305 lines
39 KiB
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305 lines
39 KiB
XML
<document id="2DB7869C13D2FEBFA5E962CA3DCCC8D6" ID-DOI="10.5281/zenodo.183126" ID-GBIF-Dataset="5ba6cec4-345d-4f95-8064-dd9dc33dac4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="183126" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459892214864" checkinUser="plazi" docAuthor="Miller, Kelly B. & Spangler, Paul J." docDate="2008" docId="03AD2179AE59FFC4FF78FB36FD3BF875" docLanguage="en" docName="zt01827p052.pdf" docOrigin="Zootaxa 1827" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Fontidessus Miller and Spangler" docType="treatment" docVersion="9" lastPageNumber="47" masterDocId="FF945901AE58FFC6FFEFFFDBFFA9FFC9" masterDocTitle="Fontidessus Miller and Spangler, a new genus of Bidessini from Venezuela (Coleoptera: Dytiscidae: Hydroporinae) with three new species" masterLastPageNumber="52" masterPageNumber="45" pageNumber="46" updateTime="1698228095007" updateUser="plazi">
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<mods:titleInfo id="9B33705D6347B8D0645B1EA6543FBFA9">
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<mods:title id="1165876066BB4A24189E4E028A88E8A0">Fontidessus Miller and Spangler, a new genus of Bidessini from Venezuela (Coleoptera: Dytiscidae: Hydroporinae) with three new species</mods:title>
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<mods:name id="9E0D446F7E1ABA31B7F9FD63A74071D7" type="personal">
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<mods:namePart id="6A0FDD278430986868F6D1239EC8C474">Miller, Kelly B.</mods:namePart>
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<mods:name id="432144B4FE0B14B3DCE3BC6DD1DDE293" type="personal">
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<mods:roleTerm id="6101E6B18F99BCF18A07FC61E2C601D9">Author</mods:roleTerm>
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<mods:namePart id="BD72108BE3B9A1263BD0E73B92EACD9F">Spangler, Paul J.</mods:namePart>
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<mods:typeOfResource id="DBCE8042408D07A232C7962CA20E0F12">text</mods:typeOfResource>
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<mods:title id="0CE4610FE359275A06366C17B35483F3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="845FF4321BDE6F36B491875E0754C022">
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<mods:date id="42DF64050DAF7B17A61318A9BF8208BA">2008</mods:date>
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<mods:detail id="AC1F93F8605771B472E19A6EC2D644AE" type="volume">
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<mods:number id="0106B9ECE87117067611FB49DE8243D5">1827</mods:number>
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<mods:classification id="669CB6E71929B35DA2A6301A2B13F47A">journal article</mods:classification>
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<mods:identifier id="551AA85D9F9320B320E4879395552925" type="DOI">10.5281/zenodo.183126</mods:identifier>
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<mods:identifier id="C574E2E47D7AD3D22BADE89B1B5FC670" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="206B7A2212DF7DDC020BC67D5ACDFF2F" type="Zenodo-Dep">183126</mods:identifier>
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<treatment id="03AD2179AE59FFC4FF78FB36FD3BF875" ID-DOI="http://doi.org/10.5281/zenodo.6233620" ID-GBIF-Taxon="119372125" ID-Zenodo-Dep="6233620" LSID="urn:lsid:plazi:treatment:03AD2179AE59FFC4FF78FB36FD3BF875" httpUri="http://treatment.plazi.org/id/03AD2179AE59FFC4FF78FB36FD3BF875" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46">
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<subSubSection id="C31EC3E4AE59FFC7FF78FB36FEB7FAE3" pageId="1" pageNumber="46" type="nomenclature">
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<paragraph id="8BBB906FAE59FFC7FF78FB36FD10FACE" blockId="1.[151,697,1261,1322]" box="[151,697,1261,1287]" pageId="1" pageNumber="46">
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<heading id="D0F32703AE59FFC7FF78FB36FD10FACE" bold="true" box="[151,697,1261,1287]" fontSize="11" level="1" pageId="1" pageNumber="46" reason="1">
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<emphasis id="B9704C7DAE59FFC7FF78FB36FD10FACE" bold="true" box="[151,697,1261,1287]" pageId="1" pageNumber="46">
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<taxonomicName id="4C04EBECAE59FFC7FF78FB36FD82FACE" ID-CoL="4KQ3" authority="Miller and Spangler" authorityName="Miller and Spangler" box="[151,555,1261,1287]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
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<emphasis id="B9704C7DAE59FFC7FF78FB36FE81FACE" bold="true" box="[151,296,1261,1287]" italics="true" pageId="1" pageNumber="46">Fontidessus</emphasis>
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Miller and Spangler
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</taxonomicName>
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, new genus
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBB906FAE59FFC7FF78FACBFEB7FAE3" blockId="1.[151,697,1261,1322]" box="[151,286,1296,1322]" pageId="1" pageNumber="46">
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(
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<figureCitation id="133F8CEAAE59FFC7FF70FACBFEBFFAE3" box="[159,278,1296,1322]" captionStart-0="FIGURES 1 – 7" captionStart-1="FIGURES 8 – 17" captionStartId-0="2.[151,269,1010,1034]" captionStartId-1="4.[151,269,1053,1077]" captionTargetBox-0="[476,1111,479,980]" captionTargetBox-1="[469,1097,360,1021]" captionTargetId-0="figure@2.[473,1113,477,983]" captionTargetId-1="figure@4.[463,1103,354,1031]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." captionText-1="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri-0="https://zenodo.org/record/183127/files/figure.png" httpUri-1="https://zenodo.org/record/183128/files/figure.png" pageId="1" pageNumber="46">Figs 1–17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31EC3E4AE59FFC7FF78FA81FBC7FABD" box="[151,1134,1370,1396]" pageId="1" pageNumber="46" type="materials_examined">
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<paragraph id="8BBB906FAE59FFC7FF78FA81FBC7FABD" blockId="1.[151,1437,1370,2036]" box="[151,1134,1370,1396]" pageId="1" pageNumber="46">
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<emphasis id="B9704C7DAE59FFC7FF78FA81FE90FABD" bold="true" box="[151,313,1370,1396]" pageId="1" pageNumber="46">
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<typeStatus id="54BF2ECDAE59FFC7FF78FA81FF7DFABD" box="[151,212,1370,1396]" pageId="1" pageNumber="46">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C04EBECAE59FFC7FEAFFA80FD69FABD" authority="Miller" authorityName="Miller" box="[320,704,1370,1396]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="toboganensis" status="sp. nov.">
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<emphasis id="B9704C7DAE59FFC7FEAFFA80FDC6FABD" box="[320,623,1371,1396]" italics="true" pageId="1" pageNumber="46">Fontidessus toboganensis</emphasis>
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Miller
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</taxonomicName>
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,
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<taxonomicNameLabel id="A243F106AE59FFC7FD23FA81FCF1FABD" box="[716,856,1370,1396]" pageId="1" pageNumber="46" rank="species">new species</taxonomicNameLabel>
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by current designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31EC3E4AE59FFC4FF2AFA59FACCFE6B" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46" type="diagnosis">
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<paragraph id="8BBB906FAE59FFC7FF2AFA59FAAFF915" blockId="1.[151,1437,1370,2036]" pageId="1" pageNumber="46">
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<emphasis id="B9704C7DAE59FFC7FF2AFA59FDBAFA55" bold="true" box="[197,531,1410,1436]" pageId="1" pageNumber="46">Diagnosis and description.</emphasis>
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<taxonomicName id="4C04EBECAE59FFC7FDF3FA58FD01FA55" box="[540,680,1411,1436]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
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<emphasis id="B9704C7DAE59FFC7FDF3FA58FD01FA55" box="[540,680,1411,1436]" italics="true" pageId="1" pageNumber="46">Fontidessus</emphasis>
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</taxonomicName>
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is clearly a member of the tribe
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<taxonomicName id="4C04EBECAE59FFC7FBDAFA59FB0BFA55" box="[1077,1186,1410,1436]" class="Insecta" family="Dytiscidae" genus="Bidessini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="47" phylum="Arthropoda" rank="genus">Bidessini</taxonomicName>
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based on the bi-segmented lateral lobes of the aedeagus (
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<figureCitation id="133F8CEAAE59FFC7FDA2FA71FD55FA0D" box="[589,764,1450,1476]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="1" pageNumber="46">Figs 10, 13, 16</figureCitation>
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) (
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<bibRefCitation id="EF95ED9EAE59FFC7FCFBFA71FC16FA0D" author="Bistrom" box="[788,959,1450,1476]" pageId="1" pageNumber="52" refString="Bistrom, O. (1988) Generic review of the Bidessini (Coleoptera, Dytiscidae). Acta Zoologica Fennica, 184, 1 - 41." type="journal article" year="1988">Biström, 1988</bibRefCitation>
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) and the presence of a distinct spermathecal spine (
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<figureCitation id="133F8CEAAE59FFC7FEF5FA09FEC7FA25" box="[282,366,1490,1516]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="1" pageNumber="46">Fig. 17</figureCitation>
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) (
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<bibRefCitation id="EF95ED9EAE59FFC7FE68FA09FDB4FA25" author="Miller" box="[391,541,1490,1516]" pageId="1" pageNumber="52" refString="Miller, K. B. (2001) On the phylogeny of the Dytiscidae (Coleoptera) with emphasis on the morphology of the female reproductive tract. Insect Systematics and Evolution, 32, 45 - 92." type="journal article" year="2001">Miller, 2001</bibRefCitation>
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).
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<taxonomicName id="4C04EBECAE59FFC7FDDEFA08FD14FA25" box="[561,701,1491,1516]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
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<emphasis id="B9704C7DAE59FFC7FDDEFA08FD14FA25" box="[561,701,1491,1516]" italics="true" pageId="1" pageNumber="46">Fontidessus</emphasis>
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</taxonomicName>
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differs from other genera in the tribe by the combination of: 1) transverse occipital line absent (
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<figureCitation id="133F8CEAAE59FFC7FDF5FA21FD2FF9DD" box="[538,646,1530,1556]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="1" pageNumber="46">Figs 1–3</figureCitation>
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), 2) basal pronotal striae present (
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<figureCitation id="133F8CEAAE59FFC7FBCAFA21FB38F9DD" box="[1061,1169,1530,1556]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="1" pageNumber="46">Figs 1–3</figureCitation>
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), 3) basal elytral stria absent (
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<figureCitation id="133F8CEAAE59FFC7FF1CF9F9FEF7F9F5" box="[243,350,1570,1596]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="1" pageNumber="46">Figs 1–3</figureCitation>
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), 4) elytral sutural stria faintly present in some specimens (e.g.
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<figureCitation id="133F8CEAAE59FFC7FB8FF9F9FB0CF9F5" box="[1120,1189,1570,1596]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
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), 5) anterior clypeal margin unmodified, 6) elytron without longitudinal carinae, 7) epipleuron without transverse carina at humeral angle, 8) lateral lobes of aedeagus two-segmented (
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<figureCitation id="133F8CEAAE59FFC7FC8CF9A9FBB5F945" box="[867,1052,1650,1676]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="1" pageNumber="46">Figs 10, 13, 16</figureCitation>
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), 9) habitus elongate, oval, with lateral pronotal and elytral margins nearly continuously and shallowly curved (
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<figureCitation id="133F8CEAAE59FFC7FBA1F941FB10F97D" box="[1102,1209,1690,1716]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="1" pageNumber="46">Figs 1–3</figureCitation>
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), and 10) metatrochanter extremely large relative to metafemur, approximately 0.6 × length of metafemur (
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<figureCitation id="133F8CEAAE59FFC7FB40F919FB51F915" box="[1199,1272,1730,1756]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="1" pageNumber="46">Fig. 4</figureCitation>
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).
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</paragraph>
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<paragraph id="8BBB906FAE59FFC7FF29F931FE5FF83D" blockId="1.[151,1437,1370,2036]" pageId="1" pageNumber="46">
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This genus is most similar to the genera
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<taxonomicName id="4C04EBECAE59FFC7FD44F930FCC0F8CD" authority="Guignot" authorityName="Guignot" box="[683,873,1770,1796]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
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<emphasis id="B9704C7DAE59FFC7FD44F930FCA8F8CD" box="[683,769,1771,1796]" italics="true" pageId="1" pageNumber="46">Uvarus</emphasis>
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Guignot
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</taxonomicName>
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and
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<taxonomicName id="4C04EBECAE59FFC7FC4AF930FB05F8CD" authority="Regimbart" authorityName="Regimbart" box="[933,1196,1770,1796]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE59FFC7FC4AF930FB8EF8CD" box="[933,1063,1771,1796]" italics="true" pageId="1" pageNumber="46">Bidessodes</emphasis>
|
||
Régimbart
|
||
</taxonomicName>
|
||
(to which it keys in Biström’s 1988 key to bidessine genera). From
|
||
<taxonomicName id="4C04EBECAE59FFC7FD2FF8C8FCEBF8E5" box="[704,834,1811,1836]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE59FFC7FD2FF8C8FCEBF8E5" box="[704,834,1811,1836]" italics="true" pageId="1" pageNumber="46">Bidessodes</emphasis>
|
||
</taxonomicName>
|
||
it differs by 1) dorsum not iridescent, and 2) metatibiae without natatory setae and metatarsi with few natatory setae.
|
||
<taxonomicName id="4C04EBECAE59FFC7FC5BF8E0FB9FF89D" box="[948,1078,1851,1876]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE59FFC7FC5BF8E0FB9FF89D" box="[948,1078,1851,1876]" italics="true" pageId="1" pageNumber="46">Bidessodes</emphasis>
|
||
</taxonomicName>
|
||
appears to be a monophyletic group. Although some of its members have dramatic modifications to the male legs or genitalia, they are superficially quite similar and have similar iridescence and color pattern on the dorsal surface.
|
||
<taxonomicName id="4C04EBECAE59FFC7FAFFF850FA35F86D" box="[1296,1436,1931,1956]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE59FFC7FAFFF850FA35F86D" box="[1296,1436,1931,1956]" italics="true" pageId="1" pageNumber="46">Fontidessus</emphasis>
|
||
</taxonomicName>
|
||
specimens lack these features. Despite the absence of a cladistic phylogenetic analysis,
|
||
<taxonomicName id="4C04EBECAE59FFC7FB7DF868FAB7F805" box="[1170,1310,1971,1996]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE59FFC7FB7DF868FAB7F805" box="[1170,1310,1971,1996]" italics="true" pageId="1" pageNumber="46">Fontidessus</emphasis>
|
||
</taxonomicName>
|
||
appears to be clearly outside
|
||
<taxonomicName id="4C04EBECAE59FFC7FE82F800FE46F83D" box="[365,495,2011,2036]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE59FFC7FE82F800FE46F83D" box="[365,495,2011,2036]" italics="true" pageId="1" pageNumber="46">Bidessodes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBB906FAE5AFFC4FF2AFF43FACCFE6B" blockId="2.[151,1436,152,418]" pageId="2" pageNumber="47">
|
||
From
|
||
<taxonomicName id="4C04EBECAE5AFFC4FEE1FF43FECDFF78" box="[270,356,152,177]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5AFFC4FEE1FF43FECDFF78" box="[270,356,152,177]" italics="true" pageId="2" pageNumber="47">Uvarus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C04EBECAE5AFFC4FE9AFF43FDA8FF78" box="[373,513,152,177]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5AFFC4FE9AFF43FDA8FF78" box="[373,513,152,177]" italics="true" pageId="2" pageNumber="47">Fontidessus</emphasis>
|
||
</taxonomicName>
|
||
genus differs in lacking basal elytra striae and natatory setae. It is, however, quite similar to
|
||
<taxonomicName id="4C04EBECAE5AFFC4FEBDFF1BFE01FF10" box="[338,424,192,217]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5AFFC4FEBDFF1BFE01FF10" box="[338,424,192,217]" italics="true" pageId="2" pageNumber="47">Uvarus</emphasis>
|
||
</taxonomicName>
|
||
genus in most characters and general features such as shape, size and coloration. The problem with these observations is that
|
||
<taxonomicName id="4C04EBECAE5AFFC4FD68FF33FD74FEC8" box="[647,733,232,257]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5AFFC4FD68FF33FD74FEC8" box="[647,733,232,257]" italics="true" pageId="2" pageNumber="47">Uvarus</emphasis>
|
||
</taxonomicName>
|
||
is a heterogeneous group with a worldwide distribution. Within
|
||
<taxonomicName id="4C04EBECAE5AFFC4FF00FECBFEF5FEE3" box="[239,348,272,298]" class="Insecta" family="Dytiscidae" genus="Bidessini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Bidessini</taxonomicName>
|
||
it is the “trashcan” group that includes those taxa without an occipital line but also without any unifying synapomorphy. Placing these new species in
|
||
<taxonomicName id="4C04EBECAE5AFFC4FCA0FEE3FC0FFE98" box="[847,934,312,337]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5AFFC4FCA0FEE3FC0FFE98" box="[847,934,312,337]" italics="true" pageId="2" pageNumber="47">Uvarus</emphasis>
|
||
</taxonomicName>
|
||
would require an expansion of an already problematic group to include a new character combination. It seems more desirable to place these species in a new genus until the classification of the
|
||
<taxonomicName id="4C04EBECAE5AFFC4FD80FE53FD75FE6B" box="[623,732,392,418]" class="Insecta" family="Dytiscidae" genus="Bidessini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Bidessini</taxonomicName>
|
||
can be more thoroughly tested (see Discussion below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7BC0E7AE5AFFC4FF78FC29FC49FB87" httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="2" pageNumber="47" targetBox="[476,1111,479,980]" targetPageId="2">
|
||
<paragraph id="8BBB906FAE5AFFC4FF78FC29FC49FB87" blockId="2.[151,1435,1010,1102]" pageId="2" pageNumber="47">
|
||
<emphasis id="B9704C7DAE5AFFC4FF78FC29FEEBFBC3" bold="true" box="[151,322,1010,1034]" pageId="2" pageNumber="47">FIGURES 1–7.</emphasis>
|
||
<taxonomicName id="4C04EBECAE5AFFC4FEA7FC29FE6EFBC0" box="[328,455,1010,1033]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5AFFC4FEA7FC29FE6EFBC0" box="[328,455,1010,1033]" italics="true" pageId="2" pageNumber="47">Fontidessus</emphasis>
|
||
</taxonomicName>
|
||
species. 1–3) Dorsal habitus, 1)
|
||
<taxonomicName id="4C04EBECAE5AFFC4FCCEFC29FC6EFBC0" box="[801,967,1010,1033]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="toboganensis">
|
||
<emphasis id="B9704C7DAE5AFFC4FCCEFC29FC6EFBC0" box="[801,967,1010,1033]" italics="true" pageId="2" pageNumber="47">F. toboganensis</emphasis>
|
||
</taxonomicName>
|
||
, 2)
|
||
<taxonomicName id="4C04EBECAE5AFFC4FC1FFC29FBF2FBC0" box="[1008,1115,1010,1033]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="B9704C7DAE5AFFC4FC1FFC29FBF2FBC0" box="[1008,1115,1010,1033]" italics="true" pageId="2" pageNumber="47">F. ornatus</emphasis>
|
||
</taxonomicName>
|
||
, 3)
|
||
<taxonomicName id="4C04EBECAE5AFFC4FB6BFC29FB50FBC0" box="[1156,1273,1010,1033]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="wheeleri">
|
||
<emphasis id="B9704C7DAE5AFFC4FB6BFC29FB50FBC0" box="[1156,1273,1010,1033]" italics="true" pageId="2" pageNumber="47">F. wheeleri</emphasis>
|
||
</taxonomicName>
|
||
. 4)
|
||
<taxonomicName id="4C04EBECAE5AFFC4FACDFC29FF65FBE5" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="toboganensis">
|
||
<emphasis id="B9704C7DAE5AFFC4FACDFC29FF65FBE5" italics="true" pageId="2" pageNumber="47">F. toboganensis</emphasis>
|
||
</taxonomicName>
|
||
, left metatrochanter and metafemur, anterior aspect. 5–7) prosternal process, 5)
|
||
<taxonomicName id="4C04EBECAE5AFFC4FBCDFBCEFB60FBE5" box="[1058,1225,1045,1068]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="toboganensis">
|
||
<emphasis id="B9704C7DAE5AFFC4FBCDFBCEFB60FBE5" box="[1058,1225,1045,1068]" italics="true" pageId="2" pageNumber="47">F. toboganensis</emphasis>
|
||
</taxonomicName>
|
||
, 6)
|
||
<taxonomicName id="4C04EBECAE5AFFC4FB1CFBCEFAF6FBE5" box="[1267,1375,1045,1068]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="B9704C7DAE5AFFC4FB1CFBCEFAF6FBE5" box="[1267,1375,1045,1068]" italics="true" pageId="2" pageNumber="47">F. ornatus</emphasis>
|
||
</taxonomicName>
|
||
, 7)
|
||
<taxonomicName id="4C04EBECAE5AFFC4FA67FBCEFF5AFB87" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="wheeleri">
|
||
<emphasis id="B9704C7DAE5AFFC4FA67FBCEFF5AFB87" italics="true" pageId="2" pageNumber="47">F. wheeleri</emphasis>
|
||
</taxonomicName>
|
||
. Scale bar a = 1.0mm for Figs 1–4. Scale bar b = 0.1mm for Figs 5–7.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31EC3E4AE5AFFC4FF2AFB5AFEC5FB25" pageId="2" pageNumber="47" type="etymology">
|
||
<paragraph id="8BBB906FAE5AFFC4FF2AFB5AFEC5FB25" blockId="2.[151,1437,1153,1980]" pageId="2" pageNumber="47">
|
||
<emphasis id="B9704C7DAE5AFFC4FF2AFB5AFEFBFB52" bold="true" box="[197,338,1153,1179]" pageId="2" pageNumber="47">Etymology.</emphasis>
|
||
This genus is named
|
||
<taxonomicName id="4C04EBECAE5AFFC4FDB8FB59FD4AFB52" box="[599,739,1154,1179]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5AFFC4FDB8FB59FD4AFB52" box="[599,739,1154,1179]" italics="true" pageId="2" pageNumber="47">Fontidessus</emphasis>
|
||
</taxonomicName>
|
||
from the Latin word
|
||
<emphasis id="B9704C7DAE5AFFC4FC0BFB59FB8CFB52" box="[996,1061,1154,1179]" italics="true" pageId="2" pageNumber="47">fontis</emphasis>
|
||
, meaning “spring,” referring to the seepage-like habitat of the
|
||
<typeStatus id="54BF2ECDAE5AFFC4FE13FB71FD86FB0D" box="[508,559,1194,1220]" pageId="2" pageNumber="47">type</typeStatus>
|
||
locality of the
|
||
<typeStatus id="54BF2ECDAE5AFFC4FD31FB71FCB8FB0D" box="[734,785,1194,1220]" pageId="2" pageNumber="47">type</typeStatus>
|
||
species, and
|
||
<emphasis id="B9704C7DAE5AFFC4FC43FB71FC51FB0A" box="[940,1016,1194,1219]" italics="true" pageId="2" pageNumber="47">dessus</emphasis>
|
||
, a common root for genus names in the tribe
|
||
<taxonomicName id="4C04EBECAE5AFFC4FF10FB09FEC5FB25" box="[255,364,1234,1260]" class="Insecta" family="Dytiscidae" genus="Bidessini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Bidessini</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31EC3E4AE5AFFC4FF2AFB22FB97FAAD" pageId="2" pageNumber="47" type="distribution">
|
||
<paragraph id="8BBB906FAE5AFFC4FF2AFB22FB97FAAD" blockId="2.[151,1437,1153,1980]" pageId="2" pageNumber="47">
|
||
<emphasis id="B9704C7DAE5AFFC4FF2AFB22FDABFADA" bold="true" box="[197,514,1273,1299]" pageId="2" pageNumber="47">Distribution and habitat.</emphasis>
|
||
The genus is known from three sympatric species from only a single site in
|
||
<collectingCountry id="F313D0FFAE5AFFC4FF78FAF9FEA6FAF5" box="[151,271,1314,1340]" name="Venezuela" pageId="2" pageNumber="47">Venezuela</collectingCountry>
|
||
(see
|
||
<typeStatus id="54BF2ECDAE5AFFC4FEA3FAF9FED6FAF5" box="[332,383,1314,1340]" pageId="2" pageNumber="47">type</typeStatus>
|
||
localities below). In each case, however, the species do not appear to occupy typical open water. Instead they appear to occur in seeps or shallow water at stream margins.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31EC3E4AE5AFFC4FF2AFAAAFD3BF875" pageId="2" pageNumber="47" type="discussion">
|
||
<paragraph id="8BBB906FAE5AFFC4FF2AFAAAFD3BF875" blockId="2.[151,1437,1153,1980]" pageId="2" pageNumber="47">
|
||
<emphasis id="B9704C7DAE5AFFC4FF2AFAAAFEF8FA42" bold="true" box="[197,337,1393,1419]" pageId="2" pageNumber="47">Discussion.</emphasis>
|
||
The genera of
|
||
<taxonomicName id="4C04EBECAE5AFFC4FDEBFAA9FDD8FA45" box="[516,625,1394,1420]" class="Insecta" family="Dytiscidae" genus="Bidessini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Bidessini</taxonomicName>
|
||
are currently defined based on the combination of several characters such as the presence or absent of an occipital line, basal sutural and pronotal striae, epipleural carinae, elytra carinae and more subjective characters such as general shape. Because of the nature of the various combinations it is clear that homoplasy, or loss, is exhibited in many of the characters throughout the tribe. There has been only limited cladistic testing of these characters (e.g.
|
||
<bibRefCitation id="EF95ED9EAE5AFFC4FCA5F9C9FB8CF9E5" author="Miller" box="[842,1061,1554,1580]" pageId="2" pageNumber="52" refString="Miller, K. B., Wolfe, G. W. & Bistrom, O. (2006) Phylogeny of the Hydroporinae and classification of the genus Peschetius Guignot, 1942 (Coleoptera: Dytiscidae). Insect Systematics and Evolution, 37, 257 - 279." type="journal article" year="2006">
|
||
Miller
|
||
<emphasis id="B9704C7DAE5AFFC4FC75F9C9FC7CF9E2" box="[922,981,1554,1579]" italics="true" pageId="2" pageNumber="47">et al.</emphasis>
|
||
, 2006
|
||
</bibRefCitation>
|
||
). When new species are discovered with a new combination of characters, such as those described here, it is not easy to decide whether to erect a new genus or expand the definition of existing genera (which may require synonymy with other genera). It seems best, in the absence of cladistic testing of relationships and putative homologies, to place anomalous species in new genera to draw attention to their unique character combination until such time as the genera can be phylogenetically revised. In further support of the placement of this taxon outside both
|
||
<taxonomicName id="4C04EBECAE5AFFC4FAAAF901FA32F93A" box="[1349,1435,1754,1779]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5AFFC4FAAAF901FA32F93A" box="[1349,1435,1754,1779]" italics="true" pageId="2" pageNumber="47">Uvarus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C04EBECAE5AFFC4FF24F8D9FEE4F8D2" box="[203,333,1794,1819]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5AFFC4FF24F8D9FEE4F8D2" box="[203,333,1794,1819]" italics="true" pageId="2" pageNumber="47">Bidessodes</emphasis>
|
||
</taxonomicName>
|
||
, a recent molecular phylogenetic analysis of
|
||
<taxonomicName id="4C04EBECAE5AFFC4FC9DF8D9FC46F8D5" box="[882,1007,1794,1820]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF95ED9EAE5AFFC4FBCEF8D9FAA7F8D5" box="[1057,1294,1794,1820]" pageId="2" pageNumber="52" refString="Ribera, I., Vogler, A. P. & Balke, M. (2008) Phylogeny and diversification of diving beetles (Coleoptera: Dytiscidae). Cladistics, in press (doi: 10.1111 / j. 1096 - 0031.2007.00192. x)." type="book">
|
||
Ribera
|
||
<emphasis id="B9704C7DAE5AFFC4FB96F8D9FB1FF8D2" box="[1145,1206,1794,1819]" italics="true" pageId="2" pageNumber="47">et al.</emphasis>
|
||
(2008)
|
||
</bibRefCitation>
|
||
included at least one member of this new genus. According to M. Balke (personal communication), “
|
||
<emphasis id="B9704C7DAE5AFFC4FB51F8F1FF0FF8A2" italics="true" pageId="2" pageNumber="47">
|
||
unknown
|
||
<taxonomicName id="4C04EBECAE5AFFC4FADFF8F1FA35F88A" box="[1328,1436,1834,1859]" class="Insecta" family="Dytiscidae" genus="Bidessini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Bidessini</taxonomicName>
|
||
a
|
||
</emphasis>
|
||
” in that analysis (MB1171,
|
||
<bibRefCitation id="EF95ED9EAE5AFFC4FE11F889FD4BF8A5" author="Ribera" box="[510,738,1874,1900]" pageId="2" pageNumber="52" refString="Ribera, I., Vogler, A. P. & Balke, M. (2008) Phylogeny and diversification of diving beetles (Coleoptera: Dytiscidae). Cladistics, in press (doi: 10.1111 / j. 1096 - 0031.2007.00192. x)." type="book" year="2008">
|
||
Ribera
|
||
<emphasis id="B9704C7DAE5AFFC4FDB8F889FD3DF8A2" box="[599,660,1874,1899]" italics="true" pageId="2" pageNumber="47">et al.</emphasis>
|
||
, 2008
|
||
</bibRefCitation>
|
||
) is the species described here as
|
||
<taxonomicName id="4C04EBECAE5AFFC4FB97F889FA9AF8A2" box="[1144,1331,1874,1899]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="toboganensis">
|
||
<emphasis id="B9704C7DAE5AFFC4FB97F889FA9AF8A2" box="[1144,1331,1874,1899]" italics="true" pageId="2" pageNumber="47">F. toboganensis</emphasis>
|
||
</taxonomicName>
|
||
. In their analysis
|
||
<taxonomicName id="4C04EBECAE5AFFC4FEEEF8A1FE17F85A" box="[257,446,1914,1939]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="toboganensis">
|
||
<emphasis id="B9704C7DAE5AFFC4FEEEF8A1FE17F85A" box="[257,446,1914,1939]" italics="true" pageId="2" pageNumber="47">F. toboganensis</emphasis>
|
||
</taxonomicName>
|
||
is resolved with two other unidentified
|
||
<taxonomicName id="4C04EBECAE5AFFC4FC5CF8A1FB89F85D" box="[947,1056,1914,1940]" class="Insecta" family="Dytiscidae" genus="Bidessini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Bidessini</taxonomicName>
|
||
and well removed from either
|
||
<taxonomicName id="4C04EBECAE5AFFC4FF78F879FF44F872" box="[151,237,1954,1979]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5AFFC4FF78F879FF44F872" box="[151,237,1954,1979]" italics="true" pageId="2" pageNumber="47">Uvarus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C04EBECAE5AFFC4FEFBF879FD26F875" authority="Ribera et al., 2008" authorityName="Ribera et al." authorityYear="2008" box="[276,655,1954,1980]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9704C7DAE5AFFC4FEFBF879FE3FF872" box="[276,406,1954,1979]" italics="true" pageId="2" pageNumber="47">Bidessodes</emphasis>
|
||
(
|
||
<bibRefCitation id="EF95ED9EAE5AFFC4FE4AF879FD2FF875" author="Ribera" box="[421,646,1954,1980]" pageId="2" pageNumber="52" refString="Ribera, I., Vogler, A. P. & Balke, M. (2008) Phylogeny and diversification of diving beetles (Coleoptera: Dytiscidae). Cladistics, in press (doi: 10.1111 / j. 1096 - 0031.2007.00192. x)." type="book" year="2008">
|
||
Ribera
|
||
<emphasis id="B9704C7DAE5AFFC4FE12F879FD9EF872" box="[509,567,1954,1979]" italics="true" pageId="2" pageNumber="47">et al.</emphasis>
|
||
, 2008
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |