232 lines
37 KiB
XML
232 lines
37 KiB
XML
<document id="3F09D688D3392ABE1B7E087946EDBFA5" ID-DOI="10.5281/zenodo.183126" ID-GBIF-Dataset="5ba6cec4-345d-4f95-8064-dd9dc33dac4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="183126" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459892214864" checkinUser="plazi" docAuthor="Miller, Kelly B. & Spangler, Paul J." docDate="2008" docId="03AD2179AE5BFFC2FF78FD19FD68F8AC" docLanguage="en" docName="zt01827p052.pdf" docOrigin="Zootaxa 1827" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Fontidessus toboganensis Miller and Spangler, new species" docType="treatment" docVersion="9" lastPageNumber="49" masterDocId="FF945901AE58FFC6FFEFFFDBFFA9FFC9" masterDocTitle="Fontidessus Miller and Spangler, a new genus of Bidessini from Venezuela (Coleoptera: Dytiscidae: Hydroporinae) with three new species" masterLastPageNumber="52" masterPageNumber="45" pageNumber="48" updateTime="1698228095007" updateUser="plazi">
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<mods:mods id="3BE7A0505DC96E7D309E2F9F999C6AD5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2152B2E3FAE7B0B561B87E1EDD30137E">
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<mods:title id="EB89CBE680070CE5D8B30ECD3D2FF40C">Fontidessus Miller and Spangler, a new genus of Bidessini from Venezuela (Coleoptera: Dytiscidae: Hydroporinae) with three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="BC3F1E05430921406F3E2B06623A5F8F" type="personal">
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<mods:role id="69741FA2B14E83F639AD563D88921108">
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<mods:roleTerm id="E824593F6664178E93B3876A60468F55">Author</mods:roleTerm>
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<mods:namePart id="1A2EB0E6CA3BD8EF11EC50B7139C0877">Miller, Kelly B.</mods:namePart>
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</mods:name>
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<mods:name id="57F4482D920EB404607D922A1775DD37" type="personal">
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<mods:role id="C7AE62A0FA6AE6112435783B58B87468">
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<mods:roleTerm id="3570F2A733F2772A68F5F37BF117D354">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A84E5B3D47DB5E5D054E8AD8CCA6E705">Spangler, Paul J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="622BACFCAC3FF67FA0A0661591246079">text</mods:typeOfResource>
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<mods:titleInfo id="3A57FF6F86D5AC2E04870FEC7AED5237">
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<mods:title id="1060B4886195F077577388C9F00DFF90">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F678A5F521A462AD7C95B3687DFCC51F">
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<mods:date id="6DC0C5FFFE3C154B8443FFF9AA4AB436">2008</mods:date>
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<mods:detail id="08D7413CE4FAFD372217E22260285C26" type="volume">
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<mods:number id="CC565834A2C24C921F9E79B32A4EBFFA">1827</mods:number>
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<mods:start id="6C62305DC12FA40AA31D94F8F4635A69">45</mods:start>
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<mods:end id="5625575571E06C8163EFF6C339BEA880">52</mods:end>
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<mods:classification id="4937A80BA793BDED3AF47C02B36CD431">journal article</mods:classification>
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<mods:identifier id="A99529FCDA1D788E6F1DDDB8C4082DF7" type="DOI">10.5281/zenodo.183126</mods:identifier>
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<mods:identifier id="7209EBF4E687828025EFD22323CCE893" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="AFBA91C76837DA97C4B9EA16E2B98F2D" type="Zenodo-Dep">183126</mods:identifier>
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<treatment id="03AD2179AE5BFFC2FF78FD19FD68F8AC" ID-DOI="http://doi.org/10.5281/zenodo.6233624" ID-GBIF-Taxon="119372128" ID-Zenodo-Dep="6233624" LSID="urn:lsid:plazi:treatment:03AD2179AE5BFFC2FF78FD19FD68F8AC" httpUri="http://treatment.plazi.org/id/03AD2179AE5BFFC2FF78FD19FD68F8AC" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
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<subSubSection id="C31EC3E4AE5BFFC5FF78FD19FE0BFD36" pageId="3" pageNumber="48" type="nomenclature">
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<paragraph id="8BBB906FAE5BFFC5FF78FD19FCC5FD15" blockId="3.[151,876,706,767]" box="[151,876,706,732]" pageId="3" pageNumber="48">
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<heading id="D0F32703AE5BFFC5FF78FD19FCC5FD15" bold="true" box="[151,876,706,732]" fontSize="11" level="1" pageId="3" pageNumber="48" reason="1">
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<emphasis id="B9704C7DAE5BFFC5FF78FD19FCC5FD15" bold="true" box="[151,876,706,732]" pageId="3" pageNumber="48">
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<taxonomicName id="4C04EBECAE5BFFC5FF78FD19FD66FD15" ID-CoL="6JDKT" authority="Miller and Spangler" authorityName="Miller and Spangler" box="[151,719,706,732]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="toboganensis" status="sp. nov.">
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<emphasis id="B9704C7DAE5BFFC5FF78FD19FE64FD15" bold="true" box="[151,461,706,732]" italics="true" pageId="3" pageNumber="48">Fontidessus toboganensis</emphasis>
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Miller and Spangler
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</taxonomicName>
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,
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<taxonomicNameLabel id="A243F106AE5BFFC5FD33FD19FCC5FD15" box="[732,876,706,732]" pageId="3" pageNumber="48" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBB906FAE5BFFC5FF78FD3EFE0BFD36" blockId="3.[151,876,706,767]" box="[151,418,741,767]" pageId="3" pageNumber="48">
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(
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<figureCitation id="133F8CEAAE5BFFC5FF70FD3EFE8CFD36" box="[159,293,741,767]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="3" pageNumber="48">Figs 1, 4, 5</figureCitation>
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,
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<figureCitation id="133F8CEAAE5BFFC5FEDDFD3EFE30FD36" box="[306,409,741,767]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="3" pageNumber="48">8–10, 17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31EC3E4AE5BFFC5FF78FCEEFA3CFC86" box="[151,1429,821,847]" pageId="3" pageNumber="48" type="materials_examined">
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<paragraph id="8BBB906FAE5BFFC5FF78FCEEFA3CFC86" blockId="3.[151,1437,821,2007]" box="[151,1429,821,847]" pageId="3" pageNumber="48">
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<emphasis id="B9704C7DAE5BFFC5FF78FCEEFE94FC86" bold="true" box="[151,317,821,847]" pageId="3" pageNumber="48">
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<typeStatus id="54BF2ECDAE5BFFC5FF78FCEEFF7DFC86" box="[151,212,821,847]" pageId="3" pageNumber="48">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F313D0FFAE5BFFC5FEABFCEEFE16FC86" box="[324,447,821,847]" name="Venezuela" pageId="3" pageNumber="48">Venezuela</collectingCountry>
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, Territorio Federal Amazonas,
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<quantity id="4CFC3D8AAE5BFFC5FCC4FCEEFCC7FC86" box="[811,878,821,847]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="48" unit="km" value="40.0">40km</quantity>
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S Puerto Ayacucho, at El Tobogán, Coromoto.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31EC3E4AE5BFFC5FF2AFC86FC54FC26" pageId="3" pageNumber="48" type="diagnosis">
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<paragraph id="8BBB906FAE5BFFC5FF2AFC86FC54FC26" blockId="3.[151,1437,821,2007]" pageId="3" pageNumber="48">
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<emphasis id="B9704C7DAE5BFFC5FF2AFC86FEECFCBE" bold="true" box="[197,325,861,887]" pageId="3" pageNumber="48">Diagnosis.</emphasis>
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This is the largest known species in the genus (TL>
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<quantity id="4CFC3D8AAE5BFFC5FC5CFC86FBA2FCBE" box="[947,1035,861,887]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="48" unit="mm" value="1.4">1.4mm</quantity>
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). The prosternal process is moderately broad and parallel-sided and has the apex broadly truncate (
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<figureCitation id="133F8CEAAE5BFFC5FC45FC5EFC5DFC56" box="[938,1012,901,927]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="3" pageNumber="48">Fig. 5</figureCitation>
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). The male genitalia are distinctive with the median lobe elongate, slender and broadly recurved in lateral aspect (
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<figureCitation id="133F8CEAAE5BFFC5FBA9FC76FB39FC0E" box="[1094,1168,941,967]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="3" pageNumber="48">Fig. 9</figureCitation>
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). There is an elongate ventral sclerite that fits into the ventral groove of the median lobe (
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<figureCitation id="133F8CEAAE5BFFC5FC48FC0EFC46FC26" box="[935,1007,981,1007]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="3" pageNumber="48">Fig. 9</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31EC3E4AE5BFFC2FF2AFC26FF6CFEE3" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48" type="description">
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<paragraph id="8BBB906FAE5BFFC5FF2AFC26FDE9FB46" blockId="3.[151,1437,821,2007]" pageId="3" pageNumber="48">
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<emphasis id="B9704C7DAE5BFFC5FF2AFC26FEF4FBDE" bold="true" box="[197,349,1021,1047]" pageId="3" pageNumber="48">Description.</emphasis>
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<emphasis id="B9704C7DAE5BFFC5FE88FC26FDB8FBDF" box="[359,529,1021,1046]" italics="true" pageId="3" pageNumber="48">Measurements</emphasis>
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. TL =
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<quantity id="4CFC3D8AAE5BFFC5FD85FC26FD55FBDE" box="[618,764,1021,1047]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="3" pageNumber="48" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
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, GW =
