260 lines
23 KiB
XML
260 lines
23 KiB
XML
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<mods:title id="230AFC443C52AC037DBEA8EA36841974">Phylogenetic structure of Holbrookia lacerata (Cope 1880) (Squamata: Phrynosomatidae): one species or two?</mods:title>
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<mods:title id="43B437514797C504E8C7433256759858">Zootaxa</mods:title>
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<mods:date id="7A61C979AD8443C6DC6470F134B37DF3">2019</mods:date>
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<treatment id="03A3C76EFFFCFFA4AC926C9A143BFD47" ID-DOI="http://doi.org/10.5281/zenodo.5625493" ID-GBIF-Taxon="159418533" ID-Zenodo-Dep="5625493" LSID="urn:lsid:plazi:treatment:03A3C76EFFFCFFA4AC926C9A143BFD47" httpUri="http://treatment.plazi.org/id/03A3C76EFFFCFFA4AC926C9A143BFD47" lastPageId="10" lastPageNumber="149" pageId="9" pageNumber="148">
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<emphasis id="B97EAA6AFFFCFFA7AC926C9A1217F8E9" bold="true" box="[151,608,1771,1798]" pageId="9" pageNumber="148">
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<emphasis id="B97EAA6AFFFCFFA7AC926C9A11CFF8EA" bold="true" box="[151,440,1771,1798]" italics="true" pageId="9" pageNumber="148">Holbrookia subcaudalis</emphasis>
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(
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<bibRefCitation id="EF9B0B89FFFCFFA7ADCD6C9A122FF8E9" author="Axtell, R. A." box="[456,600,1771,1797]" pageId="9" pageNumber="148" pagination="163 - 179" refId="ref7480" refString="Axtell, R. A. (1956) A solution to the long neglected Holbrookia lacerata problem and the destription of two new subspecies of Holbrookia. Bulletin of the Chicago Academy of Science, 10 (11), 163 - 179." type="journal article" year="1956">Axtell 1956</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BB57678FFFCFFA7AC926D7E1232F8C5" blockId="9.[151,608,1771,1869]" box="[151,581,1807,1833]" pageId="9" pageNumber="148">Tamaulipan Spot-tailed Earless Lizard</paragraph>
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</subSubSection>
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<subSubSection id="C31025F3FFFCFFA7AC926D42109CF8A1" box="[151,235,1843,1869]" pageId="9" pageNumber="148" type="description">
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<paragraph id="8BB57678FFFCFFA7AC926D42109CF8A1" blockId="9.[151,608,1771,1869]" box="[151,235,1843,1869]" pageId="9" pageNumber="148">
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(
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<figureCitation id="13316AFDFFFCFFA7ACA56D421094F8A1" box="[160,227,1843,1869]" captionStart="FIGURE 9" captionStartId="10.[151,260,1937,1964]" captionTargetBox="[213,1247,683,1910]" captionTargetId="figure@10.[340,1247,702,1910]" captionText="FIGURE 9. Dorsal (top) and ventral (bottom) views of Tamaulipan Spot-tailed Earless Lizard (Holbrookia subcaudalis) holotype specimen collected by Ralph W. Axtell on 6 June 1955 and housed at Texas Natural History Collection, University of Texas (TNHC 20000)." httpUri="https://zenodo.org/record/3405215/files/figure.png" pageId="9" pageNumber="148">Fig. 9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31025F3FFFCFFA7AC926D0612BAF842" pageId="9" pageNumber="148" type="reference_group">
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<paragraph id="8BB57678FFFCFFA7AC926D061214F863" blockId="9.[151,717,1910,1966]" box="[151,611,1910,1935]" pageId="9" pageNumber="148">
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<taxonomicName id="4C0A0DFBFFFCFFA7AC926D061214F863" authority="Axtell 1956" authorityName="Axtell" authorityYear="1956" box="[151,611,1910,1935]" class="Reptilia" family="Phrynosomatidae" genus="Holbrookia" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="148" phylum="Chordata" rank="subSpecies" species="lacerata" subSpecies="subcaudalis">
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<emphasis id="B97EAA6AFFFCFFA7AC926D061192F863" box="[151,485,1911,1935]" italics="true" pageId="9" pageNumber="148">Holbrookia lacerata subcaudalis</emphasis>
