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<document id="C2A02E592F5F06B3DAB5A8FA987CBB3E" ID-DOI="10.11646/phytotaxa.257.3.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13670670" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725398280652" checkinUser="felipe" docAuthor="Doucette, Alfonso, Portilla, Jose &amp; Cameron, Kenneth M." docDate="2016" docId="03B987A5BA50346BA7963C7951AEFD8A" docLanguage="en" docName="phytotaxa.257.3.2.pdf" docOrigin="Phytotaxa 257 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.257.3.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Pleurothallis cardiostola fm. magnidraba A. Doucette &amp; J. Portilla 2016, f. nov." docType="treatment" docVersion="2" lastPageNumber="238" masterDocId="FF80FFDDBA553463A71E3A395174FFE3" masterDocTitle="Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador" masterLastPageNumber="248" masterPageNumber="230" pageNumber="235" updateTime="1725420285918" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A269E46F91B10FC1DC23F74FB18E461B">Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador</mods:title>
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<mods:namePart id="C2EB87E5D81110C838D52933509BFE1D">Doucette, Alfonso</mods:namePart>
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<mods:namePart id="4E358E7E5395EF25ECB2FCA2C9FE532D">Portilla, Jose</mods:namePart>
<mods:affiliation id="34B6CDECAE638B35EE25CD09AA993867">Km. 2 1 / 1 Vía a Cuenca Sector Llampasay, Ecuagenera, Gualaceo, Ecuador</mods:affiliation>
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<mods:namePart id="DAFA83135E638060C88AE756B4A0A875">Cameron, Kenneth M.</mods:namePart>
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<mods:title id="353C6E3B6B6AE1166256D5805B4B2570">Phytotaxa</mods:title>
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<mods:date id="74F1D16B6494BBECD2A4EEE6233F2B27">2016</mods:date>
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<mods:number id="C8A2FC53DFE31EA6F5DBEB7CF1AA9A15">2016-04-20</mods:number>
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<treatment id="03B987A5BA50346BA7963C7951AEFD8A" ID-DOI="http://doi.org/10.5281/zenodo.13670578" ID-Zenodo-Dep="13670578" LSID="urn:lsid:plazi:treatment:03B987A5BA50346BA7963C7951AEFD8A" httpUri="http://treatment.plazi.org/id/03B987A5BA50346BA7963C7951AEFD8A" lastPageId="8" lastPageNumber="238" pageId="5" pageNumber="235">
<subSubSection id="C30A6538BA503466A7963C7955FEF9BA" box="[136,1162,1600,1625]" pageId="5" pageNumber="235" type="nomenclature">
<paragraph id="8BAF36B3BA503466A7963C7955FEF9BA" blockId="5.[136,1162,1600,1625]" box="[136,1162,1600,1625]" pageId="5" pageNumber="235">
<heading id="D0E781DFBA503466A7963C7955FEF9BA" box="[136,1162,1600,1625]" fontSize="11" level="2" pageId="5" pageNumber="235" reason="3">
<taxonomicName id="4C104D30BA503466A7963C7952EEF9BA" ID-CoL="7Z45T" authority="A.Doucette &amp; J.Portilla" authorityName="A. Doucette &amp; J. Portilla" authorityYear="2016" box="[136,922,1600,1625]" class="Liliopsida" family="Orchidaceae" form="magnidraba" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="235" phylum="Tracheophyta" rank="form" species="cardiostola" status="f. nov.">
<emphasis id="B964EAA1BA503466A7963C7953F3F9BA" bold="true" box="[136,647,1600,1625]" pageId="5" pageNumber="235">
