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<document id="B12F0256BA482BDB02E7C99187ED21D6" ID-CLB-Dataset="9015" ID-DOI="10.11646/zootaxa.4861.3.4" ID-GBIF-Dataset="e4063d74-38c7-4442-90b9-6d7b929760ae" ID-ISSN="1175-5326" ID-Zenodo-Dep="4416421" ID-ZooBank="4E0A8C82-1C7A-4454-A5C5-54B8B62593CE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609799253807" checkinUser="plazi" docAuthor="Volynkin, Anton V." docDate="2020" docId="03EAED4E1548E817018589EEB6C7F90D" docLanguage="en" docName="zootaxa.4861.3.4.pdf" docOrigin="Zootaxa 4861 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyana (Gigantovulpecula) detoulgoeti Volynkin 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="379" masterDocId="FFD39536154AE81401128F52B742FF94" masterDocTitle="On the taxonomy of the subgenus Gigantovulpecula Karisch, 2013 of the genus Cyana Walker, 1854, with description of a new species (Lepidoptera: Erebidae Arctiinae: Lithosiini)" masterLastPageNumber="384" masterPageNumber="376" pageNumber="378" updateTime="1698852199270" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="ACC62E6D72878B463FEB4BDDD6235431" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="64932EEC3A8083F28022C19778ED23F0">On the taxonomy of the subgenus Gigantovulpecula Karisch, 2013 of the genus Cyana Walker, 1854, with description of a new species (Lepidoptera: Erebidae Arctiinae: Lithosiini)</mods:title>
</mods:titleInfo>
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<mods:namePart id="CABA8AAB5E71B1D0AA83FC92C19CDE7F">Volynkin, Anton V.</mods:namePart>
<mods:affiliation id="99E0089B408F2CC48CD4066354973C67">Altai State University, Lenina Avenue, 61, RF- 656049, Barnaul, Russia. &amp; National Research Tomsk State University, Lenina Avenue, 36, RF- 634050, Tomsk, Russia</mods:affiliation>
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<mods:title id="AD78A2646AB2CD60974F5CFB5D959434">Zootaxa</mods:title>
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<mods:part id="4CE3FB60CC6E3F674E79ACB5B887D605">
<mods:date id="727A8B4ECD6A6EFA0F7F91435989EAB3">2020</mods:date>
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<mods:number id="C5102D2AE542DB2BB5A70ECC6209389B">2020-10-19</mods:number>
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<mods:number id="EC6F967F04EB4FCF968F8984A8A766AD">4861</mods:number>
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<mods:classification id="2EC8E7F3512F59D666BB47EB83BCF1B1">journal article</mods:classification>
<mods:identifier id="0A504AE5AAC0B625CDE38860C1F65673" type="CLB-Dataset">9015</mods:identifier>
<mods:identifier id="D9AD0AB640112A3910220488ABDA7E97" type="DOI">10.11646/zootaxa.4861.3.4</mods:identifier>
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<mods:identifier id="0E37CF850A5F9EF7281ABCCFAA40989F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D9D69B5BD7560726DCCB1E0316F542B8" type="Zenodo-Dep">4416421</mods:identifier>
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<treatment id="03EAED4E1548E817018589EEB6C7F90D" ID-DOI="http://doi.org/10.5281/zenodo.4428257" ID-GBIF-Taxon="176442168" ID-Zenodo-Dep="4428257" LSID="urn:lsid:plazi:treatment:03EAED4E1548E817018589EEB6C7F90D" httpUri="http://treatment.plazi.org/id/03EAED4E1548E817018589EEB6C7F90D" lastPageId="3" lastPageNumber="379" pageId="2" pageNumber="378">
<subSubSection id="C3590FD31548E816018589EEB46FF942" box="[151,813,1723,1750]" pageId="2" pageNumber="378" type="nomenclature">
<paragraph id="8BFC5C581548E816018589EEB46FF942" blockId="2.[151,813,1723,1787]" box="[151,813,1723,1750]" pageId="2" pageNumber="378">
<heading id="D0B4EB341548E816018589EEB46FF942" bold="true" box="[151,813,1723,1750]" fontSize="11" level="1" pageId="2" pageNumber="378" reason="1">
<emphasis id="B937804A1548E816018589EEB46FF942" bold="true" box="[151,813,1723,1750]" pageId="2" pageNumber="378">
<taxonomicName id="4C4327DB1548E816018589EEB59CF942" ID-CoL="BHQHX" authority="Volynkin" authorityName="Volynkin" authorityYear="2020" box="[151,734,1723,1750]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="378" phylum="Arthropoda" rank="species" species="detoulgoeti" status="sp. nov." subGenus="Gigantovulpecula">
<emphasis id="B937804A1548E816018589EEB7A7F942" bold="true" box="[151,229,1724,1750]" italics="true" pageId="2" pageNumber="378">Cyana</emphasis>
(
<emphasis id="B937804A1548E81601E589E9B68CF942" bold="true" box="[247,462,1723,1750]" italics="true" pageId="2" pageNumber="378">Gigantovulpecula</emphasis>
)
<emphasis id="B937804A1548E81600CD89E9B527F942" bold="true" box="[479,613,1723,1750]" italics="true" pageId="2" pageNumber="378">detoulgoeti</emphasis>
Volynkin
</taxonomicName>
,
<taxonomicNameLabel id="A2043D311548E81603F989EEB46FF942" box="[747,813,1724,1750]" pageId="2" pageNumber="378" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3590FD31548E816018589B2B49DF8CF" pageId="2" pageNumber="378" type="description">
<paragraph id="8BFC5C581548E816018589B2B622F96E" blockId="2.[151,813,1723,1787]" box="[151,352,1760,1787]" pageId="2" pageNumber="378">
(
<figureCitation id="137840DD1548E81601B389B2B646F96E" box="[161,260,1760,1787]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1896,1920]" captionTargetBox="[160,1435,179,1860]" captionTargetId="figure-16@4.[151,1435,179,1870]" captionTargetPageId="4" captionText="FIGURES 1–8. Cyana spp.: adults. Depositories of specimens: 1–3, 5 and 6 in ZSM; 4 in NHMUK (©); 8 in MWM/ZSM. Figure 7 is given after Hampson (1900)." figureDoi="http://doi.org/10.5281/zenodo.4416423" httpUri="https://zenodo.org/record/4416423/files/figure.png" pageId="2" pageNumber="378">Figs 5, 6</figureCitation>
,
<figureCitation id="137840DD1548E816000289B2B615F96E" box="[272,343,1760,1786]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[165,1431,181,1910]" captionTargetId="figure-16@6.[151,1435,181,1920]" captionTargetPageId="6" captionText="FIGURES 13–16. Cyana spp.: male (13, 14) and female (15, 16) genitalia. Specimens are deposited in ZSM." figureDoi="http://doi.org/10.5281/zenodo.4416427" httpUri="https://zenodo.org/record/4416427/files/figure.png" pageId="2" pageNumber="378">13, 14</figureCitation>
)
</paragraph>
<paragraph id="8BFC5C581548E81601858877B49DF8CF" blockId="2.[151,1437,1828,1883]" pageId="2" pageNumber="378">
=
<taxonomicName id="4C4327DB1548E81601BE8876B6E5F8A8" authorityName="Butler" authorityYear="1882" box="[172,423,1828,1852]" class="Insecta" family="Arctiidae" genus="Chionaema" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="378" phylum="Arthropoda" rank="species" species="saalmuelleri">
<emphasis id="B937804A1548E81601BE8876B6E5F8A8" box="[172,423,1828,1852]" italics="true" pageId="2" pageNumber="378">Chionaema saalmuelleri</emphasis>
</taxonomicName>
<materialsCitation id="3B2B56051548E81600BF8876B498F8CF" ID-GBIF-Occurrence="3013444329" collectorName="Bulletin de la Societe" country="France" county="Ambohitantely Forest" location="Central" municipality="Ankazobe" pageId="2" pageNumber="378" specimenCount="1">
forme
<emphasis id="B937804A1548E81600E28876B534F8A8" box="[496,630,1828,1852]" italics="true" pageId="2" pageNumber="378">suberythraea</emphasis>
<bibRefCitation id="EFD221A91548E816036F8876B45AF8A9" author="Toulgoet, H. de" box="[637,792,1828,1853]" pageId="2" pageNumber="378" pagination="82 - 90" refId="ref4242" refString="Toulgoet, H. de (1971) Description d'Arctiidae nouvelles de Madagascar (Lep.) (16 e note). Bulletin de la Societe entomologique de France, 76 (2), 82 - 90. [in French]" type="journal article" year="1971">Toulgoët, 1971</bibRefCitation>
,
<emphasis id="B937804A1548E81602318876B24AF8A8" box="[803,1288,1828,1852]" italics="true" pageId="2" pageNumber="378">
<collectorName id="26B6398E1548E81602318876B4BEF8A8" box="[803,1020,1828,1852]" pageId="2" pageNumber="378">Bulletin de la Société</collectorName>
entomologique de
<collectingCountry id="F3541CC81548E81605AD8877B24AF8A8" box="[1215,1288,1829,1852]" name="France" pageId="2" pageNumber="378">France</collectingCountry>
</emphasis>
<emphasis id="B937804A1548E816041D8876B26AF8A8" bold="true" box="[1295,1320,1828,1852]" pageId="2" pageNumber="378">76</emphasis>
(2): 85, fig. 6 (
<typeStatus id="54F8E2FA1548E81601F38811B657F8CF" box="[225,277,1859,1883]" pageId="2" pageNumber="378">Type</typeStatus>
locality:
<collectingCountry id="F3541CC81548E81600678811B6ADF8CF" box="[373,495,1859,1883]" name="Madagascar" pageId="2" pageNumber="378">Madagascar</collectingCountry>
<location id="8E9C0A831548E81600E78811B500F8CF" LSID="urn:lsid:plazi:treatment:03EAED4E1548E817018589EEB6C7F90D:8E9C0A831548E81600E78811B500F8CF" box="[501,578,1859,1883]" country="France" county="Ambohitantely Forest" municipality="Ankazobe" name="Central" pageId="2" pageNumber="378">Central</location>
, N of
<collectingMunicipality id="6B98C6221548E816036D8811B5AAF8CF" box="[639,744,1859,1883]" pageId="2" pageNumber="378">Ankazobe</collectingMunicipality>
,
<collectingCounty id="629D24D41548E81603E38811B491F8CF" box="[753,979,1859,1883]" pageId="2" pageNumber="378">Ambohitantely Forest</collectingCounty>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3590FD31548E817018588DBB257FF6D" lastPageId="3" lastPageNumber="379" pageId="2" pageNumber="378" type="materials_examined">
<paragraph id="8BFC5C581548E816018588DBB3FFF853" blockId="2.[151,1437,1929,2028]" pageId="2" pageNumber="378">
<emphasis id="B937804A1548E816018588DBB603F837" bold="true" box="[151,321,1929,1955]" pageId="2" pageNumber="378">Type material</emphasis>
.
<materialsCitation id="3B2B56051548E816004388DBB3FFF853" ID-GBIF-Occurrence="3013444301" collectingDate="1955-02-18" collectingDateMax="1955-02-19" collectingDateMin="1955-02-18" collectionCode="ZSM" collectorName="A. Volynkin" country="Madagascar" location="Ankazobe" municipality="Vieu" pageId="2" pageNumber="378" specimenCount="250" specimenCount-male="250" typeStatus="holotype">
<emphasis id="B937804A1548E816004388DBB6FCF837" bold="true" box="[337,446,1929,1955]" pageId="2" pageNumber="378">
<typeStatus id="54F8E2FA1548E816004388DBB6FCF837" box="[337,446,1929,1955]" pageId="2" pageNumber="378" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="137840DD1548E81600C388DBB55EF830" box="[465,540,1929,1956]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1896,1920]" captionTargetBox="[160,1435,179,1860]" captionTargetId="figure-16@4.[151,1435,179,1870]" captionTargetPageId="4" captionText="FIGURES 1–8. Cyana spp.: adults. Depositories of specimens: 1–3, 5 and 6 in ZSM; 4 in NHMUK (©); 8 in MWM/ZSM. Figure 7 is given after Hampson (1900)." figureDoi="http://doi.org/10.5281/zenodo.4416423" httpUri="https://zenodo.org/record/4416423/files/figure.png" pageId="2" pageNumber="378">Figs 5</figureCitation>
,
<figureCitation id="137840DD1548E816033888DBB50BF837" box="[554,585,1929,1955]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[165,1431,181,1910]" captionTargetId="figure-16@6.[151,1435,181,1920]" captionTargetPageId="6" captionText="FIGURES 13–16. Cyana spp.: male (13, 14) and female (15, 16) genitalia. Specimens are deposited in ZSM." figureDoi="http://doi.org/10.5281/zenodo.4416427" httpUri="https://zenodo.org/record/4416427/files/figure.png" pageId="2" pageNumber="378">13</figureCitation>
): male, “
<collectingCountry id="F3541CC81548E81603A088DBB47EF837" box="[690,828,1929,1955]" name="Madagascar" pageId="2" pageNumber="378">Madagascar</collectingCountry>
C.,
<location id="8E9C0A831548E816027D88DBB4A1F837" LSID="urn:lsid:plazi:treatment:03EAED4E1548E817018589EEB6C7F90D:8E9C0A831548E816027D88DBB4A1F837" box="[879,995,1929,1955]" country="Madagascar" municipality="Vieu" name="Ankazobe" pageId="2" pageNumber="378">Ankazobe</location>
, Frt. Ambohitanteli,
<date id="FFFD7A981548E81605C688DBB239F830" box="[1236,1403,1929,1956]" pageId="2" pageNumber="378" value="1955-02-18" valueMax="1955-02-19" valueMin="1955-02-18">
<collectingDate id="EFB983701548E81605C688DBB239F830" box="[1236,1403,1929,1956]" pageId="2" pageNumber="378" value="1955-02-18" valueMax="1955-02-19" valueMin="1955-02-18">18–19 II 1955</collectingDate>
</date>
, P. Griveaud–R.
<collectingMunicipality id="6B98C6221548E816003C88FFB624F853" box="[302,358,1965,1991]" pageId="2" pageNumber="378">Vieu</collectingMunicipality>
” / “
<taxonomicName id="4C4327DB1548E816009C88FFB562F853" authorityName="Felder" authorityYear="1874" box="[398,544,1965,1991]" class="Insecta" family="Arctiidae" genus="Paratype" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="378" phylum="Arthropoda" rank="genus">
<typeStatus id="54F8E2FA1548E816009C88FFB562F853" box="[398,544,1965,1991]" pageId="2" pageNumber="378" type="paratype">PARATYPE</typeStatus>
</taxonomicName>
” / “Slide
<collectionCode id="ED52C49D1548E816039988FFB587F853" box="[651,709,1965,1991]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="378" type="Museum">ZSM</collectionCode>
Arct. 2019-
<specimenCount id="9D4597D11548E816025E88FFB4C8F85C" box="[844,906,1965,1992]" pageId="2" pageNumber="378" type="male">250♂</specimenCount>
<collectorName id="26B6398E1548E816028288FCB358F853" box="[912,1050,1965,1992]" pageId="2" pageNumber="378">A. Volynkin</collectorName>
” (Coll.
<collectionCode id="ED52C49D1548E816056188FFB3F3F853" box="[1139,1201,1965,1991]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="378" type="Museum">ZSM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BFC5C581548E81701D58883B257FF6D" blockId="2.[151,1437,1929,2028]" lastBlockId="3.[151,1437,151,1689]" lastPageId="3" lastPageNumber="379" pageId="2" pageNumber="378">
<emphasis id="B937804A1548E81601D58883B67DF87F" bold="true" box="[199,319,2001,2027]" pageId="2" pageNumber="378">
<typeStatus id="54F8E2FA1548E81601D58883B67DF87F" box="[199,319,2001,2027]" pageId="2" pageNumber="378" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D4597D11548E81600438883B6F2F878" box="[337,432,2001,2028]" pageId="2" pageNumber="378" type="male">4 males</specimenCount>
):
<specimenCount id="9D4597D11548E81600D78883B557F87F" box="[453,533,2001,2027]" pageId="2" pageNumber="378" type="male">1 male</specimenCount>
,
<collectingCountry id="F3541CC81548E81603318883B5EEF87F" box="[547,684,2001,2027]" name="Madagascar" pageId="2" pageNumber="378">Madagascar</collectingCountry>
, central area, road from Tananarive to Majunga, 4.IV.[19]65, 80., Collection Jacques Plante /
<taxonomicName id="4C4327DB1549E81700C28FC5B519FF25" authorityName="Felder" authorityYear="1874" box="[464,603,151,177]" class="Insecta" family="Arctiidae" genus="Paratype" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="genus">
<typeStatus id="54F8E2FA1549E81700C28FC5B519FF25" box="[464,603,151,177]" pageId="3" pageNumber="379" type="paratype">PARATYPE</typeStatus>
</taxonomicName>
/
<taxonomicName id="4C4327DB1549E81703618FC5B3D3FF25" authority="Toulgoet" authorityName="Toulgoet" box="[627,1169,151,177]" class="Insecta" family="Arctiidae" genus="Chionaema" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="379" phylum="Arthropoda" rank="subSpecies" species="saalmuelleri" subSpecies="suberythraea">
<emphasis id="B937804A1549E81703618FC5B366FF25" box="[627,1060,151,177]" italics="true" pageId="3" pageNumber="379">Chionaema saalmuelleri suberythraea</emphasis>
Toulgoët
</taxonomicName>
, slide
<collectionCode id="ED52C49D1549E81705C88FC5B256FF25" box="[1242,1300,151,177]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="379" type="Museum">ZSM</collectionCode>
Arct. 2019- 376 (male) Volynkin (Coll.
<collectionCode id="ED52C49D1549E81700C08FE9B552FF41" box="[466,528,187,213]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="379" type="Museum">ZSM</collectionCode>
);
<specimenCount id="9D4597D11549E81703348FE9B5C2FF41" box="[550,640,187,213]" pageId="3" pageNumber="379" type="male">2 males</specimenCount>
, EU et P. Griveaud, Tampoketsy, 18.III.53 /
<typeStatus id="54F8E2FA1549E81705698FE9B3ABFF41" box="[1147,1257,187,213]" pageId="3" pageNumber="379" type="paratype">Paratypus</typeStatus>
(Coll.
<collectionCode id="ED52C49D1549E81704298FE9B23BFF41" box="[1339,1401,187,213]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="379" type="Museum">ZSM</collectionCode>
);
<specimenCount id="9D4597D11549E817049C8FE9B792FF6D" pageId="3" pageNumber="379" type="male">1 male</specimenCount>
, SE
<collectingCountry id="F3541CC81549E81700118F8DB6CEFF6D" box="[259,396,223,249]" name="Madagascar" pageId="3" pageNumber="379">Madagascar</collectingCountry>
, Ranomafana, km. 402, Alt.
<quantity id="4CBBF1BD1549E81703DF8F8CB454FF6D" box="[717,790,222,249]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="379" unit="m" value="650.0">650 m</quantity>
, 26.III.–1.IV.55, H. de Toulgoёt (Coll.
<collectionCode id="ED52C49D1549E81705D88F8DB24AFF6D" box="[1226,1288,223,249]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="379" type="Museum">ZSM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3590FD31549E81701D58E51B527FD3D" pageId="3" pageNumber="379" type="discussion">
<paragraph id="8BFC5C581549E81701D58E51B527FD3D" blockId="3.[151,1437,151,1689]" pageId="3" pageNumber="379">
<emphasis id="B937804A1549E81701D58E51B676FE89" bold="true" box="[199,308,259,285]" pageId="3" pageNumber="379">Remarks</emphasis>
. (1) According to the article 45.6.3 of
<bibRefCitation id="EFD221A91549E81703E38E51B4C5FE89" author="ICZN" box="[753,903,259,285]" pageId="3" pageNumber="379" refId="ref3870" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust of Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN (1999)</bibRefCitation>
, the name first published after 1960 as “form” is infrasubspecific, thus, the Toulgoët’s name
<emphasis id="B937804A1549E81703B78E75B47BFED5" box="[677,825,295,321]" italics="true" pageId="3" pageNumber="379">suberythraea</emphasis>
is unavailable. (2) The
<typeStatus id="54F8E2FA1549E817055C8E75B3F2FED5" box="[1102,1200,295,321]" pageId="3" pageNumber="379" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4327DB1549E81705C88E75B7B6FEF1" authorityName="Butler" authorityYear="1882" class="Insecta" family="Arctiidae" genus="Chionaema" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="379" phylum="Arthropoda" rank="species" species="saalmuelleri">
<emphasis id="B937804A1549E81705C88E75B7B6FEF1" italics="true" pageId="3" pageNumber="379">Chionaema saalmuelleri</emphasis>
</taxonomicName>
forme
<emphasis id="B937804A1549E81700588E19B69CFEF1" box="[330,478,331,357]" italics="true" pageId="3" pageNumber="379">suberythraea</emphasis>
is housed in the MNHN collection (https://science.mnhn.fr/institution/mnhn/collection/el/item/el29068) and remains undissected. According to the articles 72.1.2 and 72.1.3 of
<bibRefCitation id="EFD221A91549E81705C18E3DB229FE1D" author="ICZN" box="[1235,1387,367,393]" pageId="3" pageNumber="379" refId="ref3870" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust of Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN (1999)</bibRefCitation>
, the
<typeStatus id="54F8E2FA1549E81701858EC1B7BBFE39" box="[151,249,403,429]" pageId="3" pageNumber="379" type="holotype">holotype</typeStatus>