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<bibRefCitation id="EF9B0B89FFFCFFA7ADEF6D071214F863" author="Axtell, R. A." box="[490,611,1910,1935]" pageId="9" pageNumber="148" pagination="163 - 179" refId="ref7480" refString="Axtell, R. A. (1956) A solution to the long neglected Holbrookia lacerata problem and the destription of two new subspecies of Holbrookia. Bulletin of the Chicago Academy of Science, 10 (11), 163 - 179." type="journal article" year="1956">Axtell 1956</bibRefCitation>
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<paragraph id="8BB57678FFFCFFA7AC926DE712BAF842" blockId="9.[151,717,1910,1966]" box="[151,717,1941,1966]" pageId="9" pageNumber="148">
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<taxonomicName id="4C0A0DFBFFFCFFA7AC926DE71222F842" authority="Hibbitts et al. 2019" authorityName="Hibbitts" authorityYear="2019" box="[151,597,1941,1966]" class="Reptilia" family="Phrynosomatidae" genus="Holbrookia" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="148" phylum="Chordata" rank="species" species="subcaudalis">
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<emphasis id="B97EAA6AFFFCFFA7AC926DE711FDF842" box="[151,394,1942,1966]" italics="true" pageId="9" pageNumber="148">Holbrookia subcaudalis</emphasis>
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Hibbitts
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<emphasis id="B97EAA6AFFFCFFA7ADEC6DE61262F842" box="[489,533,1942,1966]" italics="true" pageId="9" pageNumber="148">et al</emphasis>
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. 2019
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(this study)
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<subSubSection id="C31025F3FFFCFFA4AC926DAD11C3FF3A" lastPageId="10" lastPageNumber="149" pageId="9" pageNumber="148" type="materials_examined">
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<paragraph id="8BB57678FFFCFFA4AC926DAD11C3FF3A" blockId="9.[151,1436,2012,2039]" lastBlockId="10.[151,1437,152,683]" lastPageId="10" lastPageNumber="149" pageId="9" pageNumber="148">
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<materialsCitation id="3B627C25FFFCFFA4AC926DAD11C3FF3A" ID-GBIF-Occurrence="2397873946" collectingDate="1955-06-06" collectionCode="TNHC" collectorName="Natural History Collection & University & Ralph W. Axtell" country="United States of America" county="Nueces County" elevation="23" lastPageId="10" lastPageNumber="149" latitude="27.6" location="Bishop" longLatPrecision="1236" longitude="-97.75" pageId="9" pageNumber="148" specimenCode="TNHC 20000" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
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<materialsCitation id="3B627C25FFFCFFA4AC926DAD11C6FF3A" ID-GBIF-Occurrence="2397873945" collectingDate="1955-06-06" collectionCode="TNHC" collectorName="Natural History Collection & University & Ralph W. Axtell" country="United States of America" county="Nueces County" elevation="23" lastPageId="10" lastPageNumber="149" latitude="27.6" location="Bishop" longLatPrecision="1236" longitude="-97.75" pageId="9" pageNumber="148" specimenCode="TNHC 20000" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
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.
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,
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of
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(
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); collected by
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<collectorName id="26FF13AEFFFCFFA7A8C16DAD150EF81A" box="[1220,1401,2012,2039]" pageId="9" pageNumber="148">Ralph W. Axtell</collectorName>
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on
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<date id="FFB450B8FFFFFFA4AC926AE91169FF5F" box="[151,286,152,179]" pageId="10" pageNumber="149" value="1955-06-06">
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6 June
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</date>
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in plowed field
|
||
<quantity id="4CF2DB9DFFFFFFA4ADD06AE8124BFF5E" box="[469,572,152,179]" metricMagnitude="3" metricUnit="m" metricValue="7.7248512" pageId="10" pageNumber="149" unit="mi" value="4.8">
|
||
<locationDeviation id="9A076F5AFFFFFFA4ADD06AE8124BFF5E" box="[469,572,152,179]" metricMagnitude="3" metricUnit="m" metricValue="7.7248512" pageId="10" pageNumber="149" unit="mi" value="4.8">4.8 miles</locationDeviation>
|
||
</quantity>
|
||
east northeast of
|
||
<location id="8ED520A3FFFFFFA4AF046AE91324FF5E" LSID="urn:lsid:plazi:treatment:03A3C76EFFFCFFA4AC926C9A143BFD47:8ED520A3FFFFFFA4AF046AE91324FF5E" box="[769,851,152,178]" country="United States of America" county="Nueces County" latitude="27.6" longLatPrecision="1236" longitude="-97.75" name="Bishop" pageId="10" pageNumber="149" stateProvince="Texas">Bishop</location>
|
||
,
|
||
<collectingCounty id="62D40EF4FFFFFFA4AF5B6AE8147CFF5F" box="[862,1035,153,179]" pageId="10" pageNumber="149">Nueces County</collectingCounty>
|
||
,
|
||
<collectingRegion id="49CEB89AFFFFFFA4A8106AE8142EFF5F" box="[1045,1113,153,179]" country="United States of America" name="Texas" pageId="10" pageNumber="149">Texas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31D36E8FFFFFFA4A8616AE814EDFF5F" box="[1124,1178,153,179]" name="United States of America" pageId="10" pageNumber="149">USA</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE3E10BFFFFFFFA4A8AC6AE81560FF5F" box="[1193,1303,152,179]" degrees="27" direction="north" minutes="36" orientation="latitude" pageId="10" pageNumber="149" precision="925" value="27.6">27º 36’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE3E10BFFFFFFFA4A9266AE815E3FF5F" box="[1315,1428,152,179]" degrees="97" direction="west" minutes="45" orientation="longitude" pageId="10" pageNumber="149" precision="925" value="-97.75">97º 45’ W</geoCoordinate>
|
||
) at an elevation of
|
||
<elevation id="0027914BFFFFFFA4AD656ACD11C6FF3A" box="[352,433,188,215]" metricMagnitude="1" metricUnit="m" metricValue="2.286" pageId="10" pageNumber="149" unit="ft" value="75.0">
|
||
<quantity id="4CF2DB9DFFFFFFA4AD656ACD11C6FF3A" box="[352,433,188,215]" metricMagnitude="1" metricUnit="m" metricValue="2.286" pageId="10" pageNumber="149" unit="ft" value="75.0">75 feet</quantity>
|
||
</elevation>
|
||
</materialsCitation>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31025F3FFFFFFA4ACC26A90140FFEF2" pageId="10" pageNumber="149" type="etymology">
|
||
<paragraph id="8BB57678FFFFFFA4ACC26A90140FFEF2" blockId="10.[151,1437,152,683]" pageId="10" pageNumber="149">
|
||
<emphasis id="B97EAA6AFFFFFFA4ACC26A901139FF17" bold="true" box="[199,334,225,251]" pageId="10" pageNumber="149">Etymology.</emphasis>
|
||
The name
|
||
<taxonomicName id="4C0A0DFBFFFFFFA4ADC96A911225FF16" box="[460,594,224,250]" class="Reptilia" family="Phrynosomatidae" genus="Holbrookia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="149" phylum="Chordata" rank="species" species="subcaudalis">
|
||
<emphasis id="B97EAA6AFFFFFFA4ADC96A911225FF16" box="[460,594,224,250]" italics="true" pageId="10" pageNumber="149">subcaudalis</emphasis>
|
||
</taxonomicName>
|
||
is derived from the latin word
|
||
<emphasis id="B97EAA6AFFFFFFA4AFAB6A9113A2FF16" box="[942,981,224,250]" italics="true" pageId="10" pageNumber="149">sub</emphasis>
|
||
which means under or below and
|
||
<emphasis id="B97EAA6AFFFFFFA4A95D6A9115EBFF16" box="[1368,1436,224,250]" italics="true" pageId="10" pageNumber="149">cauda</emphasis>
|
||
which means tail. This refers to the dark spots on the underside of the tail in this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31025F3FFFFFFA4ACC26B5911A6FE66" pageId="10" pageNumber="149" type="distribution">
|
||
<paragraph id="8BB57678FFFFFFA4ACC26B5911A6FE66" blockId="10.[151,1437,152,683]" pageId="10" pageNumber="149">
|
||
<emphasis id="B97EAA6AFFFFFFA4ACC26B591116FEAF" bold="true" box="[199,353,296,323]" pageId="10" pageNumber="149">Distribution.</emphasis>
|
||
<taxonomicName id="4C0A0DFBFFFFFFA4AD6E6B59120CFEAE" box="[363,635,296,322]" class="Reptilia" family="Phrynosomatidae" genus="Holbrookia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="149" phylum="Chordata" rank="species" species="subcaudalis">
|
||
<emphasis id="B97EAA6AFFFFFFA4AD6E6B59120CFEAE" box="[363,635,296,322]" italics="true" pageId="10" pageNumber="149">Holbrookia subcaudalis</emphasis>
|
||
</taxonomicName>
|
||
is composed of all populations south of the Balcones Escarpment in Texas and west to the Sierra Madre Oriental in
|
||
<collectingRegion id="49CEB89AFFFFFFA4AEB16B3D136BFE8A" box="[692,796,332,358]" country="Mexico" name="Coahuila de Zaragoza" pageId="10" pageNumber="149">Coahuila</collectingRegion>
|
||
,
|
||
<collectingRegion id="49CEB89AFFFFFFA4AF2D6B3C13C0FE8B" box="[808,951,333,359]" country="Mexico" name="Nuevo Leon" pageId="10" pageNumber="149">Nuevo Leon</collectingRegion>
|
||
, and
|
||
<collectingRegion id="49CEB89AFFFFFFA4AFF16B3D140FFE8A" box="[1012,1144,332,358]" country="Mexico" name="Tamaulipas" pageId="10" pageNumber="149">Tamaulipas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31D36E8FFFFFFA4A8816B3D14AAFE8A" box="[1156,1245,332,358]" name="Mexico" pageId="10" pageNumber="149">Mexico</collectingCountry>
|
||
. They are absent from areas with sandy soils.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31025F3FFFFFFA4ACC26BE5143BFD47" pageId="10" pageNumber="149" type="diagnosis">
|
||
<paragraph id="8BB57678FFFFFFA4ACC26BE5143BFD47" blockId="10.[151,1437,152,683]" pageId="10" pageNumber="149">
|
||
<emphasis id="B97EAA6AFFFFFFA4ACC26BE51135FE43" bold="true" box="[199,322,404,431]" pageId="10" pageNumber="149">Diagnosis.</emphasis>
|
||
Morphological description based on measurements and counts from
|
||
<specimenCount id="9D0CBDF1FFFFFFA4A86E6BE514AFFE42" box="[1131,1240,404,431]" pageId="10" pageNumber="149" type="adult">45 adults</specimenCount>
|
||
. This is a small, earless lizard with an average of 5 (1
|
||
<emphasis id="B97EAA6AFFFFFFA4AE3A6BC8123AFE3E" box="[575,589,441,466]" italics="true" pageId="10" pageNumber="149">–</emphasis>
|
||
10) black spots on the underside of the tail. The average snout-vent length (SVL) is
|
||
<quantity id="4CF2DB9DFFFFFFA4ACFA6BAD1139FE1B" box="[255,334,476,503]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="10" pageNumber="149" unit="mm" value="56.0">56 mm</quantity>
|
||
(31
|
||
<emphasis id="B97EAA6AFFFFFFA4AD7C6BAC11F0FE1A" box="[377,391,477,502]" italics="true" pageId="10" pageNumber="149">–</emphasis>
|
||
72), paravertebral and dorsolateral body blotches are often separated. An average of 0.7 (0
|
||
<emphasis id="B97EAA6AFFFFFFA4A9726BAC15F2FE1A" box="[1399,1413,477,502]" italics="true" pageId="10" pageNumber="149">–</emphasis>
|
||
6) blotches are fused out of an average of 6 (4
|
||
<emphasis id="B97EAA6AFFFFFFA4AE78687012FCFDF6" box="[637,651,513,538]" italics="true" pageId="10" pageNumber="149">–</emphasis>
|
||
9) blotches. The blotches form four rows of transverse mostly circular blotches. The dark blotches on the rear legs are circular in shape and do not form into bands with the average number of leg blotches being 8 (4
|
||
<emphasis id="B97EAA6AFFFFFFA4ADDA6838119AFD8E" box="[479,493,585,610]" italics="true" pageId="10" pageNumber="149">–</emphasis>
|
||
14). Most individuals have black lateral spots on the abdomen and these average 2.6 (0
|
||
<emphasis id="B97EAA6AFFFFFFA4ACAA681C10CAFD6A" box="[175,189,621,646]" italics="true" pageId="10" pageNumber="149">–</emphasis>
|
||
5). The average number of femoral pores on the left leg is 14 (10
|
||
<emphasis id="B97EAA6AFFFFFFA4AF88681C13ECFD6A" box="[909,923,621,646]" italics="true" pageId="10" pageNumber="149">–</emphasis>
|
||
19). The female body color is greenish yellow during the breeding season but they do not acquire orange on the throat in either sex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |