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<document id="6D6EDA2B154C4298DFE31C17CB8F81BD" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629656873565" checkinUser="felipe" docAuthor="Chandler, Peter J." docDate="2009" docId="03EA87E1FFB8126111BBFB655BA66D8B" docLanguage="en" docName="zt02318p506.pdf" docOrigin="Zootaxa 2318" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pyratula takkae Chandler 2001" docType="treatment" docVersion="3" masterDocId="FFD3FF99FFB1126B112CFF9D58536D53" masterDocTitle="The fungus gnats (Diptera: Bolitophilidae, Keroplatidae, Mycetophilidae) of Sardinia, with description of six new species *" updateTime="1699090310938" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D0A138F18487A2E5EB29C69BD1F49C70">The fungus gnats (Diptera: Bolitophilidae, Keroplatidae, Mycetophilidae) of Sardinia, with description of six new species *</mods:title>
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<mods:namePart id="106AB9C839BAAFC37EFC8EBA5FF314B1">Chandler, Peter J.</mods:namePart>
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<mods:date id="DC15CF2AA804862FA83E80AF5D902875">2009</mods:date>
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<treatment id="03EA87E1FFB8126111BBFB655BA66D8B" ID-DOI="http://doi.org/10.5281/zenodo.6486266" ID-Zenodo-Dep="6486266" LSID="urn:lsid:plazi:treatment:03EA87E1FFB8126111BBFB655BA66D8B" httpUri="http://treatment.plazi.org/id/03EA87E1FFB8126111BBFB655BA66D8B" lastPageId="10" pageId="9">
<subSubSection id="C359657CFFB8126211BBFB655A316841" box="[151,610,1272,1299]" pageId="9" type="nomenclature">
<paragraph id="8BFC36F7FFB8126211BBFB655A316841" blockId="9.[151,610,1272,1338]" box="[151,610,1272,1299]" pageId="9">
<heading id="D0B4819BFFB8126211BBFB655A316841" bold="true" box="[151,610,1272,1299]" fontSize="11" level="1" pageId="9" reason="1">
<emphasis id="B937EAE5FFB8126211BBFB655A316841" bold="true" box="[151,610,1272,1299]" pageId="9">
11.
<taxonomicName id="4C434D74FFB8126211EDFB645A156841" ID-CoL="4QQMK" authority="Chandler, 2001" authorityName="Chandler" authorityYear="2001" box="[193,582,1272,1299]" class="Insecta" family="Keroplatidae" genus="Pyratula" kingdom="Animalia" order="Diptera" pageId="9" phylum="Arthropoda" rank="species" species="takkae">
<emphasis id="B937EAE5FFB8126211EDFB64592C6840" bold="true" box="[193,383,1273,1299]" italics="true" pageId="9">Pyratula takkae</emphasis>
Chandler, 2001
</taxonomicName>
**
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C359657CFFB8126211BBFABD59436869" box="[151,272,1312,1338]" pageId="9" type="description">
<paragraph id="8BFC36F7FFB8126211BBFABD59436869" blockId="9.[151,610,1272,1338]" box="[151,272,1312,1338]" pageId="9">
(
<figureCitation id="13782A72FFB8126211B3FABD59546869" box="[159,263,1312,1338]" captionStart="FIGURES 18" captionStartId="10.[151,275,1745,1769]" captionTargetBox="[153,1436,284,1723]" captionTargetId="figure-124@10.[151,1436,280,1724]" captionTargetPageId="10" captionText="FIGURES 18. Male genitalia. 13. Monocentrota matilei Bechev (Sardinia, sa Duchessa). 1. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 2. Aedeagus. 3. Tergite 9 and cerci. 45. Pyratula takkae Chandler (Sardinia, Colonia Beneck). 4. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 5. Tergite 9 and cerci. 6 8. Urytalpa juliae sp. nov. (paratype, Sardinia, sa Duchessa). 6. Ventral view of gonocoxites and gonostyli. 7. Aedeagus. 8. Tergite 9 and cerci." pageId="9">Figs 45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C359657CFFB8126111BBFAF05BA66D8B" lastPageId="10" pageId="9" type="materials_examined">
<paragraph id="8BFC36F7FFB8126211BBFAF05BE168D4" blockId="9.[151,1436,1389,2034]" box="[151,946,1389,1415]" pageId="9">
<emphasis id="B937EAE5FFB8126211BBFAF059E168D4" bold="true" box="[151,434,1389,1415]" pageId="9">
Material examined.
<geoCoordinate id="EE775030FFB8126210BEFAF059E168D4" box="[402,434,1389,1415]" degrees="1" direction="south" minutes="4" orientation="latitude" pageId="9" precision="925" value="-1.0666667">S1</geoCoordinate>
</emphasis>
418.IV.06
<specimenCount id="9D45FD7EFFB81262136DFAF05A3868D5" box="[577,619,1389,1415]" pageId="9" type="male">1 ♂</specimenCount>
.
<emphasis id="B937EAE5FFB812621356FAF05A9068D4" bold="true" box="[634,707,1389,1415]" pageId="9">SAR1</emphasis>
16.II15.VI.04
<specimenCount id="9D45FD7EFFB8126212ADFAF05BF868D5" box="[897,939,1389,1415]" pageId="9" type="male">1 ♂</specimenCount>
.
</paragraph>
<paragraph id="8BFC36F7FFB8126211EAFA0E59F36887" blockId="9.[151,1436,1389,2034]" pageId="9">
<emphasis id="B937EAE5FFB8126211EAFA0E590168FE" bold="true" box="[198,338,1427,1453]" pageId="9">Chorotype.</emphasis>
<materialsCitation id="3B2B3CAAFFB812621070FA0959CF6887" country="Greece" location="Mediterranean" pageId="9" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F88855FFB812621070FA0959C568FD" box="[348,406,1428,1454]" pageId="9">Type</typeStatus>
6 [3.01 of
<bibRefCitation id="EFD24B06FFB812621333FA095B2B68FD" author="Vigna Taglianti, A. &amp; Audisio, P. A. &amp; Biondi, M. &amp; Bologna, M. A. &amp; Carpaneto, G. M. &amp; De Biase, A. &amp; Fattorini, S. &amp; Piattella, E. &amp; Sindaco, R. &amp; Venchi, A. &amp; Zapparoli, M." box="[543,888,1428,1454]" pageId="9" pagination="31 - 59" refId="ref45744" refString="Vigna Taglianti, A., Audisio, P. A., Biondi, M., Bologna, M. A., Carpaneto, G. M., De Biase, A., Fattorini, S., Piattella, E., Sindaco, R., Venchi, A. &amp; Zapparoli, M. (1999) A proposal for a chorotype classification of the Near East fauna, in the framework of the Western Palaearctic region. Biogeographia, Lavori della Societa Italiana di Biogeografia (n. s.), 20, 31 - 59." type="journal article" year="1999">
Vigna Taglianti
<emphasis id="B937EAE5FFB8126213F2FA095B4868FE" box="[734,795,1428,1453]" italics="true" pageId="9">et al.</emphasis>
(1999)
</bibRefCitation>
].
<location id="8E9C602CFFB8126212A0FA095C6968FD" LSID="urn:lsid:plazi:treatment:03EA87E1FFB8126111BBFB655BA66D8B:8E9C602CFFB8126212A0FA095C6968FD" box="[908,1082,1428,1454]" country="Greece" name="Mediterranean" pageId="9">Mediterranean</location>
, previously known only from
<collectingCountry id="F3547667FFB8126211BBFA2758B96887" box="[151,234,1466,1492]" name="Greece" pageId="9">Greece</collectingCountry>
(Pelopponese)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFC36F7FFB8126211EAFA7D5A3568A8" blockId="9.[151,1436,1389,2034]" box="[198,614,1504,1531]" pageId="9">
<emphasis id="B937EAE5FFB8126211EAFA7D59EF68A9" bold="true" box="[198,444,1504,1530]" pageId="9">Italian distribution.</emphasis>
<collectingRegion id="4987F815FFB8126210E8FA7C5A7668A8" box="[452,549,1505,1531]" country="Italy" name="Sardegna" pageId="9">Sardinia</collectingRegion>
only.
</paragraph>
<paragraph id="8BFC36F7FFB8126211EAF99A5A456A2D" blockId="9.[151,1436,1389,2034]" pageId="9">
<emphasis id="B937EAE5FFB8126211EAF99A59406B72" bold="true" box="[198,275,1543,1569]" pageId="9">Notes.</emphasis>
<bibRefCitation id="EFD24B06FFB812621036F9955A976B71" author="Chandler, P. J. &amp; Blasco-Zumeta, J." box="[282,708,1544,1570]" pageId="9" pagination="1 - 24" refId="ref42136" refString="Chandler, P. J. &amp; Blasco-Zumeta, J. (2001) The fungus gnats (Diptera, Bolitophilidae, Keroplatidae and Mycetophilidae) of the Monegros region (Zaragoza, Spain) and five other new European species of Pyratula Edwards and Sciophila Meigen. Zapateri. Revista Aragonesa de Entomologia, 9, 1 - 24." type="journal article" year="2001">Chandler and Blasco-Zumeta (2001)</bibRefCitation>
included descriptions of six species of the
<taxonomicName id="4C434D74FFB812621594F995591E6B1B" authority="Edwards, 1925" authorityName="Edwards" authorityYear="1925" class="Insecta" family="Keroplatidae" genus="Pyratula" kingdom="Animalia" order="Diptera" pageId="9" phylum="Arthropoda" rank="species" species="perpusilla">
<emphasis id="B937EAE5FFB812621594F9955DCF6B72" box="[1208,1436,1544,1569]" italics="true" pageId="9">Pyratula perpusilla</emphasis>
<bibRefCitation id="EFD24B06FFB8126211BBF9B3591E6B1B" author="Edwards, F. W." box="[151,333,1582,1608]" pageId="9" pagination="505 - 670" refId="ref42843" refString="Edwards, F. W. (1925) British Fungus-Gnats (Diptera, Mycetophilidae). With a revised Generic Classification of the Family. Transactions of the Entomological Society of London, 73, 505 - 670." type="journal article" year="1925">Edwards, 1925</bibRefCitation>
</taxonomicName>
group, mostly from the Mediterranean region, only
<taxonomicName id="4C434D74FFB8126212FDF9B25C386B1B" authorityName="Edwards" authorityYear="1925" box="[977,1131,1583,1608]" class="Insecta" family="Keroplatidae" genus="Pyratula" kingdom="Animalia" order="Diptera" pageId="9" phylum="Arthropoda" rank="species" species="perpusilla">
<emphasis id="B937EAE5FFB8126212FDF9B25C386B1B" box="[977,1131,1583,1608]" italics="true" pageId="9">P. perpusilla</emphasis>
</taxonomicName>
itself being known from northern Europe. Among the species described in that paper these specimens are concluded to be conspecific with
<taxonomicName id="4C434D74FFB8126211FEF9E1596F6BC6" authorityName="Chandler" authorityYear="2001" box="[210,316,1660,1685]" class="Insecta" family="Keroplatidae" genus="Pyratula" kingdom="Animalia" order="Diptera" pageId="9" phylum="Arthropoda" rank="species" species="takkae">
<emphasis id="B937EAE5FFB8126211FEF9E1596F6BC6" box="[210,316,1660,1685]" italics="true" pageId="9">P.takkae</emphasis>
</taxonomicName>
pending discovery of further material. They agree well in most characters of the male genitalia (
<figureCitation id="13782A72FFB8126211B3F93F595E6BEF" box="[159,269,1698,1724]" captionStart="FIGURES 18" captionStartId="10.[151,275,1745,1769]" captionTargetBox="[153,1436,284,1723]" captionTargetId="figure-124@10.[151,1436,280,1724]" captionTargetPageId="10" captionText="FIGURES 18. Male genitalia. 13. Monocentrota matilei Bechev (Sardinia, sa Duchessa). 1. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 2. Aedeagus. 3. Tergite 9 and cerci. 45. Pyratula takkae Chandler (Sardinia, Colonia Beneck). 4. Ventral view of gonocoxites and gonostyli, with aedeagus in situ. 5. Tergite 9 and cerci. 6 8. Urytalpa juliae sp. nov. (paratype, Sardinia, sa Duchessa). 6. Ventral view of gonocoxites and gonostyli. 7. Aedeagus. 8. Tergite 9 and cerci." pageId="9">Figs 45</figureCitation>
), in particular in the structure of the apical margin of the gonocoxites, which is characteristic in species of this group. The
<typeStatus id="54F88855FFB8126210CDF9545A1F6BB0" box="[481,588,1737,1763]" pageId="9" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C434D74FFB81262135BF9545AB56BB1" authorityName="Chandler" authorityYear="2001" box="[631,742,1737,1762]" class="Insecta" family="Keroplatidae" genus="Pyratula" kingdom="Animalia" order="Diptera" pageId="9" phylum="Arthropoda" rank="species" species="takkae">
<emphasis id="B937EAE5FFB81262135BF9545AB56BB1" box="[631,742,1737,1762]" italics="true" pageId="9">P. takkae</emphasis>
</taxonomicName>
was unusual in this genus in having strong setae on the mediotergite. These specimens differ in lacking these setae so in the absence of other material it is not possible to say which condition is typical of the species. The
<typeStatus id="54F88855FFB8126212B4F88B5C506A63" box="[920,1027,1814,1840]" pageId="9" type="holotype">holotype</typeStatus>
has the head damaged and lacks flagellomeres. The Sardinian specimens have the antenna 1.5 times as long as the thorax and the flagellomeres are about twice as long as broad.
</paragraph>
<paragraph id="8BFC36F7FFB8126111EAF8175BA66D8B" blockId="9.[151,1436,1389,2034]" lastBlockId="10.[151,1436,152,216]" lastPageId="10" pageId="9">
<bibRefCitation id="EFD24B06FFB8126211EAF81759CA6AF7" author="Chandler, P. J." box="[198,409,1930,1956]" pageId="9" pagination="195 - 203" refId="ref41815" refString="Chandler, P. J. (2004 b) Fungus gnats (Diptera, Sciaroidea: Ditomyiidae, Keroplatidae and Mycetophilidae). In: Cerretti, P., Hardersen, S., Mason, F., Nardi, G., Tisato, M. &amp; Zapparoli, M. (Eds), Invertebrati di una foresta della Pianura Padana, Bosco della Fontana. Secondo contributo. Conservazione Habitat Invertebrati, 3. Cierre Grafica Editore, Verona, pp. 195 - 203." type="book chapter" year="2004">Chandler (2004b)</bibRefCitation>
recorded
<taxonomicName id="4C434D74FFB812621339F8165AFF6AF7" authorityName="Edwards" authorityYear="1925" box="[533,684,1931,1956]" class="Insecta" family="Keroplatidae" genus="Pyratula" kingdom="Animalia" order="Diptera" pageId="9" phylum="Arthropoda" rank="species" species="perpusilla">
<emphasis id="B937EAE5FFB812621339F8165AFF6AF7" box="[533,684,1931,1956]" italics="true" pageId="9">P. perpusilla</emphasis>
</taxonomicName>
as new to
<collectingCountry id="F3547667FFB81262121BF8175B3E6AF7" box="[823,877,1930,1956]" name="Italy" pageId="9">Italy</collectingCountry>
from Bosco della Fontana and
<bibRefCitation id="EFD24B06FFB8126215DEF8175DCF6AF7" author="Kurina, O." box="[1266,1436,1930,1956]" pageId="9" pagination="245 - 293" refId="ref43515" refString="Kurina, O. (2008) Sciaroidea excl. Sciaridae. In: Ziegler, J. (Ed.), Diptera Stelviana. A dipterological perspective on a changing alpine landscape. Volume 1. Studia dipterologica, Supplement 16, pp. 245 - 293." type="journal article" year="2008">Kurina (2008)</bibRefCitation>
recorded two species of this group from South Tyrol,
<taxonomicName id="4C434D74FFB812621209F82C5C3B6A98" authority="Chandler, 2001" authorityName="Chandler" authorityYear="2001" box="[805,1128,1969,1995]" class="Insecta" family="Keroplatidae" genus="Pyratula" kingdom="Animalia" order="Diptera" pageId="9" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="B937EAE5FFB812621209F82C5BF56A99" box="[805,934,1969,1994]" italics="true" pageId="9">P. alpicola</emphasis>
Chandler, 2001
</taxonomicName>
(also known from
<collectingCountry id="F3547667FFB81262147DF82C59596AA1" name="Czech Republic" pageId="9">Czech Republic</collectingCountry>
, the Swiss Alps and
<collectingCountry id="F3547667FFB81262133DF8455A296AA1" box="[529,634,2008,2034]" name="Andorra" pageId="9">Andorra</collectingCountry>
) and
<taxonomicName id="4C434D74FFB8126213EDF8455C5A6AA1" authority="Chandler, 1994" authorityName="Chandler" authorityYear="1994" box="[705,1033,2008,2034]" class="Insecta" family="Keroplatidae" genus="Pyratula" kingdom="Animalia" order="Diptera" pageId="9" phylum="Arthropoda" rank="species" species="oracula">
<emphasis id="B937EAE5FFB8126213EDF8455B136AA2" box="[705,832,2008,2033]" italics="true" pageId="9">P. oracula</emphasis>
<bibRefCitation id="EFD24B06FFB812621266F8455C5A6AA1" author="Chandler, P. J." box="[842,1033,2008,2034]" pageId="9" pagination="1 - 100" refId="ref41464" refString="Chandler, P. J. (1994) Fungus gnats of Israel (Diptera: Sciaroidea, excluding Sciaridae). Israel Journal of Entomology, 28, 1 - 100." type="journal article" year="1994">Chandler, 1994</bibRefCitation>
</taxonomicName>
(mainly Mediterranean but also occurring in
<collectingCountry id="F3547667FFBB12611002FF0559BA6DE1" box="[302,489,152,178]" name="Czech Republic" pageId="10">Czech Republic</collectingCountry>
and
<collectingCountry id="F3547667FFBB12611308FF055AE66DE1" box="[548,693,152,178]" name="Switzerland" pageId="10">Switzerland</collectingCountry>
). The finding of this species in
<collectingRegion id="4987F815FFBB12611505FF055CD96DE1" box="[1065,1162,152,178]" country="Italy" name="Sardegna" pageId="10">Sardinia</collectingRegion>
is thus of considerable interest, identifying
<collectingCountry id="F3547667FFBB126110AFFF2359EA6D8B" box="[387,441,190,216]" name="Italy" pageId="10">Italy</collectingCountry>
as having the richest known fauna of this genus.
</paragraph>
</subSubSection>
</treatment>
</document>