municipality="Nai" name="Tan Phu area" pageId="4" pageNumber="365">Tân Phú area</location>
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,
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||
<location id="892C6070FFBFB207FB163A8A202AF9DB" LSID="urn:lsid:plazi:treatment:045A87BDFFBFB20AFF2F3AC3212AFB84:892C6070FFBFB207FB163A8A202AF9DB" box="[1198,1406,222,247]" country="Vietnam" county="Lava" latitude="11.250478" longLatPrecision="1" longitude="107.40399" municipality="Nai" name="Dinh Quan district" pageId="4" pageNumber="365">Dinh Quan district</location>
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of Đỗng
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<collectingMunicipality id="6C28ACD1FFBFB207FF653B56245CF837" box="[221,264,258,283]" pageId="4" pageNumber="365">Nai</collectingMunicipality>
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||
,
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<collectingCounty id="652D4E27FFBFB207FEAC3B572418F836" box="[276,332,259,282]" pageId="4" pageNumber="365">Lava</collectingCounty>
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||
tube cave (
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||
<geoCoordinate id="E9C7506CFFBFB207FE773B56273CF837" box="[463,616,258,283]" direction="north" orientation="latitude" pageId="4" pageNumber="365" precision="1" value="11.250478">11°15'1.72''N</geoCoordinate>
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||
,
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||
<geoCoordinate id="E9C7506CFFBFB207FDCD3B57267DF837" box="[629,809,259,283]" direction="east" orientation="longitude" pageId="4" pageNumber="365" precision="1" value="107.40399">107°24'14.37''E</geoCoordinate>
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), altitude
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||
<quantity id="4B0B9B4EFFBFB207FC213B5726B7F836" box="[921,995,259,283]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="365" unit="m" value="180.0">
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||
<elevation id="07DED198FFBFB207FC213B5726B7F836" box="[921,995,259,283]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="365" unit="m" value="180.0">180 m</elevation>
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</quantity>
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||
; collected by hand near bat guano, leg.
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<collectorName id="2106537DFFBFB207FF2F3B72240CF813" box="[151,344,294,319]" pageId="4" pageNumber="365">Louis Deharveng</collectorName>
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&
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<collectorName id="2106537DFFBFB207FEC43B73275DF813" box="[380,521,294,319]" pageId="4" pageNumber="365">Anne Bedos</collectorName>
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</materialsCitation>
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||
; coll. Museum National d'Histoire Naturelle, Paris, France.
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||
</paragraph>
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||
<paragraph id="8C4C36ABFFBFB207FF7F3B1F268EF84F" blockId="4.[151,1437,222,355]" box="[199,986,330,355]" pageId="4" pageNumber="365">
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Data deposition in GenBank, under
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||
<taxonomicName id="4BF34D28FFBFB207FDE23B1E27BDF84F" box="[602,745,330,355]" class="Insecta" family="Nocticolidae" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="365" phylum="Arthropoda" rank="family">Nocticolidae</taxonomicName>
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gen. n.—
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<accessionNumber id="93A0AB48FFBFB207FCEE3B1F2682F84F" box="[854,982,331,355]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KR814821" pageId="4" pageNumber="365">KR814821</accessionNumber>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<paragraph id="8C4C36ABFFBFB207FF2F3D6024E4FEAB" pageId="4" pageNumber="365">
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||
<table id="FEF3C40BFFBF4DFCFF2F3D6720C8FEAB" box="[151,1436,1843,1927]" gridcols="3" gridrows="3" pageId="4" pageNumber="365">
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||
<tr id="32C334E9FFBF4DFCFF2F3D6720C8FE66" box="[151,1436,1843,1866]" gridrow="0" pageId="4" pageNumber="365">
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||
<th id="71125D95FFBF4DFCFF2F3D672791FE66" box="[151,709,1843,1866]" gridcol="0" gridrow="0" pageId="4" pageNumber="365">
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||
<emphasis id="BE87EAB9FFBFB207FF2F3D602790FE66" bold="true" box="[151,708,1843,1866]" pageId="4" pageNumber="365">
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FIGURE 1. Male of helmet cockroach
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<emphasis id="BE87EAB9FFBFB207FDFB3D672790FE66" bold="true" box="[579,708,1843,1866]" italics="true" pageId="4" pageNumber="365">Helmablatta</emphasis>
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</emphasis>
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||
</th>
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||
<th id="71125D95FFBF4DFCFD743D67218DFE66" box="[716,1241,1843,1866]" gridcol="1" gridrow="0" pageId="4" pageNumber="365">
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||
<emphasis id="BE87EAB9FFBFB207FD753D672622FE66" bold="true" box="[717,886,1843,1866]" pageId="4" pageNumber="365">
|
||
<taxonomicName id="4BF34D28FFBFB207FD753D672666FE66" box="[717,818,1843,1866]" class="Insecta" family="Nocticolidae" genus="Helmablatta" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="365" phylum="Arthropoda" rank="species" species="louisrothi" status="sp. nov.">
|
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<emphasis id="BE87EAB9FFBFB207FD753D672666FE66" bold="true" box="[717,818,1843,1866]" italics="true" pageId="4" pageNumber="365">louisrothi</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A5B457C2FFBFB207FC833D602622FE66" box="[827,886,1844,1866]" pageId="4" pageNumber="365" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
(Holotype Vn 0 6 253)—drawing.
|
||
</th>
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||
<th id="71125D95FFBF4DFCFB583D6720C8FE66" box="[1248,1436,1843,1866]" gridcol="2" gridrow="0" pageId="4" pageNumber="365">
|
||
<emphasis id="BE87EAB9FFBFB207FB583D6021A0FE66" bold="true" box="[1248,1268,1844,1866]" pageId="4" pageNumber="365">a)</emphasis>
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||
dorsal view;
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||
<emphasis id="BE87EAB9FFBFB207FA3D3D6020C8FE66" bold="true" box="[1413,1436,1844,1866]" pageId="4" pageNumber="365">b)</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="32C334E9FFBF4DFCFF2F3D0720C8FE45" box="[151,1436,1875,1897]" gridrow="1" pageId="4" pageNumber="365">
|
||
<th id="71125D95FFBF4DFCFF2F3D072791FE45" box="[151,709,1875,1897]" gridcol="0" gridrow="1" pageId="4" pageNumber="365">ventral view without left antenna, legs, maxillary palp</th>
|
||
<td id="71125D95FFBF4DFCFD743D07218DFE45" box="[716,1241,1875,1897]" gridcol="1" gridrow="1" pageId="4" pageNumber="365">
|
||
and cercus;
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||
<emphasis id="BE87EAB9FFBFB207FCF03D072608FE45" bold="true" box="[840,860,1875,1897]" pageId="4" pageNumber="365">c)</emphasis>
|
||
view from left side without legs and
|
||
</td>
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||
<td id="71125D95FFBF4DFCFB583D0720C8FE45" box="[1248,1436,1875,1897]" gridcol="2" gridrow="1" pageId="4" pageNumber="365">
|
||
cerci;
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||
<emphasis id="BE87EAB9FFBFB207FA993D07206CFE45" bold="true" box="[1313,1336,1875,1897]" pageId="4" pageNumber="365">d)</emphasis>
|
||
abdomen
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||
</td>
|
||
</tr>
|
||
<tr id="32C334E9FFBF4DFCFF2F3D2520C8FEAB" box="[151,1436,1905,1927]" gridrow="2" pageId="4" pageNumber="365" rowspan-1="1" rowspan-2="1">
|
||
<th id="71125D95FFBF4DFCFF2F3D252791FEAB" box="[151,709,1905,1927]" gridcol="0" gridrow="2" pageId="4" pageNumber="365">without cerci—dorsal view.</th>
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||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="D88C6623FFBEB206FF2F3CEA2766FE3D" httpUri="https://zenodo.org/record/312408/files/figure.png" pageId="5" pageNumber="366" targetBox="[179,1411,193,1699]" targetPageId="5">
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||
<paragraph id="8C4C36ABFFBEB206FF2F3CEA2766FE3D" blockId="5.[151,1436,1725,1809]" pageId="5" pageNumber="366">
|
||
<emphasis id="BE87EAB9FFBEB206FF2F3CEA2630FFF8" bold="true" box="[151,868,1725,1748]" pageId="5" pageNumber="366">
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||
FIGURE 2. Male of helmet cockroach
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<taxonomicName id="4BF34D28FFBEB206FD803CE92670FFF8" box="[568,804,1725,1748]" class="Insecta" family="Nocticolidae" genus="Helmablatta" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="louisrothi" status="sp. nov.">
|
||
<emphasis id="BE87EAB9FFBEB206FD803CE92670FFF8" bold="true" box="[568,804,1725,1748]" italics="true" pageId="5" pageNumber="366">Helmablatta louisrothi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A5B457C2FFBEB206FC933CEA2630FFF8" box="[811,868,1726,1748]" pageId="5" pageNumber="366" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Holotype Vn 0 6 253)—photos.
|
||
<emphasis id="BE87EAB9FFBEB206FB0E3CEA2198FFF8" bold="true" box="[1206,1228,1726,1748]" pageId="5" pageNumber="366">a)</emphasis>
|
||
dorsal view without antennae, legs and cerci;
|
||
<emphasis id="BE87EAB9FFBEB206FE233C8824E6FFDE" bold="true" box="[411,434,1756,1778]" pageId="5" pageNumber="366">b)</emphasis>
|
||
view from left side;
|
||
<emphasis id="BE87EAB9FFBEB206FD343C8827F4FFDE" bold="true" box="[652,672,1756,1778]" pageId="5" pageNumber="366">c)</emphasis>
|
||
view from right side (partly from above);
|
||
<emphasis id="BE87EAB9FFBEB206FBE03C88213BFFDE" bold="true" box="[1112,1135,1756,1778]" pageId="5" pageNumber="366">d)</emphasis>
|
||
head and pronotum—ventral view;
|
||
<emphasis id="BE87EAB9FFBEB206FF6D3CAF25BDFE3D" bold="true" box="[213,233,1787,1809]" pageId="5" pageNumber="366">e)</emphasis>
|
||
head and pronotum—side view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C4E96520FFBEB20AFF7F3D6C25E1FB4C" lastPageId="9" lastPageNumber="370" pageId="5" pageNumber="366" type="description">
|
||
<paragraph id="8C4C36ABFFBEB205FF7F3D6C207DF86A" blockId="5.[151,1436,1848,2020]" lastBlockId="6.[151,1436,151,326]" lastPageId="6" lastPageNumber="367" pageId="5" pageNumber="366">
|
||
<emphasis id="BE87EAB9FFBEB206FF7F3D6C24F0FE7D" bold="true" box="[199,420,1848,1873]" pageId="5" pageNumber="366">Description. Male.</emphasis>
|
||
Body pale, brownish yellow, nearly translucent, only apex of maxillae darker. Head very big (one third of body length), ratio of head length/width = 1.45 (1.167/
|
||
<quantity id="4B0B9B4EFFBEB206FC643D082103FE59" box="[988,1111,1884,1909]" metricMagnitude="-4" metricUnit="m" metricValue="8.05" pageId="5" pageNumber="366" unit="mm" value="0.805">0.805 mm</quantity>
|
||
), oval, invisible from above (opisthognathous at rest); clypeus and labrum short, overlaps mandible only partly; mandibular tip with strongly sclerotized teeth; eyes small (
|
||
<quantity id="4B0B9B4EFFBEB206FDBD3DF02727FE90" box="[517,627,1956,1981]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="366" unit="mm" value="0.15">0.15 mm</quantity>
|
||
long)—about 40-faceted, oblong, narrow, strongly reduced, facets marginally obvious; lateral and central ocelli absent. Maxillary palps very long (length of palpomeres 1st–5th: 0.096, 0.172, 0.310, 0.316,
|
||
<quantity id="4B0B9B4EFFBDB205FEC63ACF24A1F99F" box="[382,501,155,179]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="6" pageNumber="367" unit="mm" value="0.34">0.340 mm</quantity>
|
||
), first and second palpomeres very short, 3th to 5th palpomeres approximately equal, apex of last segment obliquely truncated. Labial palps three-segmented (length of palpomeres 1st–3rd: 0.191, 0.220,
|
||
<quantity id="4B0B9B4EFFBDB205FF2F3AB12459F9D2" box="[151,269,229,254]" metricMagnitude="-4" metricUnit="m" metricValue="3.06" pageId="6" pageNumber="367" unit="mm" value="0.306">0.306 mm</quantity>
|
||
), last segment rounded on apex. Antennae longer than body (preserved part
|
||
<quantity id="4B0B9B4EFFBDB205FBDE3AB121E8F9D1" box="[1126,1212,229,254]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="6" pageNumber="367" unit="mm" value="5.1">5.1 mm</quantity>
|
||
long, scapus 0.145/
|
||
<quantity id="4B0B9B4EFFBDB205FF2F3B5D2458F80E" box="[151,268,265,290]" metricMagnitude="-5" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="367" unit="mm" value="0.085">0.085 mm</quantity>
|
||
), slender, complete number of antennomeres is unknown, only 47 segments remained, scape and pedicel moderate elongated, following 10 segments as long as wide, other segments two times longer than wide.
|
||
</paragraph>
|
||
<caption id="D88C6623FFBDB205FF2F3D2025A3FEC9" httpUri="https://zenodo.org/record/312413/files/figure.png" pageId="6" pageNumber="367" targetBox="[179,1417,368,1860]" targetPageId="6">
|
||
<paragraph id="8C4C36ABFFBDB205FF2F3D2025A3FEC9" blockId="6.[151,1436,1907,2021]" pageId="6" pageNumber="367">
|
||
<emphasis id="BE87EAB9FFBDB205FF2F3D2026DBFEA6" bold="true" box="[151,911,1907,1930]" pageId="6" pageNumber="367">
|
||
FIGURE 3. Male of helmet cockroach
|
||
<taxonomicName id="4BF34D28FFBDB205FDEC3D272612FEA6" box="[596,838,1907,1930]" class="Insecta" family="Nocticolidae" genus="Helmablatta" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="367" phylum="Arthropoda" rank="species" species="louisrothi" status="sp. nov.">
|
||
<emphasis id="BE87EAB9FFBDB205FDEC3D272612FEA6" bold="true" box="[596,838,1907,1930]" italics="true" pageId="6" pageNumber="367">Helmablatta louisrothi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A5B457C2FFBDB205FCE93D2026DBFEA6" box="[849,911,1908,1930]" pageId="6" pageNumber="367" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Holotype Vn 0 6 253)—photos and drawing.
|
||
<emphasis id="BE87EAB9FFBDB205FA3F3D2020C8FEA6" bold="true" box="[1415,1436,1908,1930]" pageId="6" pageNumber="367">a)</emphasis>
|
||
foreleg—anterior and posterior views;
|
||
<emphasis id="BE87EAB9FFBDB205FD9F3DC6276AFE84" bold="true" box="[551,574,1938,1960]" pageId="6" pageNumber="367">b)</emphasis>
|
||
midleg—anterior and posterior views;
|
||
<emphasis id="BE87EAB9FFBDB205FC6D3DC626BDFE84" bold="true" box="[981,1001,1938,1960]" pageId="6" pageNumber="367">c)</emphasis>
|
||
hindleg—anterior and posterior views;
|
||
<emphasis id="BE87EAB9FFBDB205FA3D3DC620C8FE84" bold="true" box="[1413,1436,1938,1960]" pageId="6" pageNumber="367">d)</emphasis>
|
||
complex of tergal gland—oblique from above view;
|
||
<emphasis id="BE87EAB9FFBDB205FD7D3DE5278DFEEB" bold="true" box="[709,729,1969,1991]" pageId="6" pageNumber="367">e)</emphasis>
|
||
right tegmen;
|
||
<emphasis id="BE87EAB9FFBDB205FCC03DE526DDFEEB" bold="true" box="[888,905,1969,1991]" pageId="6" pageNumber="367">f)</emphasis>
|
||
right wing. Abbreviations, ante = anterior, post = posterior.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D88C6623FFBCB204FF2F3C9C2767FE37" httpUri="https://zenodo.org/record/312420/files/figure.png" pageId="7" pageNumber="368" targetBox="[151,1436,193,1714]" targetPageId="7">
|
||
<paragraph id="8C4C36ABFFBCB204FF2F3C9C2767FE37" blockId="7.[151,1436,1735,1819]" pageId="7" pageNumber="368">
|
||
<emphasis id="BE87EAB9FFBCB204FF2F3C9C2637FFF2" bold="true" box="[151,867,1735,1758]" pageId="7" pageNumber="368">
|
||
FIGURE 4. Male of helmet cockroach
|
||
<taxonomicName id="4BF34D28FFBCB204FD8E3C932677FFF2" box="[566,803,1735,1758]" class="Insecta" family="Nocticolidae" genus="Helmablatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="368" phylum="Arthropoda" rank="species" species="louisrothi" status="sp. nov.">
|
||
<emphasis id="BE87EAB9FFBCB204FD8E3C932677FFF2" bold="true" box="[566,803,1735,1758]" italics="true" pageId="7" pageNumber="368">Helmablatta louisrothi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A5B457C2FFBCB204FC923C9C2637FFF2" box="[810,867,1736,1758]" pageId="7" pageNumber="368" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Holotype Vn 0 6 253)—electron microscope photos.
|
||
<emphasis id="BE87EAB9FFBCB204FA3F3C9C20C8FFF2" bold="true" box="[1415,1436,1736,1758]" pageId="7" pageNumber="368">a)</emphasis>
|
||
body without appendages—dorsal view;
|
||
<emphasis id="BE87EAB9FFBCB204FDFB3CB2270DFFD0" bold="true" box="[579,601,1766,1788]" pageId="7" pageNumber="368">b)</emphasis>
|
||
tergal gland and “nipper”—dorsal view;
|
||
<emphasis id="BE87EAB9FFBCB204FBB73CB22177FFD0" bold="true" box="[1039,1059,1766,1788]" pageId="7" pageNumber="368">c)</emphasis>
|
||
tergal gland and “nipper”—oblique from above view;
|
||
<emphasis id="BE87EAB9FFBCB204FEE83D512433FE37" bold="true" box="[336,359,1797,1819]" pageId="7" pageNumber="368">d)</emphasis>
|
||
hook—dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8C4C36ABFFBCB20BFF7F3D17214EF9F9" blockId="7.[151,1437,1858,2030]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="369" pageId="7" pageNumber="368">
|
||
Pronotum parabolic, lateral hind corner slightly oblong (width/length = 1.360/
|
||
<quantity id="4B0B9B4EFFBCB204FBEC3D16219AFE77" box="[1108,1230,1858,1883]" metricMagnitude="-4" metricUnit="m" metricValue="9.64" pageId="7" pageNumber="368" unit="mm" value="0.964">0.964 mm</quantity>
|
||
). Tegmina do not touch, shortened (
|
||
<quantity id="4B0B9B4EFFBCB204FED93D3E2481FEAD" box="[353,469,1898,1922]" metricMagnitude="-3" metricUnit="m" metricValue="1.944" pageId="7" pageNumber="368" unit="mm" value="1.944">1.944 mm</quantity>
|
||
long), reaching to posterior margin of 6th abdominal segment, in apical third strongly tapered (width in middle 0.655 and in apical third
|
||
<quantity id="4B0B9B4EFFBCB204FD693DD92611FE8A" box="[721,837,1933,1958]" metricMagnitude="-4" metricUnit="m" metricValue="3.61" pageId="7" pageNumber="368" unit="mm" value="0.361">0.361 mm</quantity>
|
||
) and sinuous on inner side, uncovering (outflanking) the tergal gland, apex volute internal, venation invisible with the exception of indistinct Sc vein, upper surface and inner margin of sinuous part covered by setae, basis with a few spines (
|
||
<figureCitation id="14C82A2EFFBCB204FC033D81215EFEC2" box="[955,1034,2005,2030]" captionStart="FIGURE 2" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[179,1411,193,1699]" captionTargetId="figure@5.[151,1436,171,1705]" captionTargetPageId="5" captionText="FIGURE 2. Male of helmet cockroach Helmablatta louisrothi sp. n. (Holotype Vn 0 6 253) — photos. a) dorsal view without antennae, legs and cerci; b) view from left side; c) view from right side (partly from above); d) head and pronotum—ventral view; e) head and pronotum—side view." httpUri="https://zenodo.org/record/312408/files/figure.png" pageId="7" pageNumber="368">Figs. 2</figureCitation>
|
||
e, 3e). Hind wings strongly reduced (0.990/
|
||
<quantity id="4B0B9B4EFFB3B20BFF5F3ACC240BF99C" box="[231,351,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="4.28" pageId="8" pageNumber="369" unit="mm" value="0.428">0.428 mm</quantity>
|
||
), widest in apical third, without visible venation, with unique very deep incision in apical part, upper surface with very sparse and small setae, basis with a few spines (
|
||
<figureCitation id="14C82A2EFFB3B20BFC073AE82155F9F9" box="[959,1025,188,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1908,1930]" captionTargetBox="[179,1417,368,1860]" captionTargetId="figure@6.[151,1436,352,1885]" captionTargetPageId="6" captionText="FIGURE 3. Male of helmet cockroach Helmablatta louisrothi sp. n. (Holotype Vn 0 6 253) — photos and drawing. a) foreleg—anterior and posterior views; b) midleg—anterior and posterior views; c) hindleg—anterior and posterior views; d) complex of tergal gland—oblique from above view; e) right tegmen; f) right wing. Abbreviations, ante = anterior, post = posterior." httpUri="https://zenodo.org/record/312413/files/figure.png" pageId="8" pageNumber="369">Fig. 3</figureCitation>
|
||
f).
|
||
</paragraph>
|
||
<caption id="D88C6623FFB3B20BFF2F3C2C2642FFC6" httpUri="https://zenodo.org/record/312424/files/figure.png" pageId="8" pageNumber="369" targetBox="[194,1417,296,1570]" targetPageId="8">
|
||
<paragraph id="8C4C36ABFFB3B20BFF2F3C2C2642FFC6" blockId="8.[151,1436,1655,1770]" pageId="8" pageNumber="369">
|
||
<emphasis id="BE87EAB9FFB3B20BFF2F3C2C2659FFA2" bold="true" box="[151,781,1655,1678]" pageId="8" pageNumber="369">
|
||
FIGURE 5. Helmet cockroach
|
||
<taxonomicName id="4BF34D28FFB3B20BFE5A3C23279AFFA2" box="[482,718,1655,1678]" class="Insecta" family="Nocticolidae" genus="Helmablatta" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="369" phylum="Arthropoda" rank="species" species="louisrothi" status="sp. nov.">
|
||
<emphasis id="BE87EAB9FFB3B20BFE5A3C23279AFFA2" bold="true" box="[482,718,1655,1678]" italics="true" pageId="8" pageNumber="369">Helmablatta louisrothi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A5B457C2FFB3B20BFD6C3C2C2659FFA2" box="[724,781,1656,1678]" pageId="8" pageNumber="369" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
—CT 3D representation.
|
||
<emphasis id="BE87EAB9FFB3B20BFBB53C2C216EFFA2" bold="true" box="[1037,1082,1656,1678]" pageId="8" pageNumber="369">a–c)</emphasis>
|
||
male body without appendages:
|
||
<emphasis id="BE87EAB9FFB3B20BFA3F3C2C20C8FFA2" bold="true" box="[1415,1436,1656,1678]" pageId="8" pageNumber="369">a)</emphasis>
|
||
ventral view;
|
||
<emphasis id="BE87EAB9FFB3B20BFE9A3CC3246DFF81" bold="true" box="[290,313,1687,1709]" pageId="8" pageNumber="369">b)</emphasis>
|
||
dorsal view;
|
||
<emphasis id="BE87EAB9FFB3B20BFE793CC32481FF81" bold="true" box="[449,469,1687,1709]" pageId="8" pageNumber="369">c)</emphasis>
|
||
views from right and left side;
|
||
<emphasis id="BE87EAB9FFB3B20BFCAB3CC32615FF81" bold="true" box="[787,833,1687,1709]" pageId="8" pageNumber="369">d–e)</emphasis>
|
||
hypothetical reconstruction of body position during female feeding on the male's tergal gland secretion:
|
||
<emphasis id="BE87EAB9FFB3B20BFDCD3CE127D8FFE7" bold="true" box="[629,652,1717,1739]" pageId="8" pageNumber="369">d)</emphasis>
|
||
drawing reconstruction,
|
||
<emphasis id="BE87EAB9FFB3B20BFC2C3CE126CBFFE7" bold="true" box="[916,927,1717,1739]" pageId="8" pageNumber="369">e</emphasis>
|
||
) CT 3D reconstruction;
|
||
<emphasis id="BE87EAB9FFB3B20BFB103CE121E4FFE7" bold="true" box="[1192,1200,1717,1739]" pageId="8" pageNumber="369">f</emphasis>
|
||
) CT representation of digestive tract visualising food particles (fungal spores or soil).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8C4C36ABFFB3B20AFF7F3D42201EF9D4" blockId="8.[151,1437,1814,2022]" lastBlockId="9.[151,1437,151,680]" lastPageId="9" lastPageNumber="370" pageId="8" pageNumber="369">
|
||
All legs, in proportion to body, very large. Procoxae elongated, tapered, meso- and metacoxae triangular, robust, proximally with a few small spines; femora on anterior side covered with small spines, genicular spines of femora present, length of fore-, mid- and hind femora 1.214, 1.275 and
|
||
<quantity id="4B0B9B4EFFB3B20BFC5F3D0A210BFE5B" box="[999,1119,1886,1911]" metricMagnitude="-3" metricUnit="m" metricValue="1.579" pageId="8" pageNumber="369" unit="mm" value="1.579">1.579 mm</quantity>
|
||
, respectively; anteroventral margin of fore femur with alternate large and small spines irregularly disposed, and a large distal spine (intermediate
|
||
<typeStatus id="53488809FFB3B20BFE8D3DFC2439FEEC" box="[309,365,1960,1984]" pageId="8" pageNumber="369">Type</typeStatus>
|
||
A1
|
||
<emphasis id="BE87EAB9FFB3B20BFE2B3DF224F5FE93" bold="true" box="[403,417,1958,1983]" pageId="8" pageNumber="369">–</emphasis>
|
||
B1—terminology follow
|
||
<bibRefCitation id="E8624B5AFFB3B20BFD043DF2266DFE93" author="Roth" box="[700,825,1958,1983]" pageId="8" pageNumber="369" refString="Roth, L. M. (2003) Systematics and phylogeny of cockroaches (Dictyoptera: Blattaria). Oriental Insects, 37, 1 - 186. https: // doi. org / 10.1080 / 00305316.2003.10417344" type="journal article" year="2003">Roth 2003</bibRefCitation>
|
||
); tibia with several small spines, distal end with four large spines, length of fore-, mid- and hind tibia 0.801, 1.020 and
|
||
<quantity id="4B0B9B4EFFB3B20BFC313D9926ABFEC9" box="[905,1023,1997,2022]" metricMagnitude="-3" metricUnit="m" metricValue="1.603" pageId="8" pageNumber="369" unit="mm" value="1.603">1.603 mm</quantity>
|
||
, respectively; tarsomeres with small spines in three rows (
|
||
<figureCitation id="14C82A2EFFB2B20AFE323AC3248DF99C" box="[394,473,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,250,1908,1930]" captionTargetBox="[179,1417,368,1860]" captionTargetId="figure@6.[151,1436,352,1885]" captionTargetPageId="6" captionText="FIGURE 3. Male of helmet cockroach Helmablatta louisrothi sp. n. (Holotype Vn 0 6 253) — photos and drawing. a) foreleg—anterior and posterior views; b) midleg—anterior and posterior views; c) hindleg—anterior and posterior views; d) complex of tergal gland—oblique from above view; e) right tegmen; f) right wing. Abbreviations, ante = anterior, post = posterior." httpUri="https://zenodo.org/record/312413/files/figure.png" pageId="9" pageNumber="370">Figs. 3</figureCitation>
|
||
a, b, c); tarsomere 1
|
||
<emphasis id="BE87EAB9FFB2B20AFD043AC3279EF99C" bold="true" box="[700,714,151,176]" pageId="9" pageNumber="370">–</emphasis>
|
||
5 length: (front-, mid- and hind-leg): (0.607, 0.777,
|
||
<quantity id="4B0B9B4EFFB2B20AFAA13ACC20C4F99C" box="[1305,1424,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="9.23" pageId="9" pageNumber="370" unit="mm" value="0.923">0.923 mm</quantity>
|
||
), (0.218, 0.231, 0.316), (0.085, 0.121, 0.151), (0.073, 0.085, 0.090), (0.170, 0.170, 0.180); tarsal claws simple, symmetrical; pulvilli and arolia absent; basitarsi of all legs longer than the four other tarsomeres combined.
|
||
</paragraph>
|
||
<paragraph id="8C4C36ABFFB2B20AFF7F3B502015F881" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="370">
|
||
Abdomen very short (1.756/
|
||
<quantity id="4B0B9B4EFFB2B20AFDAC3B5027D9F830" box="[532,653,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" pageId="9" pageNumber="370" unit="mm" value="1.09">1.090 mm</quantity>
|
||
) with very distinct tergal gland. Abdominal terga T3–T5 strongly specialized (Figs. 1a, 2a, b, c, 3d, 4a, b, c), together form very sophisticated and unique tergal gland equipment, where T4 is extremely extruded over surface of abdomen; abdominal tergite T8 create in the middle big anteriorly incurved hook (“nipper”)—extremely rigid part of the insect, interfere with cavity on T7; cerci (incomplete) with very long segments in posterior part and long sensilla on the apex, surface covered with setae (Figs. 1a, b).
|
||
</paragraph>
|
||
<paragraph id="8C4C36ABFFB2B20AFF7F3BE325E1FB4C" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="370">
|
||
Supra-anal plate wide, subtrapezoidal, with central concave hind margin (Figs. 1a, d, 2a, 4a, b, d). Sub-genital plate symmetrical, at the sides with two similar very short styles, inter-stylar margin convexly rounded (Fig. 1b). Genital hook (L3) projects (
|
||
<quantity id="4B0B9B4EFFB2B20AFE6938542711FB3B" box="[465,581,512,536]" metricMagnitude="-4" metricUnit="m" metricValue="3.88" pageId="9" pageNumber="370" unit="mm" value="0.388">0.388 mm</quantity>
|
||
long) on the left side, subapicali wide (
|
||
<quantity id="4B0B9B4EFFB2B20AFBBA3BAB2123FB34" box="[1026,1143,511,536]" metricMagnitude="-5" metricUnit="m" metricValue="6.2" pageId="9" pageNumber="370" unit="mm" value="0.062">0.062 mm</quantity>
|
||
) with several strong setae in inner margin, the apex of hook strongly narrow (
|
||
<quantity id="4B0B9B4EFFB2B20AFD6038702604FB10" box="[728,848,548,572]" metricMagnitude="-5" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="370" unit="mm" value="0.021">0.021 mm</quantity>
|
||
), long (
|
||
<quantity id="4B0B9B4EFFB2B20AFC1C3870214DFB10" box="[932,1049,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="1.28" pageId="9" pageNumber="370" unit="mm" value="0.128">0.128 mm</quantity>
|
||
) and rectangular curved (
|
||
<figureCitation id="14C82A2EFFB2B20AFA82387020DCFB11" box="[1338,1416,548,573]" captionStart="FIGURE 4" captionStartId="7.[151,250,1736,1758]" captionTargetBox="[151,1436,193,1714]" captionTargetId="figure@7.[151,1436,193,1714]" captionTargetPageId="7" captionText="FIGURE 4. Male of helmet cockroach Helmablatta louisrothi sp. n. (Holotype Vn 0 6 253) — electron microscope photos. a) body without appendages—dorsal view; b) tergal gland and “ nipper ” — dorsal view; c) tergal gland and “ nipper ” — oblique from above view; d) hook—dorsal view." httpUri="https://zenodo.org/record/312420/files/figure.png" pageId="9" pageNumber="370">Figs. 4</figureCitation>
|
||
b, d).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4E96520FFB2B20AFF7F3838212AFB84" pageId="9" pageNumber="370" type="etymology">
|
||
<paragraph id="8C4C36ABFFB2B20AFF7F3838212AFB84" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="370">
|
||
<emphasis id="BE87EAB9FFB2B20AFF7F3838241AFBA9" bold="true" box="[199,334,620,645]" pageId="9" pageNumber="370">Etymology.</emphasis>
|
||
New cockroach species is named in honour of Louis M. Roth (*1918
|
||
<emphasis id="BE87EAB9FFB2B20AFBD83838213AFBA9" bold="true" box="[1120,1134,620,645]" pageId="9" pageNumber="370">–</emphasis>
|
||
†2013), one of the greatest “cockroachologist” of the 20th century who published the latest revision of
|
||
<taxonomicName id="4BF34D28FFB2B20AFC5F38DB212EFB84" box="[999,1146,655,680]" class="Insecta" family="Nocticolidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="370" phylum="Arthropoda" rank="family">Nocticolidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |