292 lines
46 KiB
XML
292 lines
46 KiB
XML
<document id="C6D3364B596DB51484F80D3FA2302B28" ID-DOI="10.37520/aemnp.2023.024" ID-ISSN="1804-6487" ID-Zenodo-Dep="10621434" ID-ZooBank="B94E1D89-5827-404D-821F-2DA8339FFB58" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707165741765" checkinUser="felipe" docAuthor="Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A. & Rafael, José Albertino" docDate="2023" docId="C00B8781CC03F1535AFA58EE2532151F" docLanguage="en" docName="ActaEntMusNatPra.63.2.383-395.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 63 (2)" docSource="http://dx.doi.org/10.37520/aemnp.2023.024" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Chalarus boyacensis Ramos-Pastrana & Marques & Rafael 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="388" masterDocId="3C32FFF9CC01F156593658522770174A" masterDocTitle="Chalarus (Diptera: Pipunculidae) of Colombia, with description of two new species and an updated key to males of the Neotropical species" masterLastPageNumber="395" masterPageNumber="383" pageNumber="385" updateTime="1707166334007" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:titleInfo id="EA9CE0884D384C7AD19293D783DD9FA0">
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<mods:title id="A19BE99DD30DA89397B47A50F1F8FCA4">Chalarus (Diptera: Pipunculidae) of Colombia, with description of two new species and an updated key to males of the Neotropical species</mods:title>
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<mods:name id="E660101B2E6F0DBEFF396C86C7EF39B0" type="personal">
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<mods:roleTerm id="DF101A8E6750F20597DF3D87C7B324FE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5BF737DA7D0813A3998653079FDCF5BF">Ramos-Pastrana, Yardany</mods:namePart>
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<mods:affiliation id="1A54AE2E12A52FEAEA08794AD2992CBC">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEUA-, Laboratorio de Entomología, - LEUA-, Florencia, Caquetá, Colombia: e-mail: ya. ramos @ udla. edu. co; ORCID: 0000 - 0002 - 3193 - 6659 & Instituto Nacional de Pesquisas da Amazônia, INPA, Manaus, Amazonas, Brazil; e-mails and ORCIDs: willkenia @ gmail. com, 0000 - 0001 - 7260 - 5760; jarafael @ inpa. gov. br, 0000 - 0002 - 0170 - 0514</mods:affiliation>
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<mods:roleTerm id="442A80D86EDECA53748C3EA80A7DA25A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="45CAF33C59740789F72805CD08698A1C">Marques, Dayse Willkenia A.</mods:namePart>
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<mods:affiliation id="44911AF033B0CC5F0625F43E25CCB6F4">Instituto Nacional de Pesquisas da Amazônia, INPA, Manaus, Amazonas, Brazil; e-mails and ORCIDs: willkenia @ gmail. com, 0000 - 0001 - 7260 - 5760; jarafael @ inpa. gov. br, 0000 - 0002 - 0170 - 0514</mods:affiliation>
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<mods:namePart id="A2092F4E79AD48724FB94B162C0DA462">Rafael, José Albertino</mods:namePart>
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<mods:affiliation id="B06B2C25B4F8BF819589F3F4F578E26A">Instituto Nacional de Pesquisas da Amazônia, INPA, Manaus, Amazonas, Brazil; e-mails and ORCIDs: willkenia @ gmail. com, 0000 - 0001 - 7260 - 5760; jarafael @ inpa. gov. br, 0000 - 0002 - 0170 - 0514</mods:affiliation>
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<mods:title id="7DAB8796A51541C5FC28DB8593B9DA17">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="BB452AED45EEAE8EF59C8A8948597DB0">
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<mods:date id="C4F694A3309EC0D577D6FD6BBE2AE299">2023</mods:date>
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<mods:number id="A3A60B9FFD28BC776C00E38F004F2E1E">2023-12-28</mods:number>
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<mods:number id="2D35D64EB609AC433AED33D545FC47E6">63</mods:number>
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<mods:url id="B2D53A7E5D2922CBABCD3DF5D4363E74">http://dx.doi.org/10.37520/aemnp.2023.024</mods:url>
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<mods:classification id="E4E30A22DE4D6EB5F5C5F2798895AF32">journal article</mods:classification>
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<mods:identifier id="66B42590A46A7C029B911045EF8A1A2C" type="DOI">10.37520/aemnp.2023.024</mods:identifier>
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<treatment id="C00B8781CC03F1535AFA58EE2532151F" ID-DOI="http://doi.org/10.5281/zenodo.10621486" ID-Zenodo-Dep="10621486" LSID="urn:lsid:plazi:treatment:C00B8781CC03F1535AFA58EE2532151F" httpUri="http://treatment.plazi.org/id/C00B8781CC03F1535AFA58EE2532151F" lastPageId="5" lastPageNumber="388" pageId="2" pageNumber="385">
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<subSubSection id="00B8651CCC03F1545AFA58EE227C179E" box="[972,1292,188,212]" pageId="2" pageNumber="385" type="nomenclature">
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<paragraph id="481D3697CC03F1545AFA58EE227C179E" blockId="2.[972,1292,188,212]" box="[972,1292,188,212]" pageId="2" pageNumber="385">
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<heading id="135581FBCC03F1545AFA58EE227C179E" box="[972,1292,188,212]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="385" reason="3">
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<emphasis id="7AD6EA85CC03F1545AFA58EE227C179E" bold="true" box="[972,1292,188,212]" pageId="2" pageNumber="385">
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<taxonomicName id="8FA24D14CC03F1545AFA58EE23C0179E" authority="Ramos-Pastrana & Marques & Rafael, 2023" authorityName="Ramos-Pastrana & Marques & Rafael" authorityYear="2023" box="[972,1200,188,212]" class="Insecta" family="Pipunculidae" genus="Chalarus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="boyacensis" status="sp. nov.">
|
||
<emphasis id="7AD6EA85CC03F1545AFA58EE23C0179E" bold="true" box="[972,1200,188,212]" italics="true" pageId="2" pageNumber="385">Chalarus boyacensis</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="61E557FECC03F1545D8058EF227C179E" box="[1206,1292,189,212]" pageId="2" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="00B8651CCC03F1545D13588E23C317A5" box="[1061,1203,220,239]" pageId="2" pageNumber="385" type="description">
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<paragraph id="481D3697CC03F1545D13588E23C317A5" blockId="2.[828,1438,220,494]" box="[1061,1203,220,239]" pageId="2" pageNumber="385">
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||
(
|
||
<figureCitation id="D0992A12CC03F1545D1D588E23FE17A5" box="[1067,1166,220,239]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Figs 12–22</figureCitation>
|
||
,
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||
<figureCitation id="D0992A12CC03F1545DA0588E23DC17A5" box="[1174,1196,220,239]" captionStart="Fig" captionStartId="10.[151,182,1792,1811]" captionTargetBox="[151,1436,754,1762]" captionTargetId="figure-164@10.[151,1436,754,1762]" captionTargetPageId="10" captionText="Fig. 55. Geographical records of Chalarus species in Colombia. Chalarus absonus Rafael, 1990, C. boyacensis sp. nov., C. chairensis sp. nov., C. connexus Rafael, 1988, and C. delicatus Rafael, 1990." figureDoi="http://doi.org/10.5281/zenodo.10621452" httpUri="https://zenodo.org/record/10621452/files/figure.png" pageId="2" pageNumber="385">55</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="00B8651CCC03F1545A0A58AC233F16A4" pageId="2" pageNumber="385" type="materials_examined">
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||
<paragraph id="481D3697CC03F1545A0A58AC24BA1666" blockId="2.[828,1438,220,494]" pageId="2" pageNumber="385">
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||
<emphasis id="7AD6EA85CC03F1545A0A58AC24C5165B" bold="true" box="[828,949,254,273]" pageId="2" pageNumber="385">
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||
<typeStatus id="97198835CC03F1545A0A58AC2419165B" box="[828,873,254,273]" pageId="2" pageNumber="385">Type</typeStatus>
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||
locality.
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||
</emphasis>
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||
<collectingCountry id="30B57607CC03F1545A8D58AC2363165B" box="[955,1043,254,273]" name="Colombia" pageId="2" pageNumber="385">Colombia</collectingCountry>
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||
:
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||
<collectingRegion id="8A66F875CC03F1545D2A58AC232F165B" box="[1052,1119,254,273]" country="Colombia" name="Boyaca" pageId="2" pageNumber="385">Boyacá</collectingRegion>
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||
: Cabaña Chaina, Santuario de Fauna y Flora Iguaque.
|
||
</paragraph>
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||
<paragraph id="481D3697CC03F1545A0A5969233F16A4" blockId="2.[828,1438,220,494]" pageId="2" pageNumber="385">
|
||
<emphasis id="7AD6EA85CC03F1545A0A596924BB1604" bold="true" box="[828,971,315,334]" pageId="2" pageNumber="385">Type material</emphasis>
|
||
(
|
||
<materialsCitation id="F8CA3CCACC03F1545AEA5969233F16A4" collectionCode="II, LEUA" collectorName="A. Roberto & Canada & Left" country="Colombia" elevation="2550" latitude="5.4166665" location="Chaina" longLatPrecision="1305" longitude="-73.45" municipality="Iguaque" pageId="2" pageNumber="385" specimenCount="1" stateProvince="Boyaca" typeStatus="holotype">
|
||
2 JJ). HOLOTYඉE: J,
|
||
<emphasis id="7AD6EA85CC03F1545D8A5969224D1604" bold="true" box="[1212,1341,315,334]" pageId="2" pageNumber="385">
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||
<collectingCountry id="30B57607CC03F1545D8A5969224D1604" box="[1212,1341,315,334]" name="Colombia" pageId="2" pageNumber="385">COLOMBIA</collectingCountry>
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||
</emphasis>
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||
,
|
||
<emphasis id="7AD6EA85CC03F1545C78596922E61604" bold="true" box="[1358,1430,315,334]" pageId="2" pageNumber="385">
|
||
<collectingRegion id="8A66F875CC03F1545C78596922E61604" box="[1358,1430,315,334]" country="Colombia" name="Boyaca" pageId="2" pageNumber="385">Boyacá</collectingRegion>
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||
</emphasis>
|
||
, SFF[Santuario de Fauna y Flora] /
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||
<collectingMunicipality id="A879ACEDCC03F1545D4F590723B11622" box="[1145,1217,341,360]" pageId="2" pageNumber="385">Iguaque</collectingMunicipality>
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||
, Cab.[Cabaña]
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||
<location id="4D7D604CCC03F1545C7A590722FB1622" LSID="urn:lsid:plazi:treatment:C00B8781CC03F1535AFA58EE2532151F:4D7D604CCC03F1545C7A590722FB1622" box="[1356,1419,341,360]" country="Colombia" latitude="5.4166665" longLatPrecision="1305" longitude="-73.45" municipality="Iguaque" name="Chaina" pageId="2" pageNumber="385" stateProvince="Boyaca">Chaina</location>
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||
, /
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||
<geoCoordinate id="2D965050CC03F1545A0A5922240816C9" box="[828,888,368,387]" degrees="5" direction="north" minutes="25" orientation="latitude" pageId="2" pageNumber="385" precision="925" value="5.4166665">5º25’N</geoCoordinate>
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/
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||
<geoCoordinate id="2D965050CC03F1545AB6592224BF16C9" box="[896,975,368,387]" degrees="73" direction="west" minutes="27" orientation="longitude" pageId="2" pageNumber="385" precision="925" value="-73.45">73º27’W</geoCoordinate>
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,
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<quantity id="8F5A9B72CC03F1545AED5922235216C9" box="[987,1058,368,387]" metricMagnitude="3" metricUnit="m" metricValue="2.55" pageId="2" pageNumber="385" unit="m" value="2550.0">
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<elevation id="C38FD1A4CC03F1545AED5922235216C9" box="[987,1058,368,387]" metricMagnitude="3" metricUnit="m" metricValue="2.55" pageId="2" pageNumber="385" unit="m" value="2550.0">2550 m</elevation>
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</quantity>
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||
[eters], / 01–14.feb[
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||
<collectionCode id="2EB3AE52CC03F1545DE15922239716C9" box="[1239,1255,368,387]" pageId="2" pageNumber="385">II</collectionCode>
|
||
].2001, /
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||
<collectorName id="E5575341CC03F1545C0E592222EC16C9" box="[1336,1436,368,387]" pageId="2" pageNumber="385">A. Roberto</collectorName>
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||
leg. M1271 (IavH) (photographed specimen).
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||
<typeStatus id="97198835CC03F1545DFA59D9226B16D4" box="[1228,1307,395,414]" pageId="2" pageNumber="385" type="holotype">Holotype</typeStatus>
|
||
with left wing mounted on microslide with
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||
<collectorName id="E5575341CC03F1545D1A59F7231C16F2" box="[1068,1132,421,440]" pageId="2" pageNumber="385">Canada</collectorName>
|
||
balsam.
|
||
<collectorName id="E5575341CC03F1545D8359F723A916F2" box="[1205,1241,421,440]" pageId="2" pageNumber="385">Left</collectorName>
|
||
antenna and terminalia placed in a microvial with glycerin, both pinned along with the specimen. – PARATYඉE: 1 J,
|
||
<emphasis id="7AD6EA85CC03F1545AE45989248B16A4" box="[978,1019,475,494]" italics="true" pageId="2" pageNumber="385">idem</emphasis>
|
||
(
|
||
<collectionCode id="2EB3AE52CC03F1545D315989233516A4" box="[1031,1093,475,494]" pageId="2" pageNumber="385">LEUA</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00B8651CCC03F1545A0A5A5424991477" pageId="2" pageNumber="385" type="diagnosis">
|
||
<paragraph id="481D3697CC03F1545A0A5A5424991477" blockId="2.[828,1437,517,2077]" pageId="2" pageNumber="385">
|
||
<emphasis id="7AD6EA85CC03F1545A0A5A5424811557" bold="true" box="[828,1009,517,541]" pageId="2" pageNumber="385">
|
||
Diagnosis.
|
||
<emphasis id="7AD6EA85CC03F1545A855A57249B1557" bold="true" box="[947,1003,517,541]" italics="true" pageId="2" pageNumber="385">Male</emphasis>
|
||
.
|
||
</emphasis>
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||
Occiput dark brown, gray-brown pruinose. Both surstyli with one protuberance subapically, inward directed, outer margins slightly straight, apices rounded and downward directed; both surstyli with apices truncated and small sinus at junction with epandrium in lateral view. Left gonopod slightly shorter, wider, and with apex more rounded than right one. Tip of distiphallus short with apex hook-shaped and two symmetrical phallic processes, broad and dorsally with setae. Phallus trifid, all ejaculatory ducts situated at apex.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00B8651CCC03F1545A0A5B14235B10F7" pageId="2" pageNumber="385" type="description">
|
||
<paragraph id="481D3697CC03F1545A0A5B14232910D7" blockId="2.[828,1437,517,2077]" pageId="2" pageNumber="385">
|
||
<emphasis id="7AD6EA85CC03F1545A0A5B1423731417" bold="true" box="[828,1027,837,861]" pageId="2" pageNumber="385">
|
||
Description.
|
||
<emphasis id="7AD6EA85CC03F1545AFD5B1723731417" bold="true" box="[971,1027,837,861]" italics="true" pageId="2" pageNumber="385">Male</emphasis>
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="97198835CC03F1545D395B1423021417" box="[1039,1138,838,861]" pageId="2" pageNumber="385" type="holotype">holotype</typeStatus>
|
||
). Body length
|
||
<quantity id="8F5A9B72CC03F1545C3E5B14222B1417" box="[1288,1371,838,861]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="385" unit="mm" value="2.6">2.6 mm</quantity>
|
||
.
|
||
<emphasis id="7AD6EA85CC03F1545C555B1422ED1417" box="[1379,1437,838,861]" italics="true" pageId="2" pageNumber="385">Head</emphasis>
|
||
(
|
||
<figureCitation id="D0992A12CC03F1545A755B3424CB1437" box="[835,955,870,893]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Figs 12–13</figureCitation>
|
||
). Frons with upper 3/4 dark brown, dark brown pruinose, lower 1/4 gray-brown pruinose. Occiput dark brown, gray-brown pruinose. Antenna (
|
||
<figureCitation id="D0992A12CC03F1545DD95BF4224F14F7" box="[1263,1343,934,957]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Fig. 14</figureCitation>
|
||
) brown, pedicel with two dorsal and two ventral setae; LPP/WPP = 1.5.
|
||
<emphasis id="7AD6EA85CC03F1545ABE5BB424A614B7" box="[904,982,998,1021]" italics="true" pageId="2" pageNumber="385">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="D0992A12CC03F1545ADE5BB4231D14B7" box="[1000,1133,998,1021]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Figs 13, 15</figureCitation>
|
||
). Postpronotal lobe brown, brown pruinose. Scutum dark brown, gray-brown pruinose. Notopleuron and scutellum concolorous with scutum. Mesopleuron brown, gray-brown pruinose. Mediotergite concolorous with mesopleuron.
|
||
<emphasis id="7AD6EA85CC03F1545DAB5C3423A41337" box="[1181,1236,1126,1149]" italics="true" pageId="2" pageNumber="385">Wing</emphasis>
|
||
(
|
||
<figureCitation id="D0992A12CC03F1545DD05C34224A1337" box="[1254,1338,1126,1149]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Fig. 16</figureCitation>
|
||
). Length
|
||
<quantity id="8F5A9B72CC03F1545A0A5CD424E413D7" box="[828,916,1158,1181]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="385" unit="mm" value="2.6">2.6 mm</quantity>
|
||
. LW/MWW = 2.7; LTC/LFC = 5.1. Membrane brown infuscated. Halter stem dark brown, knob brown.
|
||
<emphasis id="7AD6EA85CC03F1545A0A5C94241F1397" box="[828,879,1222,1245]" italics="true" pageId="2" pageNumber="385">Legs</emphasis>
|
||
(
|
||
<figureCitation id="D0992A12CC03F1545AB75C9424A41397" box="[897,980,1222,1245]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Fig. 12</figureCitation>
|
||
) entirely dark brown, except tarsomeres 1–4 brown; pulvilli light brown.
|
||
<emphasis id="7AD6EA85CC03F1545D935CB4227A13B7" box="[1189,1290,1254,1277]" italics="true" pageId="2" pageNumber="385">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="D0992A12CC03F1545C2C5CB424281257" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Figs 12–13, 17</figureCitation>
|
||
) dark brown, gray-brown pruinose, with long setae laterally and short scattered setae dorsally; tergite 1 light brown basally, dark brown posterolaterally; tergites 1–5 with transverse brown pruinose bands; tergite 6 slightly shorter than tergite 5; syntergosternite 8 brown pruinose without long apical setae (
|
||
<figureCitation id="D0992A12CC03F1545D6E5DF423A612F7" box="[1112,1238,1446,1469]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Figs 17–18</figureCitation>
|
||
).
|
||
<emphasis id="7AD6EA85CC03F1545DDF5DF4222D12F7" box="[1257,1373,1446,1469]" italics="true" pageId="2" pageNumber="385">Terminalia</emphasis>
|
||
(
|
||
<figureCitation id="D0992A12CC03F1545C5B5DF424F21297" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Figs 18–22</figureCitation>
|
||
). Epandrium and surstyli brown (
|
||
<figureCitation id="D0992A12CC03F1545DD15D9422451297" box="[1255,1333,1478,1501]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Fig. 18</figureCitation>
|
||
). Surstyli (
|
||
<figureCitation id="D0992A12CC03F1545A755DB424CE12B7" box="[835,958,1510,1533]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Figs 18–20</figureCitation>
|
||
) subsymmetrical and setose, with long setae apically, apices rounded and downward directed; both surstyli with one protuberance subapically on inner margins; outer margins nearly straight (
|
||
<figureCitation id="D0992A12CC03F1545D805E1422741117" box="[1206,1284,1606,1629]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Fig. 18</figureCitation>
|
||
); both surstyli with apices truncated; left surstylus slightly thinner than right one, and with small sinus at junction with epandrium in lateral view (
|
||
<figureCitation id="D0992A12CC03F1545AD05EF4231311F7" box="[998,1123,1702,1725]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Figs 19–20</figureCitation>
|
||
). Gonopods subsymmetrical; left gonopod slightly shorter, wider, and with apex more rounded than right one (
|
||
<figureCitation id="D0992A12CC03F1545D725EB423B211B7" box="[1092,1218,1766,1789]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Figs 19–20</figureCitation>
|
||
). Tip of distiphallus short with apex hook-shaped and two symmetrical phallic processes, broad and with setae dorsally (
|
||
<figureCitation id="D0992A12CC03F1545DCF5F74222D1077" box="[1273,1373,1830,1853]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Fig. 21A</figureCitation>
|
||
). Ejaculatory apodeme parasol-shaped (
|
||
<figureCitation id="D0992A12CC03F1545D9B5F14238B1017" box="[1197,1275,1862,1885]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Fig. 22</figureCitation>
|
||
). Phallus trifid, with all ejaculatory ducts situated at apex in lateral and dorsal view (
|
||
<figureCitation id="D0992A12CC03F1545AF05FD4233D10D7" box="[966,1101,1926,1949]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="2" pageNumber="385">Figs 21A–B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="481D3697CC03F1545A6C5FF7235B10F7" blockId="2.[828,1437,517,2077]" box="[858,1067,1957,1981]" pageId="2" pageNumber="385">
|
||
<emphasis id="7AD6EA85CC03F1545A6C5FF724C410F7" bold="true" box="[858,948,1957,1981]" pageId="2" pageNumber="385">
|
||
<emphasis id="7AD6EA85CC03F1545A6C5FF724DD10F7" bold="true" box="[858,941,1957,1981]" italics="true" pageId="2" pageNumber="385">Female</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00B8651CCC03F1535A0A5F942532151F" lastPageId="5" lastPageNumber="388" pageId="2" pageNumber="385" type="etymology">
|
||
<paragraph id="481D3697CC03F1545A0A5F94227810B7" blockId="2.[828,1437,517,2077]" pageId="2" pageNumber="385">
|
||
<emphasis id="7AD6EA85CC03F1545A0A5F9424CC1097" bold="true" box="[828,956,1990,2013]" pageId="2" pageNumber="385">Etymology.</emphasis>
|
||
The specific epithet refers to the region of the
|
||
<typeStatus id="97198835CC03F1545A0A5FB4241A10B7" box="[828,874,2022,2045]" pageId="2" pageNumber="385">type</typeStatus>
|
||
locality,
|
||
<collectingRegion id="8A66F875CC03F1545AFA5FB4236F10B7" box="[972,1055,2022,2045]" country="Colombia" name="Boyaca" pageId="2" pageNumber="385">Boyacá</collectingRegion>
|
||
,
|
||
<collectingCountry id="30B57607CC03F1545D1F5FB423E610B7" box="[1065,1174,2022,2045]" name="Colombia" pageId="2" pageNumber="385">Colombia</collectingCountry>
|
||
; adjective.
|
||
</paragraph>
|
||
<paragraph id="481D3697CC03F1535A0A50542793165F" blockId="2.[828,1437,517,2077]" lastBlockId="5.[151,760,190,597]" lastPageId="5" lastPageNumber="388" pageId="2" pageNumber="385">
|
||
<emphasis id="7AD6EA85CC03F1545A0A505424EA1F57" bold="true" box="[828,922,2054,2077]" pageId="2" pageNumber="385">Habitat.</emphasis>
|
||
The specimens were collected in the Santuario de Fauna y Flora Iguaque reserve, where the vegetation is composed of forests of the cordillera of the Northeast region.
|
||
</paragraph>
|
||
<caption id="1CDD661FCC02F15559A15F0B224110EB" ID-DOI="http://doi.org/10.5281/zenodo.10621436" ID-Zenodo-Dep="10621436" httpUri="https://zenodo.org/record/10621436/files/figure.png" pageId="3" pageNumber="386" startId="3.[151,188,1881,1900]" targetBox="[153,1431,184,1863]" targetPageId="3" targetType="figure">
|
||
<paragraph id="481D3697CC02F15559A15F0B224110EB" blockId="3.[151,1437,1881,1953]" pageId="3" pageNumber="386">
|
||
Figs 1–11.
|
||
<taxonomicName id="8FA24D14CC02F15559C35F0B25721026" authority="Rafael, 1990" authorityName="Rafael" authorityYear="1990" box="[245,514,1881,1900]" class="Insecta" family="Pipunculidae" genus="Chalarus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="absonus">
|
||
<emphasis id="7AD6EA85CC02F15559C35F0B26E01026" box="[245,400,1881,1900]" italics="true" pageId="3" pageNumber="386">Chalarus absonus</emphasis>
|
||
<bibRefCitation id="2C334B66CC02F15558A35F0B25721026" author="RAFAEL J. A." box="[405,514,1881,1900]" pageId="3" pageNumber="386" pagination="45 - 53" refId="ref9661" refString="RAFAEL J. A. 1990: Revisao das especies Neotropicais do genero Chalarus Walker, 1834 (Diptera: Pipunculidae). Iheringa, Serie Zoologia 70: 45 - 53." type="journal article" year="1990">Rafael, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
(IAvH–M1271). Male. 1 – habitus, left lateral view; 2 – habitus, dorsal view; 3 – antenna; 4 – thorax, dorsal view; 5 – wing; 6 – abdomen, dorsal view; 7 – terminalia, dorsal view; 8 – left surstylus and gonopod, lateral view; 9 – right surstylus and gonopod, lateral view; 10a – phallic guide, subapical processes and phallus, right lateral view, 10b – phallus dorsal view; 11 – ejaculatory apodeme.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1CDD661FCC05F15259A15FBF22EE1F7F" ID-DOI="http://doi.org/10.5281/zenodo.10621440" ID-Zenodo-Dep="10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="4" pageNumber="387" startId="4.[151,188,2029,2048]" targetBox="[153,1433,188,2012]" targetPageId="4" targetType="figure">
|
||
<paragraph id="481D3697CC05F15259A15FBF22EE1F7F" blockId="4.[151,1438,2029,2101]" pageId="4" pageNumber="387">
|
||
Figs 12–22.
|
||
<taxonomicName id="8FA24D14CC05F15258375FBF26C41F4A" authority="Ramos-Pastrana & Marques & Rafael, 2023" authorityName="Ramos-Pastrana & Marques & Rafael" authorityYear="2023" box="[257,436,2029,2048]" class="Insecta" family="Pipunculidae" genus="Chalarus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="boyacensis" status="sp. nov.">
|
||
<emphasis id="7AD6EA85CC05F15258375FBF26C41F4A" box="[257,436,2029,2048]" italics="true" pageId="4" pageNumber="387">Chalarus boyacensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="61E557FECC05F152588F5FBF268C1F4A" box="[441,508,2029,2048]" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="481D3697CC04F15359A1594C2532151F" blockId="5.[151,760,190,597]" pageId="5" pageNumber="388">
|
||
<emphasis id="7AD6EA85CC04F15359A1594C26CF167F" bold="true" box="[151,447,286,309]" pageId="5" pageNumber="388">Geographical distribution.</emphasis>
|
||
<collectingCountry id="30B57607CC04F15358F3594C255D167F" box="[453,557,286,309]" name="Colombia" pageId="5" pageNumber="388">Colombia</collectingCountry>
|
||
(
|
||
<collectingRegion id="8A66F875CC04F1535B0F594C25FB167F" box="[569,651,286,309]" country="Colombia" name="Boyaca" pageId="5" pageNumber="388">Boyacá</collectingRegion>
|
||
) (
|
||
<figureCitation id="D0992A12CC04F1535BA9594C259C167F" box="[671,748,286,309]" captionStart="Fig" captionStartId="10.[151,182,1792,1811]" captionTargetBox="[151,1436,754,1762]" captionTargetId="figure-164@10.[151,1436,754,1762]" captionTargetPageId="10" captionText="Fig. 55. Geographical records of Chalarus species in Colombia. Chalarus absonus Rafael, 1990, C. boyacensis sp. nov., C. chairensis sp. nov., C. connexus Rafael, 1988, and C. delicatus Rafael, 1990." figureDoi="http://doi.org/10.5281/zenodo.10621452" httpUri="https://zenodo.org/record/10621452/files/figure.png" pageId="5" pageNumber="388">Fig. 55</figureCitation>
|
||
).
|
||
<emphasis id="7AD6EA85CC04F15359A1596C262A161F" bold="true" box="[151,346,318,341]" pageId="5" pageNumber="388">Taxonomic notes.</emphasis>
|
||
<taxonomicName id="8FA24D14CC04F1535868596C254A161F" authority="Ramos-Pastrana & Marques & Rafael, 2023" authorityName="Ramos-Pastrana & Marques & Rafael" authorityYear="2023" box="[350,570,318,341]" class="Insecta" family="Pipunculidae" genus="Chalarus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="boyacensis" status="sp. nov.">
|
||
<emphasis id="7AD6EA85CC04F1535868596C254A161F" box="[350,570,318,341]" italics="true" pageId="5" pageNumber="388">Chalarus boyacensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="61E557FECC04F1535B09596C25FF161F" box="[575,655,318,341]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
|
||
runs to
|
||
<taxonomicName id="8FA24D14CC04F1535BE9596C260E163F" authority="Collin, 1931" authorityName="Collin" authorityYear="1931" class="Insecta" family="Pipunculidae" genus="Chalarus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="chilensis">
|
||
<emphasis id="7AD6EA85CC04F1535BE9596C2784163F" italics="true" pageId="5" pageNumber="388">C. chilensis</emphasis>
|
||
<bibRefCitation id="2C334B66CC04F15359CC590C260E163F" author="COLLIN J. E." box="[250,382,350,373]" pageId="5" pageNumber="388" pagination="49 - 61" refId="ref8852" refString="COLLIN J. E. 1931: Platypezidae, Pipunculidae. Diptera of Patagonia and South Chile 6: 49 - 61." type="journal article" year="1931">
|
||
Collin,
|
||
<quantity id="8F5A9B72CC04F153587F590C260E163F" box="[329,382,350,373]" metricMagnitude="1" metricUnit="m" metricValue="4.904739999999999" pageId="5" pageNumber="388" unit="in" value="1931.0">1931</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in couplet 7 of the key presented by
|
||
<bibRefCitation id="2C334B66CC04F15359A1592C264916DF" author="RAFAEL J. A." box="[151,313,382,405]" pageId="5" pageNumber="388" pagination="45 - 53" refId="ref9661" refString="RAFAEL J. A. 1990: Revisao das especies Neotropicais do genero Chalarus Walker, 1834 (Diptera: Pipunculidae). Iheringa, Serie Zoologia 70: 45 - 53." type="journal article" year="1990">RAFAEL (1990)</bibRefCitation>
|
||
. It differs from
|
||
<taxonomicName id="8FA24D14CC04F15358D2592C251216DF" authorityName="Collin" authorityYear="1931" box="[484,610,382,405]" class="Insecta" family="Pipunculidae" genus="Chalarus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="chilensis">
|
||
<emphasis id="7AD6EA85CC04F15358D2592C251216DF" box="[484,610,382,405]" italics="true" pageId="5" pageNumber="388">C. chilensis</emphasis>
|
||
</taxonomicName>
|
||
in having the antenna brown; pedicel with two setae dorsally and two ventrally (
|
||
<figureCitation id="D0992A12CC04F153583659EC263A169F" box="[256,330,446,469]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="5" pageNumber="388">Fig. 14</figureCitation>
|
||
) [
|
||
<emphasis id="7AD6EA85CC04F153586959EC26D2169F" box="[351,418,446,469]" italics="true" pageId="5" pageNumber="388">versus</emphasis>
|
||
antenna black, pedicel with three setae dorsally and four ventrally in
|
||
<taxonomicName id="8FA24D14CC04F1535B16598C27A8155F" authority="RAFAEL (1988" authorityName="RAFAEL" authorityYear="1988" class="Insecta" family="Pipunculidae" genus="Chalarus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="chilensis">
|
||
<emphasis id="7AD6EA85CC04F1535B16598C254216BF" box="[544,562,478,501]" italics="true" pageId="5" pageNumber="388">C</emphasis>
|
||
.
|
||
<emphasis id="7AD6EA85CC04F1535B74598C25EF16BF" box="[578,671,478,501]" italics="true" pageId="5" pageNumber="388">chilensis</emphasis>
|
||
<bibRefCitation id="2C334B66CC04F1535B9E598C27A8155F" author="RAFAEL J. A." pageId="5" pageNumber="388" pagination="1 - 9" refId="ref9615" refString="RAFAEL J. A. 1988: Pipunculidae (Diptera) da regiao Neotropical. I. Redescricao de Chalarus chilensis Collin, comb. n. e descricao de duas especies novas da Amazonia. Revista Brasileira de Entomologia 5: 1 - 9." type="journal article" year="1988">RAFAEL (1988</bibRefCitation>
|
||
</taxonomicName>
|
||
:
|
||
<figureCitation id="D0992A12CC04F15359D759AC266B155F" box="[225,283,510,533]" captionStart="Figs 1–11" captionStartId="3.[151,188,1881,1900]" captionTargetBox="[153,1431,184,1863]" captionTargetId="figure-12@3.[138,1449,170,1881]" captionTargetPageId="3" captionText="Figs 1–11. Chalarus absonus Rafael, 1990 (IAvH–M1271). Male. 1 – habitus, left lateral view; 2 – habitus, dorsal view; 3 – antenna; 4 – thorax, dorsal view; 5 – wing; 6 – abdomen, dorsal view; 7 – terminalia, dorsal view; 8 – left surstylus and gonopod, lateral view; 9 – right surstylus and gonopod, lateral view; 10a – phallic guide, subapical processes and phallus, right lateral view, 10b – phallus dorsal view; 11 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621436" httpUri="https://zenodo.org/record/10621436/files/figure.png" pageId="5" pageNumber="388">Fig. 1</figureCitation>
|
||
)]; scutum and scutellum dark brown (
|
||
<figureCitation id="D0992A12CC04F1535B9359AC2580155F" box="[677,752,510,533]" captionStart="Figs 12–22" captionStartId="4.[151,188,2029,2048]" captionTargetBox="[153,1433,188,2012]" captionTargetId="figure-13@4.[136,1447,172,2026]" captionTargetPageId="4" captionText="Figs 12–22. Chalarus boyacensis sp. nov. (IAvH–M1271). Holotype male. 12 – habitus, left lateral view; 13 – habitus, dorsal view; 14 – antenna; 15 – thorax, dorsal view; 16 – wing; 17 – abdomen, dorsal view; 18 – terminalia, dorsal view; 19 – left surstylus and gonopod, lateral view; 20 – right surstylus and gonopod, lateral view; 21a – phallic guide, subapical processes and phallus, left lateral view, 21b – phallus dorsal view; 22 – ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.10621440" httpUri="https://zenodo.org/record/10621440/files/figure.png" pageId="5" pageNumber="388">Fig. 15</figureCitation>
|
||
) (
|
||
<emphasis id="7AD6EA85CC04F15359965A4C2794157F" box="[160,228,542,565]" italics="true" pageId="5" pageNumber="388">versus</emphasis>
|
||
scutum and scutellum black); the other different characters can be seen in the key below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |