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<document id="0A083D484F86A2DA4E606193BF8392B4" ID-DOI="10.11646/zootaxa.4759.4.1" ID-GBIF-Dataset="4569c2ca-cbbb-49cb-8460-494916d5f3d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3740949" ID-ZooBank="131E86B0-BB4B-4D91-8F48-6F2A2207B424" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586154517882" checkinUser="plazi" docAuthor="Park, Kyu-Tek, Mey, Wolfram, Koo, Jun-Mo, Prins, Jurate De, Akite, Perpetra &amp; Cho, Soowon" docDate="2020" docId="03D46157FFABFFBFFF58FBFBFDB3FE5F" docLanguage="en" docName="zootaxa.4759.4.1.pdf" docOrigin="Zootaxa 4759 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thubdora umbratilis Park 2020, sp. nov." docType="treatment" docUuid="DC61858A-5586-439C-BE99-BC54179A0EC4" docUuidSource="ZooBank" docVersion="5" lastPageNumber="469" masterDocId="FFED192FFFBAFFADFFCFFF9EFF97FFF0" masterDocTitle="Fourteen new species of the genus Thubdora Park, 2018 (Lepidoptera Gelechioidea: Lecithoceridae) from Uganda, and three new combinations in Ptilothyris Walsingham, 1897 from DR Congo" masterLastPageNumber="487" masterPageNumber="451" pageNumber="468" updateTime="1698735467765" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="34268536369B0B0F6AC37B01EC23D648">Fourteen new species of the genus Thubdora Park, 2018 (Lepidoptera Gelechioidea: Lecithoceridae) from Uganda, and three new combinations in Ptilothyris Walsingham, 1897 from DR Congo</mods:title>
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<mods:namePart id="429F355D98588D0B96E1DB145E6D5AC1">Park, Kyu-Tek</mods:namePart>
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<mods:namePart id="2BA5F7C249C2C76CC1671D007FF3F6A7">Mey, Wolfram</mods:namePart>
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<mods:namePart id="87A066209AFD9EF6F46067E876F4C6F0">Koo, Jun-Mo</mods:namePart>
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<mods:namePart id="A888E80AFB1E4D10E2829494D4F4BB4C">Prins, Jurate De</mods:namePart>
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<mods:namePart id="8C8AC14B23E919FA1A7B7FF48416817F">Akite, Perpetra</mods:namePart>
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<mods:namePart id="6DB5F94BAA8417FDA73AC6A1697281AC">Cho, Soowon</mods:namePart>
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<mods:title id="6EA39B6D753A5FA09D3569603C746BAB">Zootaxa</mods:title>
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<mods:date id="DF4220EE41F7CF66CFB79602616A18B6">2020</mods:date>
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<mods:number id="E424D076962E4CD614BADAA11331D55C">2020-04-06</mods:number>
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<subSubSection id="C36783CAFFABFFBCFF58FBFBFDF8FB8F" box="[151,623,1125,1152]" pageId="17" pageNumber="468" type="nomenclature">
<paragraph id="8BC2D041FFABFFBCFF58FBFBFDF8FB8F" blockId="17.[151,623,1125,1187]" box="[151,623,1125,1152]" pageId="17" pageNumber="468">
<heading id="D08A672DFFABFFBCFF58FBFBFDF8FB8F" bold="true" box="[151,623,1125,1152]" fontSize="11" level="1" pageId="17" pageNumber="468" reason="1">
<emphasis id="B9090C53FFABFFBCFF58FBFBFDF8FB8F" bold="true" box="[151,623,1125,1152]" pageId="17" pageNumber="468">
11.
<taxonomicName id="4C7DABC2FFABFFBCFF0DFBFBFD92FB8F" ID-CoL="9FSGF" authority="Park" authorityName="Park" authorityYear="2020" box="[194,517,1125,1152]" class="Insecta" family="Lecithoceridae" genus="Thubdora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="umbratilis" status="sp. nov.">
<emphasis id="B9090C53FFABFFBCFF0DFBFBFE29FB70" bold="true" box="[194,446,1125,1152]" italics="true" pageId="17" pageNumber="468">Thubdora umbratilis</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="A23AB128FFABFFBCFDDEFBFBFDF8FB8F" box="[529,623,1125,1151]" pageId="17" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C36783CAFFABFFBCFF58FB17FEACFB53" box="[151,315,1161,1187]" pageId="17" pageNumber="468" type="description">
<paragraph id="8BC2D041FFABFFBCFF58FB17FEACFB53" blockId="17.[151,623,1125,1187]" box="[151,315,1161,1187]" pageId="17" pageNumber="468">
(
<figureCitation id="1346CCC4FFABFFBCFF50FB17FF6EFB53" box="[159,249,1161,1187]" captionStart="FIGURE 12" captionStartId="18.[151,250,1613,1638]" captionTargetBox="[215,1364,465,1575]" captionTargetId="figure@18.[208,1380,453,1588]" captionTargetPageId="18" captionText="FIGURE 12. Thubdora umbratilis Park, sp. nov.: A, adult, holotype; B, head and antenna, dorsal view; C, forewing venation; D, male genitalia, holotype, gen. slide no. CIS-7048, E, lateral view, holotype; F, aedeagus; G, abdominal segments VI-VIII. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3740973" httpUri="https://zenodo.org/record/3740973/files/figure.png" pageId="17" pageNumber="468">Figs. 12</figureCitation>
AG)
</paragraph>
</subSubSection>
<subSubSection id="C36783CAFFABFFBCFF58FB4FFB8BFAFF" pageId="17" pageNumber="468" type="materials_examined">
<paragraph id="8BC2D041FFABFFBCFF58FB4FFB8BFAFF" blockId="17.[151,1437,1232,2015]" pageId="17" pageNumber="468">
<emphasis id="B9090C53FFABFFBCFF58FB4FFED8FB1B" bold="true" box="[151,335,1232,1259]" pageId="17" pageNumber="468">Type specimen.</emphasis>
<materialsCitation id="3B15DA1CFFABFFBCFE98FB4FFB8BFAFF" ID-GBIF-Occurrence="2595809614" collectingDate="2014-11-25" collectingDateMax="2014-11-30" collectingDateMin="2014-11-25" collectionCode="MfN" collectorName="W. Mey" country="Uganda" location="Mpanga Forest" pageId="17" pageNumber="468" specimenCount="1" stateProvince="Mpigi" typeStatus="holotype">
<typeStatus id="54C66EE3FFABFFBCFE98FB4FFE53FB1B" box="[343,452,1233,1259]" pageId="17" pageNumber="468" type="holotype">Holotype</typeStatus>
: male,
<emphasis id="B9090C53FFABFFBCFDDBFB4FFDEFFB1B" bold="true" box="[532,632,1233,1259]" pageId="17" pageNumber="468">
<collectingCountry id="F36A90D1FFABFFBCFDDBFB4FFDE4FB1B" box="[532,627,1233,1259]" name="Uganda" pageId="17" pageNumber="468">Uganda</collectingCountry>
,
</emphasis>
<collectingRegion id="49B91EA3FFABFFBCFD4FFB4FFD50FB1B" box="[640,711,1233,1259]" country="Uganda" name="Mpigi" pageId="17" pageNumber="468">Mpigi</collectingRegion>
,
<location id="8EA2869AFFABFFBCFD1CFB4FFC17FB1B" LSID="urn:lsid:plazi:treatment:03D46157FFABFFBFFF58FBFBFDB3FE5F:8EA2869AFFABFFBCFD1CFB4FFC17FB1B" box="[723,896,1233,1259]" country="Uganda" name="Mpanga Forest" pageId="17" pageNumber="468" stateProvince="Mpigi">Mpanga Forest</location>
,
<date id="FFC3F681FFABFFBCFC43FB4EFBA5FB1B" box="[908,1074,1232,1259]" pageId="17" pageNumber="468" value="2014-11-25" valueMax="2014-11-30" valueMin="2014-11-25">
<collectingDate id="EF870F69FFABFFBCFC43FB4EFBA5FB1B" box="[908,1074,1232,1259]" pageId="17" pageNumber="468" value="2014-11-25" valueMax="2014-11-30" valueMin="2014-11-25">2530 xi 2014</collectingDate>
</date>
, LF, leg.
<collectorName id="2688B597FFABFFBCFB53FB4FFB6EFB1B" box="[1180,1273,1233,1259]" pageId="17" pageNumber="468">W. Mey</collectorName>
; gen. slide no. CIS-7048, wing slide no. CIS-7054, COI barcode CBNU008, deposited in
<collectionCode id="ED6C4884FFABFFBCFC11FB6BFB8FFAFF" box="[990,1048,1269,1295]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="17" pageNumber="468" type="Museum">MfN</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36783CAFFABFFBCFF08FA86FC15FA8B" pageId="17" pageNumber="468" type="diagnosis">
<paragraph id="8BC2D041FFABFFBCFF08FA86FC15FA8B" blockId="17.[151,1437,1232,2015]" pageId="17" pageNumber="468">
<emphasis id="B9090C53FFABFFBCFF08FA86FED5FAC3" bold="true" box="[199,322,1304,1331]" pageId="17" pageNumber="468">Diagnosis.</emphasis>
This species is one of the smallest species of the genus, with forewing length of
<quantity id="4C857DA4FFABFFBCFB00FA86FABEFAC3" box="[1231,1321,1304,1331]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="17" pageNumber="468" unit="mm" value="6.5">6.5 mm</quantity>
. The male genitalia can be distinguished from those of all the preceding species by the cucullus narrowed toward apex and the aedeagus with a large, heavily sclerotized, sword-shaped cornutus.
</paragraph>
</subSubSection>
<subSubSection id="C36783CAFFABFFBFFF08FA1AFE06FEB3" lastPageId="18" lastPageNumber="469" pageId="17" pageNumber="468" type="description">
<paragraph id="8BC2D041FFABFFBCFF08FA1AFC2DF9DF" blockId="17.[151,1437,1232,2015]" pageId="17" pageNumber="468">
<emphasis id="B9090C53FFABFFBCFF08FA1AFECEFA6F" bold="true" box="[199,345,1412,1439]" pageId="17" pageNumber="468">Description.</emphasis>
Male (
<figureCitation id="1346CCC4FFABFFBCFE61FA1BFD9CFA6F" box="[430,523,1413,1439]" captionStart="FIGURE 12" captionStartId="18.[151,250,1613,1638]" captionTargetBox="[215,1364,465,1575]" captionTargetId="figure@18.[208,1380,453,1588]" captionTargetPageId="18" captionText="FIGURE 12. Thubdora umbratilis Park, sp. nov.: A, adult, holotype; B, head and antenna, dorsal view; C, forewing venation; D, male genitalia, holotype, gen. slide no. CIS-7048, E, lateral view, holotype; F, aedeagus; G, abdominal segments VI-VIII. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3740973" httpUri="https://zenodo.org/record/3740973/files/figure.png" pageId="17" pageNumber="468">Figs. 12</figureCitation>
AC). Forewing length
<quantity id="4C857DA4FFABFFBCFCD0FA1AFCECFA6F" box="[799,891,1412,1439]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="17" pageNumber="468" unit="mm" value="6.5">6.5 mm</quantity>
.
<emphasis id="B9090C53FFABFFBCFC47FA1BFC52FA6F" box="[904,965,1413,1439]" italics="true" pageId="17" pageNumber="468">Head</emphasis>
: Vertex covered with dark-brown scales, with orange-white, erect scales laterally. Antenna shorter than forewing; basal segment elongated, slightly dilated distally, orange white; flagellum serrate with fine cilia ventrally, often with dark-brown annulations (
<figureCitation id="1346CCC4FFABFFBCFB31FA53FAC0FA17" box="[1278,1367,1485,1511]" captionStart="FIGURE 4" captionStartId="6.[151,250,1401,1426]" captionTargetBox="[250,1332,190,1361]" captionTargetId="figure@6.[243,1345,181,1376]" captionTargetPageId="6" captionText="FIGURE 4. Thubdora ealaensis Park &amp; De Prins, 2019:A, adult; B, head part with labial palpus, lateral view; C, male genitalia, gen. slide no. CIS-7274; D, aedeagus; E, abdominal segment VIII; F, female genitalia, gen. slide no. CIS-7010 (with a close-up of signum). Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3740957" httpUri="https://zenodo.org/record/3740957/files/figure.png" pageId="17" pageNumber="468">Fig. 4B</figureCitation>
). Second segment of labial palpus thickened, saber-shaped, yellowish brown, speckled with irregularly scattered, darkbrown scales; 3
<superScript id="7C087D09FFABFFBCFE8AF98AFEC4F9D2" attach="left" box="[325,339,1556,1570]" fontSize="6" pageId="17" pageNumber="468">rd</superScript>
segment as long as 2
<superScript id="7C087D09FFABFFBCFD8DF98AFDC5F9D2" attach="left" box="[578,594,1556,1570]" fontSize="6" pageId="17" pageNumber="468">nd</superScript>
segment, brown, acute apically.
</paragraph>
<paragraph id="8BC2D041FFABFFBCFF08F9A7FB50F8DB" blockId="17.[151,1437,1232,2015]" pageId="17" pageNumber="468">
<emphasis id="B9090C53FFABFFBCFF08F9A7FE8FF9A3" box="[199,280,1593,1619]" italics="true" pageId="17" pageNumber="468">Thorax</emphasis>
: Notum and tegula dark brown. Forewing ground color dark brown uniformly; antemedian fascia large, usually well developed, with black scales; postmedian fascia large, rectangular, black; costa slightly arched in basal 1/3 and beyond 2/3, with a small, crescent, yellowish costal patch beyond 3/4; apex obtuse; termen slightly oblique; fringe concolorous with ground color, with narrow, orange-white basal line; venation (
<figureCitation id="1346CCC4FFABFFBCFBB8F93BFB47F94F" box="[1143,1232,1701,1727]" captionStart="FIGURE 4" captionStartId="6.[151,250,1401,1426]" captionTargetBox="[250,1332,190,1361]" captionTargetId="figure@6.[243,1345,181,1376]" captionTargetPageId="6" captionText="FIGURE 4. Thubdora ealaensis Park &amp; De Prins, 2019:A, adult; B, head part with labial palpus, lateral view; C, male genitalia, gen. slide no. CIS-7274; D, aedeagus; E, abdominal segment VIII; F, female genitalia, gen. slide no. CIS-7010 (with a close-up of signum). Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3740957" httpUri="https://zenodo.org/record/3740957/files/figure.png" pageId="17" pageNumber="468">Fig. 4C</figureCitation>
) with R
<subScript id="17F9D204FFABFFBCFAE0F928FAA0F934" attach="left" box="[1327,1335,1718,1732]" fontSize="6" pageId="17" pageNumber="468">1</subScript>
strongly bent outwardly near basal 1/3, R
<subScript id="17F9D204FFABFFBCFDCBF944FD9BF918" attach="left" box="[516,524,1754,1768]" fontSize="6" pageId="17" pageNumber="468">3</subScript>
and R
<subScript id="17F9D204FFABFFBCFD9AF944FDCAF918" attach="left" box="[597,605,1754,1768]" fontSize="6" pageId="17" pageNumber="468">4</subScript>
stalked for basal 2/5, R
<subScript id="17F9D204FFABFFBCFCA7F944FCE7F918" attach="left" box="[872,880,1754,1768]" fontSize="6" pageId="17" pageNumber="468">4</subScript>
reaching before apex; R
<subScript id="17F9D204FFABFFBCFB4BF944FB1BF918" attach="left" box="[1156,1164,1754,1768]" fontSize="6" pageId="17" pageNumber="468">5</subScript>
absent; M
<subScript id="17F9D204FFABFFBCFACCF944FA9CF918" attach="left" box="[1283,1291,1754,1768]" fontSize="6" pageId="17" pageNumber="468">1</subScript>
close to R
<subScript id="17F9D204FFABFFBCFA4CF944FA0AF918" attach="left" box="[1411,1437,1754,1768]" fontSize="6" pageId="17" pageNumber="468">3+4</subScript>
at base; M
<subScript id="17F9D204FFABFFBCFEC1F960FE81F8FC" attach="left" box="[270,278,1790,1804]" fontSize="6" pageId="17" pageNumber="468">2</subScript>
arising from lower corner of cell; M
<subScript id="17F9D204FFABFFBCFD74F960FD54F8FC" attach="left" box="[699,707,1790,1804]" fontSize="6" pageId="17" pageNumber="468">3</subScript>
parallel to M
<subScript id="17F9D204FFABFFBCFC91F960FCF1F8FC" attach="left" box="[862,870,1790,1804]" fontSize="6" pageId="17" pageNumber="468">2</subScript>
; M
<subScript id="17F9D204FFABFFBCFC40F960FC00F8FC" attach="left" box="[911,919,1790,1804]" fontSize="6" pageId="17" pageNumber="468">3</subScript>
, CuA
<subScript id="17F9D204FFABFFBCFC13F960FC73F8FC" attach="left" box="[988,996,1790,1804]" fontSize="6" pageId="17" pageNumber="468">1</subScript>
and CuA
<subScript id="17F9D204FFABFFBCFB9DF960FBCDF8FC" attach="left" box="[1106,1114,1790,1804]" fontSize="6" pageId="17" pageNumber="468">2</subScript>
on a common stalk, 1A+2A shortly forked at base. Hind wing pale yellowish brown, with a bundle of hair-scale tuft at base.
</paragraph>
<paragraph id="8BC2D041FFABFFBFFF08F8ABFC91FEEF" blockId="17.[151,1437,1232,2015]" lastBlockId="18.[151,1437,153,431]" lastPageId="18" lastPageNumber="469" pageId="17" pageNumber="468">
<emphasis id="B9090C53FFABFFBCFF08F8ABFEA6F8BF" box="[199,305,1845,1871]" italics="true" pageId="17" pageNumber="468">Abdomen</emphasis>
(
<figureCitation id="1346CCC4FFABFFBCFE8DF8ABFE39F8BF" box="[322,430,1845,1871]" captionStart="FIGURE 12" captionStartId="18.[151,250,1613,1638]" captionTargetBox="[215,1364,465,1575]" captionTargetId="figure@18.[208,1380,453,1588]" captionTargetPageId="18" captionText="FIGURE 12. Thubdora umbratilis Park, sp. nov.: A, adult, holotype; B, head and antenna, dorsal view; C, forewing venation; D, male genitalia, holotype, gen. slide no. CIS-7048, E, lateral view, holotype; F, aedeagus; G, abdominal segments VI-VIII. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3740973" httpUri="https://zenodo.org/record/3740973/files/figure.png" pageId="17" pageNumber="468">Fig. 12G</figureCitation>
): Spinous zones on tergites elongated longitudinally; tergite VIII heavily sclerotized, Yshaped, with a few spines. Male genitalia (
<figureCitation id="1346CCC4FFABFFBCFDA5F8C7FD54F883" box="[618,707,1881,1907]" captionStart="FIGURE 12" captionStartId="18.[151,250,1613,1638]" captionTargetBox="[215,1364,465,1575]" captionTargetId="figure@18.[208,1380,453,1588]" captionTargetPageId="18" captionText="FIGURE 12. Thubdora umbratilis Park, sp. nov.: A, adult, holotype; B, head and antenna, dorsal view; C, forewing venation; D, male genitalia, holotype, gen. slide no. CIS-7048, E, lateral view, holotype; F, aedeagus; G, abdominal segments VI-VIII. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3740973" httpUri="https://zenodo.org/record/3740973/files/figure.png" pageId="17" pageNumber="468">Figs. 12</figureCitation>
DF): Uncus broadened distally, as wide as basal plate of gnathos, nearly truncate caudal margin. Basal plate of gnathos rounded on caudal margin; median process strongly curved downward beyond 2/3, with acute apex. Tegumen weakly sclerotized, with inverted V-shaped incision anteriorly. Valva elongated, slightly expanded anteriorly in basal 1/4 of costa; cucullus elongated, densely setose, upturned, narrowed toward apex from middle, arched before middle on ventral margin. Juxta lacerated, indiscernible. Vinculum band-like, with nearly flat margin anteriorly. Aedeagus about 4/5 the length of valva, nearly straight, but with wavy marginal line due to variable width; cornuti consisting of a large, heavily sclerotized, dagger-like plate with hooked tip distally, and a row of short bristles medially.
</paragraph>
<paragraph id="8BC2D041FFA8FFBFFF08FEB7FE06FEB3" blockId="18.[151,1437,153,431]" box="[199,401,297,323]" pageId="18" pageNumber="469">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C36783CAFFA8FFBFFF08FED2FDFEFE98" box="[199,617,332,360]" pageId="18" pageNumber="469" type="distribution">
<paragraph id="8BC2D041FFA8FFBFFF08FED2FDFEFE98" blockId="18.[151,1437,153,431]" box="[199,617,332,360]" pageId="18" pageNumber="469">
<emphasis id="B9090C53FFA8FFBFFF08FED2FEF6FE97" bold="true" box="[199,353,332,359]" pageId="18" pageNumber="469">Distribution.</emphasis>
<collectingCountry id="F36A90D1FFA8FFBFFEA8FED3FE29FE97" box="[359,446,333,359]" name="Uganda" pageId="18" pageNumber="469">Uganda</collectingCountry>
(
<collectingRegion id="49B91EA3FFA8FFBFFE02FED3FDCAFE98" box="[461,605,333,360]" country="Uganda" name="Mpigi" pageId="18" pageNumber="469">Mpigi Distr.</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36783CAFFA8FFBFFF08FEEFFDB3FE5F" pageId="18" pageNumber="469" type="etymology">
<paragraph id="8BC2D041FFA8FFBFFF08FEEFFDB3FE5F" blockId="18.[151,1437,153,431]" pageId="18" pageNumber="469">
<emphasis id="B9090C53FFA8FFBFFF08FEEFFED9FE7B" bold="true" box="[199,334,369,395]" pageId="18" pageNumber="469">Etymology.</emphasis>
This species name is derived from Latin
<emphasis id="B9090C53FFA8FFBFFCD3FEEFFCF2FE7B" box="[796,869,369,395]" italics="true" pageId="18" pageNumber="469">umbra</emphasis>
(= shade), with Latin adjectival suffix
<emphasis id="B9090C53FFA8FFBFFAD6FEEFFAC6FE7B" box="[1305,1361,369,395]" italics="true" pageId="18" pageNumber="469">atilis</emphasis>
, referring the fuscous patch on forewing.
</paragraph>
</subSubSection>
</treatment>
</document>