<emphasis id="B964EAA1BA503466A7963C7950D0F9BA" bold="true" box="[136,420,1600,1625]" italics="true" pageId="5" pageNumber="235">Pleurothallis cardiostola</emphasis>
forma
<emphasis id="B964EAA1BA503466A6E53C7953F3F9BA" bold="true" box="[507,647,1600,1625]" italics="true" pageId="5" pageNumber="235">magnidraba</emphasis>
</emphasis>
A.Doucette &amp; J.Portilla
</taxonomicName>
,
<emphasis id="B964EAA1BA503466A4BA3C79529CF9BA" box="[932,1000,1600,1625]" italics="true" pageId="5" pageNumber="235">
<taxonomicNameLabel id="A25757DABA503466A4BA3C79529CF9BA" box="[932,1000,1600,1625]" pageId="5" pageNumber="235" rank="form">f. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132B2A36BA503466A4E93C795532F9BA" box="[1015,1094,1600,1625]" captionStart="FIGURE 5" captionStartId="7.[136,229,1781,1802]" captionTargetBox="[264,1321,216,1740]" captionTargetId="figure-14@7.[232,1355,190,1756]" captionTargetPageId="7" captionText="FIGURE 5. Pleurothallis cardiostola f. magnidraba. A. Habit, drawn to 5 cm scale. B. Flower. C. Dorsal sepal. D. Lateral sepals. E. Petal. F. Lip. G. Flower with petals and sepals removed. All drawn to 1 mm scale." figureDoi="http://doi.org/10.5281/zenodo.13670688" httpUri="https://zenodo.org/record/13670688/files/figure.png" pageId="5" pageNumber="235">Figs. 5</figureCitation>
,
<figureCitation id="132B2A36BA503466A34C3C7955F6F9BA" box="[1106,1154,1600,1625]" captionStart="FIGURE 11" captionStartId="15.[136,229,1891,1912]" captionTargetBox="[208,1380,202,1851]" captionTargetId="figure-14@15.[197,1391,190,1866]" captionTargetPageId="15" captionText="FIGURE 11. A. Masdevallia constricta f. glabra. B. Masdevallia pachyura f. pardinella. C. Masdevallia posadae f. rugosepala. D. Acianthera nikoleana. E. Pleurothallis papillingua. F. Pleurothallis gigiportillae. G. Pleurothallis cardiostola f. magnidraba. H. Zootrophion machaqway. I. Stelis glaucus. J. Trisetella portillae." figureDoi="http://doi.org/10.5281/zenodo.13670708" httpUri="https://zenodo.org/record/13670708/files/figure.png" pageId="5" pageNumber="235">11G</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A6538BA503465A7963CB354E7F7E6" lastPageId="6" lastPageNumber="236" pageId="5" pageNumber="235" type="materials_examined">
<paragraph id="8BAF36B3BA503466A7963CB3502CF920" blockId="5.[136,1451,1674,1803]" pageId="5" pageNumber="235">
Type:—
<collectingCountry id="F3077623BA503466A7CB3CB35039F97C" box="[213,333,1674,1695]" name="Ecuador" pageId="5" pageNumber="235">ECUADOR</collectingCountry>
. Without a specific locality, flowered in cultivation at Ecuagenera Orchid Nursery,
<date id="FFAE1073BA503466A3793CB35463F97C" box="[1127,1303,1674,1695]" pageId="5" pageNumber="235" value="2015-09-04">4 September 2015</date>
,
<emphasis id="B964EAA1BA503466A23F3CB354DFF97C" box="[1313,1451,1674,1695]" italics="true" pageId="5" pageNumber="235">Doucette 0311</emphasis>
(
<typeStatus id="54AB8811BA503466A7DA3C975069F920" box="[196,285,1710,1731]" pageId="5" pageNumber="235" type="holotype">holotype</typeStatus>
:
<collectionCode id="ED01AE76BA503466A6383C975039F920" box="[294,333,1710,1731]" country="Ecuador" lsid="urn:lsid:biocol.org:col:14290" name="Universidad del Azuay" pageId="5" pageNumber="235" type="Herbarium">HA</collectionCode>
).
</paragraph>
<paragraph id="8BAF36B3BA503466A7963CEB50DFF8E8" blockId="5.[136,1451,1674,1803]" pageId="5" pageNumber="235">
Diagnosis: The new
<taxonomicName id="4C104D30BA503466A6523CEB50CBF904" box="[332,447,1746,1767]" form="differs" pageId="5" pageNumber="235" rank="form">form differs</taxonomicName>
from
<taxonomicName id="4C104D30BA503466A6E73CEB5510F904" authority="f. cardiostola Reichenbach (1854: 128)" authorityName="Reichenbach" authorityPageNumber="128" authorityYear="1854" box="[505,1124,1746,1767]" class="Liliopsida" family="Orchidaceae" form="cardiostola" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="235" phylum="Tracheophyta" rank="form" species="cardiostola">
<emphasis id="B964EAA1BA503466A6E73CEB5393F904" box="[505,743,1746,1767]" italics="true" pageId="5" pageNumber="235">Pleurothallis cardiostola</emphasis>
f.
<emphasis id="B964EAA1BA503466A41F3CEB5219F904" box="[769,877,1746,1767]" italics="true" pageId="5" pageNumber="235">cardiostola</emphasis>
Reichenbach (1854: 128)
</taxonomicName>
in its larger flower and number of veins in the dorsal sepal.
</paragraph>
<caption id="DF6F663BBA533465A7963C935233F900" ID-DOI="http://doi.org/10.5281/zenodo.13670684" ID-Zenodo-Dep="13670684" httpUri="https://zenodo.org/record/13670684/files/figure.png" pageId="6" pageNumber="236" startId="6.[136,229,1706,1727]" targetBox="[269,1325,213,1654]" targetPageId="6" targetType="figure">
<paragraph id="8BAF36B3BA533465A7963C935233F900" blockId="6.[136,1452,1706,1763]" pageId="6" pageNumber="236">
<emphasis id="B964EAA1BA533465A7963C93518AF95C" bold="true" box="[136,254,1706,1727]" pageId="6" pageNumber="236">FIGURE 4.</emphasis>
<taxonomicName id="4C104D30BA533465A61A3C935323F95C" authorityName="A. Doucette &amp; J. Portilla" authorityYear="2016" box="[260,599,1706,1727]" class="Liliopsida" family="Orchidaceae" form="rugosepala" genus="Masdevallia" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="236" phylum="Tracheophyta" rank="form" species="posadae">
<emphasis id="B964EAA1BA533465A61A3C9350A5F95C" box="[260,465,1706,1727]" italics="true" pageId="6" pageNumber="236">Masdevallia posadae</emphasis>
f.
<emphasis id="B964EAA1BA533465A6F53C935323F95C" box="[491,599,1706,1727]" italics="true" pageId="6" pageNumber="236">rugosepala</emphasis>
</taxonomicName>
. A. Habit, drawn to 5cm scale. B. Flower. C. Dorsal sepal. D. Lateral sepals. E. Petal. F. Lip. G. Flower with petals and sepals removed. All drawn to 1 mm scale.
</paragraph>
</caption>
<paragraph id="8BAF36B3BA533465A7963D2D54E7F7E6" blockId="6.[136,1452,1812,2053]" pageId="6" pageNumber="236">
Epiphytic, caespitose herbs; roots
<quantity id="4CE89B56BA533465A50F3D2D5320F8CE" box="[529,596,1812,1837]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="236" unit="mm" value="1.0">1 mm</quantity>
in diam. The stem 180–260 5
<quantity id="4CE89B56BA533465A4A73D2D5567F8CE" box="[953,1043,1812,1837]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="236" unit="mm" value="2.5">2.5 mm</quantity>
, enveloped by two to three tubular, papery sheaths 10–60 5
<quantity id="4CE89B56BA533465A68D3D01509FF8B2" box="[403,491,1848,1873]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="236" unit="mm" value="2.5">2.5 mm</quantity>
. Leaves sessile, cordate, 130–192 5
<quantity id="4CE89B56BA533465A4653D015282F8B2" box="[891,1014,1848,1873]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="6" pageNumber="236" unit="mm" value="25.0" valueMax="28.0" valueMin="22.0">22–28 mm</quantity>
. Inflorescence fasciculate, protected by an erect, papery sheath,
<quantity id="4CE89B56BA533465A6883D655091F896" box="[406,485,1884,1909]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="236" unit="mm" value="18.0">18 mm</quantity>
long; peduncle 3 5
<quantity id="4CE89B56BA533465A5DF3D655270F896" box="[705,772,1884,1909]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="236" unit="mm" value="1.0">1 mm</quantity>
, with a tubular bract before the pedicel, 1–2 5
<quantity id="4CE89B56BA533465A2123D65543BF896" box="[1292,1359,1884,1909]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="236" unit="mm" value="2.0">2 mm</quantity>
; pedicel 10 5
<quantity id="4CE89B56BA533465A7DD3DB9507CF87A" box="[195,264,1920,1945]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="236" unit="mm" value="1.0">1 mm</quantity>
; ovary papillose, 7 5
<quantity id="4CE89B56BA533465A6E33DB95335F87A" box="[509,577,1920,1945]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="236" unit="mm" value="2.0">2 mm</quantity>
, flowers non-resupinate, dorsal sepal broadly ovate, 11-veined, tan, minutely pubescent, 20 5
<quantity id="4CE89B56BA533465A6253D9D50F8F85E" box="[315,396,1956,1981]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="236" unit="mm" value="15.0">15 mm</quantity>
; synsepalum round, concave below the middle, similar in colour and texture to the dorsal sepal, 14 5
<quantity id="4CE89B56BA533465A7DF3DF15067F802" box="[193,275,1992,2017]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="236" unit="mm" value="15.0">15 mm</quantity>
; petals lanceolate, similar in colour and texture to the sepals, 14 5
<quantity id="4CE89B56BA533465A31E3DF15531F802" box="[1024,1093,1992,2017]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="236" unit="mm" value="5.0">5 mm</quantity>
; lip white suffused with yellow, shiny, heater-shaped, 5 5
<quantity id="4CE89B56BA533465A6BA3DD5509CF7E6" box="[420,488,2028,2053]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="236" unit="mm" value="4.0">4 mm</quantity>
; column white suffused with yellow, with a shiny yellow bilobed stigma, 3 5
<quantity id="4CE89B56BA533465A2553DD554FBF7E6" box="[1355,1423,2028,2053]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="236" unit="mm" value="3.0">3 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF6F663BBA523464A7963CCC522BF8CD" ID-DOI="http://doi.org/10.5281/zenodo.13670688" ID-Zenodo-Dep="13670688" httpUri="https://zenodo.org/record/13670688/files/figure.png" pageId="7" pageNumber="237" startId="7.[136,229,1781,1802]" targetBox="[264,1321,216,1740]" targetPageId="7" targetType="figure">
<paragraph id="8BAF36B3BA523464A7963CCC522BF8CD" blockId="7.[136,1453,1781,1838]" pageId="7" pageNumber="237">
<emphasis id="B964EAA1BA523464A7963CCC5189F8E9" bold="true" box="[136,253,1781,1802]" pageId="7" pageNumber="237">FIGURE 5.</emphasis>
<taxonomicName id="4C104D30BA523464A61C3CCC5309F8E9" authorityName="A. Doucette &amp; J. Portilla" authorityYear="2016" box="[258,637,1781,1802]" class="Liliopsida" family="Orchidaceae" form="magnidraba" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="237" phylum="Tracheophyta" rank="form" species="cardiostola">
<emphasis id="B964EAA1BA523464A61C3CCC509BF8E9" box="[258,495,1781,1802]" italics="true" pageId="7" pageNumber="237">Pleurothallis cardiostola</emphasis>
f.
<emphasis id="B964EAA1BA523464A5163CCC5309F8E9" box="[520,637,1781,1802]" italics="true" pageId="7" pageNumber="237">magnidraba</emphasis>
</taxonomicName>
. A. Habit, drawn to 5 cm scale. B. Flower. C. Dorsal sepal. D. Lateral sepals. E. Petal. F. Lip. G. Flower with petals and sepals removed. All drawn to 1 mm scale.
</paragraph>
</caption>
<subSubSection id="C30A6538BA52346BA7963D6751AEFD8A" lastPageId="8" lastPageNumber="238" pageId="7" pageNumber="237" type="etymology">
<paragraph id="8BAF36B3BA523464A7963D6750FFF878" blockId="7.[136,1452,1886,2091]" pageId="7" pageNumber="237">
<emphasis id="B964EAA1BA523464A7963D67505AF894" bold="true" box="[136,302,1886,1911]" pageId="7" pageNumber="237">Etymology:—</emphasis>
From the Latin
<emphasis id="B964EAA1BA523464A6C33D67535AF894" box="[477,558,1886,1911]" italics="true" pageId="7" pageNumber="237">magnis</emphasis>
meaning “large” and the English “drab” meaning a dull brown colour, referring to the large dull sepals.
</paragraph>
<paragraph id="8BAF36B3BA52346BA7A33D9F52DBFF3E" blockId="7.[136,1452,1886,2091]" lastBlockId="8.[136,1452,160,617]" lastPageId="8" lastPageNumber="238" pageId="7" pageNumber="237">
<emphasis id="B964EAA1BA523464A7A33D9F5052F85C" bold="true" box="[189,294,1958,1983]" pageId="7" pageNumber="237">Notes:—</emphasis>
The genus
<taxonomicName id="4C104D30BA523464A6BB3D9F5342F85C" box="[421,566,1958,1983]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="237" phylum="Tracheophyta" rank="genus">
<emphasis id="B964EAA1BA523464A6BB3D9F5342F85C" box="[421,566,1958,1983]" italics="true" pageId="7" pageNumber="237">Pleurothallis</emphasis>
</taxonomicName>
is one of the largest plant genera including over 500 species (The Plant List 2015). Phylogenetic analyses confirmed the genus to be non-monophyletic (Pridgeon
<emphasis id="B964EAA1BA523464A37C3DF355E9F800" box="[1122,1181,1994,2019]" italics="true" pageId="7" pageNumber="237">et al.</emphasis>
2001). We hypothesize the new
<taxonomicName id="4C104D30BA523464A7F13DD750EFF7E4" box="[239,411,2030,2055]" form="attributed" pageId="7" pageNumber="237" rank="form">form attributed</taxonomicName>
to
<taxonomicName id="4C104D30BA523464A6D93DD75396F7E4" box="[455,738,2030,2055]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="237" phylum="Tracheophyta" rank="species" species="cardiostola">
<emphasis id="B964EAA1BA523464A6D93DD75396F7E4" box="[455,738,2030,2055]" italics="true" pageId="7" pageNumber="237">Pleurothallis cardiostola</emphasis>
</taxonomicName>
to belong to
<taxonomicName id="4C104D30BA523464A49B3DD75562F7E4" box="[901,1046,2030,2055]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="237" phylum="Tracheophyta" rank="genus" sensu="stricto">
<emphasis id="B964EAA1BA523464A49B3DD75562F7E4" box="[901,1046,2030,2055]" italics="true" pageId="7" pageNumber="237">Pleurothallis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757DABA523464A33C3DD75531F7E4" box="[1058,1093,2030,2055]" pageId="7" pageNumber="237" sensu="stricto">s.s.</taxonomicNameLabel>
based on morphology and the placement of
<taxonomicName id="4C104D30BA523464A63E322B5231F7C8" authority="Reichenbach (1876: 15)" authorityName="Reichenbach" authorityPageNumber="15" authorityYear="1876" box="[288,837,2066,2091]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="237" phylum="Tracheophyta" rank="species" species="cardiantha">
<emphasis id="B964EAA1BA523464A63E322B5346F7C8" box="[288,562,2066,2091]" italics="true" pageId="7" pageNumber="237">Pleurothallis cardiantha</emphasis>
Reichenbach (1876: 15)
</taxonomicName>
sister to the
<typeStatus id="54AB8811BA523464A4CA322B5570F7C8" box="[980,1028,2066,2091]" pageId="7" pageNumber="237">type</typeStatus>
<taxonomicName id="4C104D30BA523464A314322B54D8F7C8" authority="(Jacquin 1760: 29) R. Br." authorityName="R. Br." baseAuthorityName="Jacquin" baseAuthorityPageNumber="29" baseAuthorityYear="1760" box="[1034,1452,2066,2091]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="237" phylum="Tracheophyta" rank="species" species="ruscifolia">
<emphasis id="B964EAA1BA523464A314322B55E5F7C8" box="[1034,1169,2066,2091]" italics="true" pageId="7" pageNumber="237">P. ruscifolia</emphasis>
(
<bibRefCitation id="EF814B42BA523464A381322B5415F7C8" author="Jacquin, N." box="[1183,1377,2066,2091]" pageId="7" pageNumber="237" refId="ref7791" refString="Jacquin, N. (1760) Enumeratio systematica plantarum, quas in insulis Caribaeis vicinaque Americes. Haak, New York, 41 pp." type="book" year="1760">Jacquin 1760: 29</bibRefCitation>
) R.Br.
</taxonomicName>
in W.T.
<bibRefCitation id="EF814B42BA5D346BA7C83A9950EBFF5A" author="Aiton, W. T." box="[214,415,160,185]" pageId="8" pageNumber="238" refId="ref7425" refString="Aiton, W. T. (1813) Hortus Kewensis, volume 5. Longman, Hurst, Rees, Orme, and Brown, London, 568 pp." type="book" year="1813">Aiton (1813: 211)</bibRefCitation>
with 100% bootstrap support based on a combined data set including two plastid genes (
<emphasis id="B964EAA1BA5D346BA2713A9954D8FF5A" box="[1391,1452,160,185]" italics="true" pageId="8" pageNumber="238">matK</emphasis>
and
<emphasis id="B964EAA1BA5D346BA7A93AFD5076FF3E" box="[183,258,196,221]" italics="true" pageId="8" pageNumber="238">trnL-F</emphasis>
) and a nuclear ribosomal gene (nrITS; Pridgeon
<emphasis id="B964EAA1BA5D346BA4393AFD522CFF3E" box="[807,856,196,221]" italics="true" pageId="8" pageNumber="238">et al</emphasis>
. 2001).
</paragraph>
<paragraph id="8BAF36B3BA5D346BA7A33AD151A9FE72" blockId="8.[136,1452,160,617]" pageId="8" pageNumber="238">
Although there are morphological characters that easily distinguish the taxon described here from typical
<taxonomicName id="4C104D30BA5D346BA7963B3550D6FEC6" box="[136,418,268,293]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="238" phylum="Tracheophyta" rank="species" species="cardiostola">
<emphasis id="B964EAA1BA5D346BA7963B3550D6FEC6" box="[136,418,268,293]" italics="true" pageId="8" pageNumber="238">Pleurothallis cardiostola</emphasis>
</taxonomicName>
we hypothesize that it represents an aberrant individual embedded in
<taxonomicName id="4C104D30BA5D346BA3CC3B35541BFEC6" box="[1234,1391,268,293]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="238" phylum="Tracheophyta" rank="species" species="cardiostola">
<emphasis id="B964EAA1BA5D346BA3CC3B35541BFEC6" box="[1234,1391,268,293]" italics="true" pageId="8" pageNumber="238">P. cardiostola</emphasis>
</taxonomicName>
. The individual described here differs in traits that have been documented to vary within the species (
<bibRefCitation id="EF814B42BA5D346BA3C33B095423FEAA" author="Luer, C." box="[1245,1367,304,329]" pageId="8" pageNumber="238" pagination="1 - 311" refId="ref8520" refString="Luer, C. (2005) Icones pleurothallidinarum XVII: Dryadella and Acronia section Macrophyllae-Fasciculatae. Monographs in Systematic Botany 103: 1 - 311." type="journal article" year="2005">Luer 2005</bibRefCitation>
) and is not reported from a locality disjunct from reported populations of
<taxonomicName id="4C104D30BA5D346BA46C3B6D557AFE8E" box="[882,1038,340,365]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="238" phylum="Tracheophyta" rank="species" species="cardiostola">
<emphasis id="B964EAA1BA5D346BA46C3B6D557AFE8E" box="[882,1038,340,365]" italics="true" pageId="8" pageNumber="238">P. cardiostola</emphasis>
</taxonomicName>
(i.e. Caribbean or Atlantic forests of
<collectingCountry id="F3077623BA5D346BA7963B4151BAFE72" box="[136,206,376,401]" name="Brazil" pageId="8" pageNumber="238">Brazil</collectingCountry>
).
</paragraph>
<paragraph id="8BAF36B3BA5D346BA7A33BA551AEFD8A" blockId="8.[136,1452,160,617]" pageId="8" pageNumber="238">
We feel that naming this new form of
<taxonomicName id="4C104D30BA5D346BA5993BA55251FE56" box="[647,805,412,437]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="238" phylum="Tracheophyta" rank="species" species="cardiostola">
<emphasis id="B964EAA1BA5D346BA5993BA55251FE56" box="[647,805,412,437]" italics="true" pageId="8" pageNumber="238">P. cardiostola</emphasis>
</taxonomicName>
brings it to the attention of scientists who can test the taxonomic hypothesis presented here using a molecular systematic approach on individuals representing different populations across the range of
<emphasis id="B964EAA1BA5D346BA6EC3BDD53E1FE1E" box="[498,661,484,509]" italics="true" pageId="8" pageNumber="238">
<taxonomicName id="4C104D30BA5D346BA6EC3BDD53E5FE1E" box="[498,657,484,509]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="238" phylum="Tracheophyta" rank="species" species="cardiostola">P. cardiostola</taxonomicName>
.
</emphasis>
If the taxon described here is not embedded within
<taxonomicName id="4C104D30BA5D346BA3F93BDD54F7FE1E" box="[1255,1411,484,509]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="238" phylum="Tracheophyta" rank="species" species="cardiostola">
<emphasis id="B964EAA1BA5D346BA3F93BDD54F7FE1E" box="[1255,1411,484,509]" italics="true" pageId="8" pageNumber="238">P. cardiostola</emphasis>
</taxonomicName>
we feel that the number of veins of the dorsal sepal and larger flower size are sufficient to warrant recognition at the rank of species. No forms of
<taxonomicName id="4C104D30BA5D346BA6BD381553C9FDA6" box="[419,701,556,581]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="238" phylum="Tracheophyta" rank="species" species="cardiostola">
<emphasis id="B964EAA1BA5D346BA6BD381553C9FDA6" box="[419,701,556,581]" italics="true" pageId="8" pageNumber="238">Pleurothallis cardiostola</emphasis>
</taxonomicName>
are reported by
<emphasis id="B964EAA1BA5D346BA46138155583FDA6" box="[895,1271,556,581]" italics="true" pageId="8" pageNumber="238">
International Plant
<taxonomicName id="4C104D30BA5D346BA37D38155583FDA6" box="[1123,1271,556,581]" class="Liliopsida" family="Orchidaceae" genus="Names" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="238" phylum="Tracheophyta" rank="species" species="index">Names Index</taxonomicName>
</emphasis>
(2016) or
<bibRefCitation id="EF814B42BA5D346BA268381551A2FD8A" author="Luer, C." pageId="8" pageNumber="238" pagination="1 - 311" refId="ref8520" refString="Luer, C. (2005) Icones pleurothallidinarum XVII: Dryadella and Acronia section Macrophyllae-Fasciculatae. Monographs in Systematic Botany 103: 1 - 311." type="journal article" year="2005">Luer (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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