and not
<typeStatus id="54F8E2FA1549E817004B8EC1B686FE39" box="[345,452,403,429]" pageId="3" pageNumber="379" type="paratype">paratypes</typeStatus>
has a name-bearing function only, from which it follows that in order for the description of the species to be correct, it is necessary to examine, describe and illustrate the genitalia of the
<typeStatus id="54F8E2FA1549E81705C48EE5B27EFE45" box="[1238,1340,439,465]" pageId="3" pageNumber="379" type="holotype">holotype</typeStatus>
. For this reason, here I designate the dissected male specimen housed in the ZSM collection as
<typeStatus id="54F8E2FA1549E817057D8E89B397FE61" box="[1135,1237,475,501]" pageId="3" pageNumber="379" type="holotype">holotype</typeStatus>
. (3) According to the original description (
<bibRefCitation id="EFD221A91549E81700A28EADB518FD8D" author="Toulgoet, H. de" box="[432,602,511,537]" pageId="3" pageNumber="379" pagination="82 - 90" refId="ref4242" refString="Toulgoet, H. de (1971) Description d'Arctiidae nouvelles de Madagascar (Lep.) (16 e note). Bulletin de la Societe entomologique de France, 76 (2), 82 - 90. [in French]" type="journal article" year="1971">Toulgoët 1971</bibRefCitation>
), besides the
<typeStatus id="54F8E2FA1549E81703EA8EADB41FFD8D" box="[760,861,511,537]" pageId="3" pageNumber="379" type="holotype">holotype</typeStatus>
, the type series of
<taxonomicName id="4C4327DB1549E817052B8EADB212FD8D" authorityName="Butler" authorityYear="1882" box="[1081,1360,511,537]" class="Insecta" family="Arctiidae" genus="Chionaema" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="379" phylum="Arthropoda" rank="species" species="saalmuelleri">
<emphasis id="B937804A1549E817052B8EADB212FD8D" box="[1081,1360,511,537]" italics="true" pageId="3" pageNumber="379">Chionaema saalmuelleri</emphasis>
</taxonomicName>
forme
<emphasis id="B937804A1549E81701858D71B669FDA9" box="[151,299,547,573]" italics="true" pageId="3" pageNumber="379">suberythraea</emphasis>
includes seven male
<typeStatus id="54F8E2FA1549E817030E8D71B5C5FDA9" box="[540,647,547,573]" pageId="3" pageNumber="379" type="paratype">paratypes</typeStatus>
from the Ambohitantely Forest being collected by P. Griveaud and R. Vieu in
<date id="FFFD7A981549E81701FD8D15B63BFDF5" box="[239,377,582,609]" pageId="3" pageNumber="379" value="1955-03">March 1955</date>
, and a male
<taxonomicName id="4C4327DB1549E81703118D15B521FDF5" authorityName="Felder" authorityYear="1874" box="[515,611,583,609]" class="Insecta" family="Arctiidae" genus="Paratype" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="genus">
<typeStatus id="54F8E2FA1549E81703118D15B521FDF5" box="[515,611,583,609]" pageId="3" pageNumber="379" type="paratype">paratype</typeStatus>
</taxonomicName>
from the same locality but collected by C. Moinier on
<date id="FFFD7A981549E81705D48D14B27AFDF5" box="[1222,1336,582,609]" pageId="3" pageNumber="379" value="1965-04-04">4.IV.1965</date>
. None of the specimens housed in the ZSM collection and labeled as
<typeStatus id="54F8E2FA1549E81702238D39B4DEFD11" box="[817,924,619,645]" pageId="3" pageNumber="379" type="paratype">paratypes</typeStatus>
has such label data therefore these specimens are not
<typeStatus id="54F8E2FA1549E81701F98DDDB614FD3D" box="[235,342,655,681]" pageId="3" pageNumber="379" type="paratype">paratypes</typeStatus>
of
<emphasis id="B937804A1549E817006E8DDDB552FD3D" box="[380,528,655,681]" italics="true" pageId="3" pageNumber="379">suberythraea</emphasis>
in fact.
</paragraph>
</subSubSection>
<subSubSection id="C3590FD31549E81701D58DE0B5E9FC16" pageId="3" pageNumber="379" type="diagnosis">
<paragraph id="8BFC5C581549E81701D58DE0B5E9FC16" blockId="3.[151,1437,151,1689]" pageId="3" pageNumber="379">
<emphasis id="B937804A1549E81701D58DE0B678FD58" bold="true" box="[199,314,690,716]" pageId="3" pageNumber="379">Diagnosis</emphasis>
. The new species differs clearly from two other members of the subgenus
<taxonomicName id="4C4327DB1549E817059F8DE1B214FD59" authorityName="Karisch" authorityYear="2013" box="[1165,1366,691,717]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="379" phylum="Arthropoda" rank="subGenus" subGenus="Gigantovulpecula">
<emphasis id="B937804A1549E817059F8DE1B214FD59" box="[1165,1366,691,717]" italics="true" pageId="3" pageNumber="379">Gigantovulpecula</emphasis>
</taxonomicName>
by its relatively small size and intense pale red suffusion on hindwing. The male genitalia of
<taxonomicName id="4C4327DB1549E81705978D85B264FD65" authority="Volynkin, 2020" authorityName="Volynkin" authorityYear="2020" box="[1157,1318,727,753]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="379" phylum="Arthropoda" rank="species" species="detoulgoeti" status="sp. nov.">
<emphasis id="B937804A1549E81705978D85B264FD65" box="[1157,1318,727,753]" italics="true" pageId="3" pageNumber="379">C. detoulgoeti</emphasis>
</taxonomicName>
<emphasis id="B937804A1549E817043D8D85B233FD65" bold="true" box="[1327,1393,727,753]" pageId="3" pageNumber="379">
<taxonomicNameLabel id="A2043D311549E817043D8D85B233FD65" box="[1327,1393,727,753]" pageId="3" pageNumber="379" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="4C4327DB1549E81700798DA9B55AFC81" authorityName="Butler" authorityYear="1882" box="[363,536,763,789]" class="Insecta" family="Arctiidae" genus="Chionaema" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="379" phylum="Arthropoda" rank="species" species="saalmuelleri">
<emphasis id="B937804A1549E81700798DA9B55AFC81" box="[363,536,763,789]" italics="true" pageId="3" pageNumber="379">C. saalmuelleri</emphasis>
</taxonomicName>
, but differ by the more robust distal saccular process, narrower aedeagus with a convex dorsal margin subapically (in
<taxonomicName id="4C4327DB1549E817032B8C4DB5A7FCAD" authorityName="Butler" authorityYear="1882" box="[569,741,799,825]" class="Insecta" family="Arctiidae" genus="Chionaema" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="379" phylum="Arthropoda" rank="species" species="saalmuelleri">
<emphasis id="B937804A1549E817032B8C4DB5A7FCAD" box="[569,741,799,825]" italics="true" pageId="3" pageNumber="379">C. saalmuelleri</emphasis>
</taxonomicName>
that is broader and almost straight), smaller spines on both the dorsal and distal diverticula, and the presence of the 3
<superScript id="7C36F1101549E81703E88C10B44AFCC4" attach="left" box="[762,776,834,848]" fontSize="6" pageId="3" pageNumber="379">rd</superScript>
subdiverticulum in the dorsal diverticulum (whereas in
<taxonomicName id="4C4327DB1549E81704918C11B661FC15" authorityName="Butler" authorityYear="1882" class="Insecta" family="Arctiidae" genus="Chionaema" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="379" phylum="Arthropoda" rank="species" species="saalmuelleri">
<emphasis id="B937804A1549E81704918C11B661FC15" italics="true" pageId="3" pageNumber="379">C. saalmuelleri</emphasis>
</taxonomicName>
there are two subdiverticula only).
</paragraph>
</subSubSection>
<subSubSection id="C3590FD31549E81701D58CD8B6D3F99D" pageId="3" pageNumber="379" type="description">
<paragraph id="8BFC5C581549E81701D58CD8B374FA71" blockId="3.[151,1437,151,1689]" pageId="3" pageNumber="379">
<emphasis id="B937804A1549E81701D58CD8B599FC31" bold="true" box="[199,731,906,933]" pageId="3" pageNumber="379">Description. External morphology of adults</emphasis>
. Forewing length
<quantity id="4CBBF1BD1549E81702A18CD8B36DFC31" box="[947,1071,906,933]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="3" pageNumber="379" unit="mm" value="14.5" valueMax="15.0" valueMin="14.0">14–15 mm</quantity>
in males. Male antenna ciliate. Head creamy white. Thorax ground color creamy white, with crimson spots medially and posteriorly; posterior part of patagia crimson; tegula with crimson sot anteriorly. Abdomen pink medially and creamy whitish basally and apically. Forewing ground color creamy white, transverse lines crimson. Subbasal spot represents large black dot. Subbasal line represents broad but short stroke at costa. Antemedial line nearly straight medially and posteriorly and V-like curved anteriorly. Postmedial line nearly straight, narrowed anteriorly and wavy at costa. Terminal line represents series of various sized spots between veins at wing apex. Cilia creamy white. Hindwing with intense pale red suffusion, but creamy whitish along costa. Cilia creamy whitish.
<emphasis id="B937804A1549E817029E8BD5B377FB34" bold="true" box="[908,1077,1158,1185]" pageId="3" pageNumber="379">Male genitalia</emphasis>
. Uncus short, narrowly trigonal, dorso-ventrally flattened, apically pointed, fully separated from tuba analis. Tuba analis broad, scaphium absent, subscaphium represent broad weakly setose area. Tegumen short and moderately broad. Anellus very broad, heavily scobinated. Juxta broadly X-shaped, moderately sclerotised. Vinculum short, U-shaped, robust. Valva moderately long and broad, broadened medially and narrowed distally, apically rounded. Costa convex medially. Medial crest of valva long but narrow, slightly prominent. Sacculus narrow, its distal process robust, slightly curved dorsally, with short thorn-like tip. Aedeagus moderately long, broadened distally, with convex dorsal wall. Aedeagus vesica broad, curved ventrally; dorsal diverticulum large, bears broad cluster of robust various sized spines and three short but broad subdiverticula; medial diverticulum moderately long, broad, with weakly scobinated area laterally; distal diverticulum short, bears broad and elongate cluster of robust various sized spines.
</paragraph>
<paragraph id="8BFC5C581549E81701D58ABDB6D3F99D" blockId="3.[151,1437,151,1689]" box="[199,401,1519,1545]" pageId="3" pageNumber="379">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C3590FD31549E81701D58940B417F9C5" pageId="3" pageNumber="379" type="distribution">
<paragraph id="8BFC5C581549E81701D58940B417F9C5" blockId="3.[151,1437,151,1689]" pageId="3" pageNumber="379">
<emphasis id="B937804A1549E81701D58940B61BF9B8" bold="true" box="[199,345,1554,1580]" pageId="3" pageNumber="379">Distribution</emphasis>
. Central and southeastern
<collectingCountry id="F3541CC81549E81703968941B449F9B9" box="[644,779,1555,1581]" name="Madagascar" pageId="3" pageNumber="379">Madagascar</collectingCountry>
(
<bibRefCitation id="EFD221A91549E81702088941B481F9B9" author="Toulgoet, H. de" box="[794,963,1555,1581]" pageId="3" pageNumber="379" pagination="82 - 90" refId="ref4242" refString="Toulgoet, H. de (1971) Description d'Arctiidae nouvelles de Madagascar (Lep.) (16 e note). Bulletin de la Societe entomologique de France, 76 (2), 82 - 90. [in French]" type="journal article" year="1971">Toulgoët 1971</bibRefCitation>
). In southeastern
<collectingCountry id="F3541CC81549E817059A8941B24DF9B9" box="[1160,1295,1555,1581]" name="Madagascar" pageId="3" pageNumber="379">Madagascar</collectingCountry>
(Ranomafa- na) the new species occurs sympatrically with
<taxonomicName id="4C4327DB1549E81703B38965B40CF9C5" authorityName="Butler" authorityYear="1882" box="[673,846,1591,1617]" class="Insecta" family="Arctiidae" genus="Chionaema" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="379" phylum="Arthropoda" rank="species" species="saalmuelleri">
<emphasis id="B937804A1549E81703B38965B40CF9C5" box="[673,846,1591,1617]" italics="true" pageId="3" pageNumber="379">C. saalmuelleri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3590FD31549E81701D58909B6C7F90D" pageId="3" pageNumber="379" type="etymology">
<paragraph id="8BFC5C581549E81701D58909B6C7F90D" blockId="3.[151,1437,151,1689]" pageId="3" pageNumber="379">
<emphasis id="B937804A1549E81701D58909B60AF9E1" bold="true" box="[199,328,1627,1653]" pageId="3" pageNumber="379">Etymology</emphasis>
. The species is named after Hervé de Toulgoët, who first described it as ‘
<taxonomicName id="4C4327DB1549E817059A8909B2DEF9E1" authorityName="Butler" authorityYear="1882" box="[1160,1436,1627,1653]" class="Insecta" family="Arctiidae" genus="Chionaema" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="379" phylum="Arthropoda" rank="species" species="saalmuelleri">
<emphasis id="B937804A1549E817059A8909B2DEF9E1" box="[1160,1436,1627,1653]" italics="true" pageId="3" pageNumber="379">Chionaema saalmuelleri</emphasis>
</taxonomicName>
forme
<emphasis id="B937804A1549E81701F3892DB637F90D" box="[225,373,1663,1689]" italics="true" pageId="3" pageNumber="379">suberythraea</emphasis>
’.
</paragraph>
</subSubSection>
</treatment>
</document>
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