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<quantity id="4CFC3D8AAE5BFFC5FCB0FC26FC5BFBDE" box="[863,1010,1021,1047]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="3" pageNumber="48" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 mm</quantity>
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, PW =
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<quantity id="4CFC3D8AAE5BFFC5FBA0FC26FB4BFBDE" box="[1103,1250,1021,1047]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="3" pageNumber="48" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
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, HW =
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<quantity id="4CFC3D8AAE5BFFC5FAABFC26FF61FBF6" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="3" pageNumber="48" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
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, EW =
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<quantity id="4CFC3D8AAE5BFFC5FECBFBFEFE1DFBF6" box="[292,436,1061,1087]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="48" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
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, TL/GW = 1.8–1.7, HW/EW = 1.6–1.7. Body (
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<figureCitation id="133F8CEAAE5BFFC5FC05FBFEFB86FBF6" box="[1002,1071,1061,1087]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="3" pageNumber="48">Fig. 1</figureCitation>
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) oval, elongate; lateral outline nearly continuous between pronotum and elytron; lateral margins of pronotum gently curved; lateral margins of elytron evenly and gently curved.
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</paragraph>
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<paragraph id="8BBB906FAE5BFFC5FF29FB46FC82FAB6" blockId="3.[151,1437,821,2007]" pageId="3" pageNumber="48">
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<emphasis id="B9704C7DAE5BFFC5FF29FB46FEFCFB7E" bold="true" box="[198,341,1181,1207]" pageId="3" pageNumber="48">Coloration.</emphasis>
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Head yellow brown, pronotum yellow brown, brown medially along posterior margin (
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<figureCitation id="133F8CEAAE5BFFC5FA83FB46FF0CFB16" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="3" pageNumber="48">Fig. 1</figureCitation>
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); elytron brown with the following yellow maculae: 1) a basal macula in a band extending from lateral to sutural margins, posterior margin of macula irregular, 2) a lateral longitudinal macula at humeral angle, 3) a narrow longitudinal macula along lateral margin, and 4) a subtriangular subapical macula (
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<figureCitation id="133F8CEAAE5BFFC5FB58FACEFB52FAE6" box="[1207,1275,1301,1327]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="3" pageNumber="48">Fig. 1</figureCitation>
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). Ventral surfaces of thorax and abdomen brown except prosternum, prosternal process, propleuron and pronotal epipleuron yellow-brown; appendages yellow to yellow-brown.
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</paragraph>
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<paragraph id="8BBB906FAE5BFFC5FF29FA56FF71F976" blockId="3.[151,1437,821,2007]" pageId="3" pageNumber="48">
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<emphasis id="B9704C7DAE5BFFC5FF29FA56FE73FA6F" box="[198,474,1421,1446]" italics="true" pageId="3" pageNumber="48">Sculpture and structure</emphasis>
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. Head with very fine, inconspicuous, irregular punctation, surface between punctures shiny with indistinct microsculpture in the form of small cells; eyes medium in size (
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<figureCitation id="133F8CEAAE5BFFC5FB28FA6EFAA7FA06" box="[1223,1294,1461,1487]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="3" pageNumber="48">Fig. 1</figureCitation>
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, HW/EW = 1.6–1.7). Pronotal surface similar to that of head; with posterior angles obtuse; lateral bead narrow, of even width throughout; pronotal striae finely incised, extending nearly 1/2 distance across pronotum (
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<figureCitation id="133F8CEAAE5BFFC5FAAFF9DEFA23F9D6" box="[1344,1418,1541,1567]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="3" pageNumber="48">Fig. 1</figureCitation>
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). Elytron with anterolateral angle obtuse, not extended anteriorly (
|
||
<figureCitation id="133F8CEAAE5BFFC5FC62F9F6FC7BF98E" box="[909,978,1581,1607]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="3" pageNumber="48">Fig. 1</figureCitation>
|
||
); surface similar to pronotum. Prosternal process broad, lateral margins subparallel, apex of process broadly rounded (
|
||
<figureCitation id="133F8CEAAE5BFFC5FBB0F98EFB01F9A6" box="[1119,1192,1621,1647]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="3" pageNumber="48">Fig. 5</figureCitation>
|
||
); metacoxal process with lateral lobe minute. Pro- and mesotarsi relatively narrow in both male and female, but slightly broader in male.
|
||
</paragraph>
|
||
<paragraph id="8BBB906FAE5BFFC5FF29F916FF54F84E" blockId="3.[151,1437,821,2007]" pageId="3" pageNumber="48">
|
||
<emphasis id="B9704C7DAE5BFFC5FF29F916FEDEF92F" box="[198,375,1741,1766]" italics="true" pageId="3" pageNumber="48">Male genitalia</emphasis>
|
||
. Median lobe in ventral aspect extremely slender and long, apex slender and sharply pointed (
|
||
<figureCitation id="133F8CEAAE5BFFC5FEEFF92EFEEEF8C6" box="[256,327,1781,1807]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="3" pageNumber="48">Fig. 8</figureCitation>
|
||
); in lateral aspect narrow, extremely slender and long in apical half, abruptly curved medially to sharply pointed, recurved apex (
|
||
<figureCitation id="133F8CEAAE5BFFC5FDC3F8C6FDDDF8FE" box="[556,628,1821,1847]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="3" pageNumber="48">Fig. 9</figureCitation>
|
||
); with elongate, slender dorsal sclerite (
|
||
<figureCitation id="133F8CEAAE5BFFC5FBABF8C6FB25F8FE" box="[1092,1164,1821,1847]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="3" pageNumber="48">Fig. 9</figureCitation>
|
||
). Lateral lobe in lateral aspect broad basally, bent medially with apical segment elongate and parallel sided to narrowly rounded apex (
|
||
<figureCitation id="133F8CEAAE5BFFC5FF70F8B6FF59F84E" box="[159,240,1901,1927]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="3" pageNumber="48">Fig 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBB906FAE5BFFC2FF29F84EFF6CFEE3" blockId="3.[151,1437,821,2007]" lastBlockId="4.[151,1436,152,298]" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
|
||
<emphasis id="B9704C7DAE5BFFC5FF29F84EFE24F867" box="[198,397,1941,1966]" italics="true" pageId="3" pageNumber="48">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="133F8CEAAE5BFFC5FE71F84EFE5CF866" box="[414,501,1941,1967]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="3" pageNumber="48">Fig. 17</figureCitation>
|
||
). Bursa copulatrix elongate and narrow; spermathecal duct long, slender, somewhat coiled, expanded prior to receptacle; receptacle small, about half size of spermatheca, intermediate duct between receptacle and spermatheca broad; spermatheca globular, spermathecal spine prominent, but not elongate; fertilization duct elongate, tightly curved in spiral, attaching to large fertilization sac; vagina elongate, moderately narrow; gonocoxae elongate, slender with abrupt, small, triangular expansion at apex (
|
||
<figureCitation id="133F8CEAAE5CFFC2FA82FF33FF1EFEE3" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="4" pageNumber="49">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7BC0E7AE5CFFC2FF78FBC6FAABFB76" httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="4" pageNumber="49" targetBox="[469,1097,360,1021]" targetPageId="4">
|
||
<paragraph id="8BBB906FAE5CFFC2FF78FBC6FAABFB76" blockId="4.[151,1436,1053,1215]" pageId="4" pageNumber="49">
|
||
<emphasis id="B9704C7DAE5CFFC2FF78FBC6FEFBFBFC" bold="true" box="[151,338,1053,1077]" pageId="4" pageNumber="49">FIGURES 8–17.</emphasis>
|
||
<taxonomicName id="4C04EBECAE5CFFC2FEB5FBC5FE70FBFC" box="[346,473,1054,1077]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5CFFC2FEB5FBC5FE70FBFC" box="[346,473,1054,1077]" italics="true" pageId="4" pageNumber="49">Fontidessus</emphasis>
|
||
</taxonomicName>
|
||
species. 8–10)
|
||
<taxonomicName id="4C04EBECAE5CFFC2FD6BFBC5FC85FBFC" box="[644,812,1054,1077]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="toboganensis">
|
||
<emphasis id="B9704C7DAE5CFFC2FD6BFBC5FC85FBFC" box="[644,812,1054,1077]" italics="true" pageId="4" pageNumber="49">F. toboganensis</emphasis>
|
||
</taxonomicName>
|
||
, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11–13)
|
||
<taxonomicName id="4C04EBECAE5CFFC2FC46FB9BFBBDFB9E" box="[937,1044,1088,1111]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="B9704C7DAE5CFFC2FC46FB9BFBBDFB9E" box="[937,1044,1088,1111]" italics="true" pageId="4" pageNumber="49">F. ornatus</emphasis>
|
||
</taxonomicName>
|
||
, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14–16)
|
||
<taxonomicName id="4C04EBECAE5CFFC2FA67FBB8FF5AFB54" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="wheeleri">
|
||
<emphasis id="B9704C7DAE5CFFC2FA67FBB8FF5AFB54" italics="true" pageId="4" pageNumber="49">F. wheeleri</emphasis>
|
||
</taxonomicName>
|
||
, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17)
|
||
<taxonomicName id="4C04EBECAE5CFFC2FDF7FB73FD16FB76" box="[536,703,1192,1215]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="toboganensis">
|
||
<emphasis id="B9704C7DAE5CFFC2FDF7FB73FD8FFB76" box="[536,550,1192,1215]" italics="true" pageId="4" pageNumber="49">F</emphasis>
|
||
.
|
||
<emphasis id="B9704C7DAE5CFFC2FDDCFB73FD16FB76" box="[563,703,1192,1215]" italics="true" pageId="4" pageNumber="49">toboganensis</emphasis>
|
||
</taxonomicName>
|
||
, female genitalia, ventral aspect. Scale bars = 0.1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31EC3E4AE5CFFC2FF2AFB29FB7FFAC4" box="[197,1238,1266,1293]" pageId="4" pageNumber="49" type="etymology">
|
||
<paragraph id="8BBB906FAE5CFFC2FF2AFB29FB7FFAC4" blockId="4.[151,1436,1266,1893]" box="[197,1238,1266,1293]" pageId="4" pageNumber="49">
|
||
<emphasis id="B9704C7DAE5CFFC2FF2AFB29FEFBFAC5" bold="true" box="[197,338,1266,1292]" pageId="4" pageNumber="49">Etymology.</emphasis>
|
||
The species name is derived from the name of the
|
||
<typeStatus id="54BF2ECDAE5CFFC2FC47FB28FC72FAC4" box="[936,987,1267,1293]" pageId="4" pageNumber="49">type</typeStatus>
|
||
locality, El Tobogán.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31EC3E4AE5CFFC2FF2AFAC1FD35F9BC" pageId="4" pageNumber="49" type="distribution">
|
||
<paragraph id="8BBB906FAE5CFFC2FF2AFAC1FEA9FA34" blockId="4.[151,1436,1266,1893]" pageId="4" pageNumber="49">
|
||
<emphasis id="B9704C7DAE5CFFC2FF2AFAC1FDA8FAFD" bold="true" box="[197,513,1306,1332]" pageId="4" pageNumber="49">Distribution and habitat.</emphasis>
|
||
According to notes discovered in P.J. Spangler’s office: “The series of [
|
||
<taxonomicName id="4C04EBECAE5CFFC2FA8CFAC0FE3EFA95" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="toboganensis">
|
||
<emphasis id="B9704C7DAE5CFFC2FA8CFAC0FE3EFA95" italics="true" pageId="4" pageNumber="49">Fontidessus toboganensis</emphasis>
|
||
</taxonomicName>
|
||
] was collected from seepage areas alongside a cascade. The specimens were obtained by pulling matted plant roots from granitic bedrock and using an aspirator to suck up the water that accumulated in rock depressions as runoff from the roots. The dirty concentrated residue was then filtered through fine-meshed, nylon cloth and the very small adults and larvae were again aspirated as they crawled out of the residue.”
|
||
</paragraph>
|
||
<paragraph id="8BBB906FAE5CFFC2FF2AF9D0FD35F9BC" blockId="4.[151,1436,1266,1893]" pageId="4" pageNumber="49">Label data indicate the collection of specimens of the species from the following habitats: “seep,” “sunlit stream”, “bare rock streambed in sunlight; beneath wet leaves at margin,” “sunlit stream; slow water over sand and gravel,” and “leaves from stream.”</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31EC3E4AE5CFFC2FF2AF959FD68F8AC" pageId="4" pageNumber="49" type="materials_examined">
|
||
<paragraph id="8BBB906FAE5CFFC2FF2AF959FD68F8AC" blockId="4.[151,1436,1266,1893]" pageId="4" pageNumber="49">
|
||
<emphasis id="B9704C7DAE5CFFC2FF2AF959FE69F955" bold="true" box="[197,448,1666,1692]" pageId="4" pageNumber="49">Material examined.</emphasis>
|
||
<typeStatus id="54BF2ECDAE5CFFC2FE20F958FDC3F954" box="[463,618,1667,1693]" pageId="4" pageNumber="49" type="holotype">HOLOTYPE</typeStatus>
|
||
in
|
||
<collectionCode id="ED1508AAAE5CFFC2FD72F958FD5FF954" LSID="urn:lsid:biocol.org:col:34871" box="[669,758,1667,1693]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="49">USNM</collectionCode>
|
||
: male labeled, “
|
||
<collectingCountry id="F313D0FFAE5CFFC2FC2CF958FBD0F954" box="[963,1145,1667,1693]" name="Venezuela" pageId="4" pageNumber="49">VENEZUELA</collectingCountry>
|
||
: T.F. Amazonas Puerto Ayacucho (
|
||
<quantity id="4CFC3D8AAE5CFFC2FEF5F970FECCF90C" box="[282,357,1707,1733]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="49" unit="km" value="40.0">40km</quantity>
|
||
.S) El Tobogán, Cano Coromoto
|
||
<date id="FFBAB6AFAE5CFFC2FD30F970FC36F90C" box="[735,927,1707,1733]" pageId="4" pageNumber="49" value="1989-01-26">26 January 1989</date>
|
||
/ seep at upper shelter collected by PJSPangler RAFaitoute&CBBarr/
|
||
<typeStatus id="54BF2ECDAE5CFFC2FE39F908FDDFF924" box="[470,630,1747,1773]" pageId="4" pageNumber="49" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
<taxonomicName id="4C04EBECAE5CFFC2FD6FF908FB40F924" authority="Miller and Spangler, 2008" authorityName="Miller and Spangler" authorityYear="2008" box="[640,1257,1747,1773]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="toboganensis">
|
||
<emphasis id="B9704C7DAE5CFFC2FD6FF908FC06F925" box="[640,943,1747,1772]" italics="true" pageId="4" pageNumber="49">Fontidessus toboganensis</emphasis>
|
||
Miller and Spangler, 2008
|
||
</taxonomicName>
|
||
[red label with double black line border].”
|
||
<typeStatus id="54BF2ECDAE5CFFC2FE38F920FD2BF8DC" box="[471,642,1787,1813]" pageId="4" pageNumber="49" type="paratype">PARATYPES</typeStatus>
|
||
, 250 total, all with same locality data as
|
||
<typeStatus id="54BF2ECDAE5CFFC2FBB6F920FB6AF8DC" box="[1113,1219,1787,1813]" pageId="4" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
, 11 with same collecting date as
|
||
<typeStatus id="54BF2ECDAE5CFFC2FEABF8F8FE07F8F4" box="[324,430,1827,1853]" pageId="4" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
; 1 with
|
||
<date id="FFBAB6AFAE5CFFC2FDE6F8F8FD75F8F4" box="[521,732,1827,1853]" pageId="4" pageNumber="49" value="1986-02-22">22 February 1986</date>
|
||
; 21 with
|
||
<date id="FFBAB6AFAE5CFFC2FCA9F8F8FBB0F8F4" box="[838,1049,1827,1853]" pageId="4" pageNumber="49" value="1986-02-25">25 February 1986</date>
|
||
; 24 with
|
||
<date id="FFBAB6AFAE5CFFC2FB6CF8F8FAEEF8F4" box="[1155,1351,1827,1853]" pageId="4" pageNumber="49" value="1985-01-21">21 January 1985</date>
|
||
; 1 with
|
||
<date id="FFBAB6AFAE5CFFC2FF78F890FED4F8AC" box="[151,381,1867,1893]" pageId="4" pageNumber="49" value="1987-11-16">16 November 1987</date>
|
||
; 192 with
|
||
<date id="FFBAB6AFAE5CFFC2FE18F890FD14F8AC" box="[503,701,1867,1893]" pageId="4" pageNumber="49" value="1989-01-23">23 January 1989</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |