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<document id="1630584DDF3B9C3E4B57ECB0E6C4622D" ID-DOI="10.11646/zootaxa.4756.1.1" ID-GBIF-Dataset="4d48d52e-6505-4f62-909f-5da5c2cfb489" ID-ISSN="1175-5334" ID-Zenodo-Dep="3736895" ID-ZooBank="urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585767638433" checkinUser="carolina" docAuthor="Kluge, Nikita J." docDate="2020" docId="039787A6FFBC844F8CFBFE49DF79FF5E" docLanguage="en" docName="zootaxa.4756.1.1.pdf.imf" docOrigin="Zootaxa 4756 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Thraulodes telegraphicus Needham &amp; Murphy 1924" docType="treatment" docVersion="7" lastPageNumber="43" masterDocId="FFAEFFDEFF9D84648C6CFFE6DB3EFFA2" masterDocTitle="Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species" masterLastPageNumber="142" masterPageNumber="1" pageNumber="34" updateTime="1698734461607" updateUser="plazi">
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<mods:title id="1F83C5DECEE417F757237B9F3D4C08C6">Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species</mods:title>
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<treatment id="039787A6FFBC844F8CFBFE49DF79FF5E" ID-DOI="http://doi.org/10.5281/zenodo.3811777" ID-GBIF-Taxon="163420784" ID-Zenodo-Dep="3811777" LSID="urn:lsid:plazi:treatment:039787A6FFBC844F8CFBFE49DF79FF5E" httpUri="http://treatment.plazi.org/id/039787A6FFBC844F8CFBFE49DF79FF5E" lastPageId="43" lastPageNumber="43" pageId="33" pageNumber="34">
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<heading id="D0C981DCFFBC84458CFBFE49D807FE6B" bold="true" box="[151,825,430,457]" fontSize="10" level="2" pageId="33" pageNumber="34" reason="2">
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3.
<taxonomicName id="4C3E4D33FFBC84458CD9FE48D807FE6B" ID-CoL="56LL8" authority="Needham &amp; Murphy 1924" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[181,825,430,457]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFBC84458CD9FE48DAD7FE6B" bold="true" box="[181,489,430,457]" italics="true" pageId="33" pageNumber="34">Thraulodes telegraphicus</emphasis>
<bibRefCitation id="EFAF4B41FFBC84458D9DFE49D807FE6B" author="Needham, J. G. &amp; Murphy, H. E." box="[497,825,431,457]" pageId="33" pageNumber="34" pagination="1 - 79" refId="ref71682" refString="Needham, J. G. &amp; Murphy, H. E. (1924) Neotropical mayflies. Bulletin of the Lloyd Library, Entomological Series 4, 24, 1 - 79." type="journal article" year="1924">Needham &amp; Murphy 1924</bibRefCitation>
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<subSubSection id="C324653BFFBC84458CFBFE35DA0BFE4C" box="[151,309,467,494]" pageId="33" pageNumber="34" type="description">
<paragraph id="8B8136B0FFBC84458CFBFE35DA0BFE4C" blockId="33.[151,825,430,494]" box="[151,309,467,494]" pageId="33" pageNumber="34">
(
<figureCitation id="13052A35FFBC84458CF3FE35DA13FE4C" box="[159,301,467,494]" captionStart-0="FIGURES 66–69" captionStart-1="FIGURES 70–85" captionStart-2="FIGURES 86–93" captionStart-3="FIGURES 94–100" captionStart-4="FIGURES 101–105" captionStart-5="FIGURES 106–111" captionStart-6="FIGURES 112–120" captionStart-7="FIGURE 121" captionStartId-0="34.[151,264,1320,1345]" captionStartId-1="36.[151,264,188,213]" captionStartId-2="37.[151,264,188,213]" captionStartId-3="38.[151,264,1799,1824]" captionStartId-4="39.[151,264,1784,1809]" captionStartId-5="40.[151,264,1834,1859]" captionStartId-6="41.[151,264,1889,1914]" captionStartId-7="42.[151,250,1423,1448]" captionTargetBox-0="[151,1436,166,1301]" captionTargetBox-3="[151,1436,222,1780]" captionTargetBox-4="[151,1436,166,1764]" captionTargetBox-5="[159,1390,175,1807]" captionTargetBox-6="[151,1423,170,1862]" captionTargetBox-7="[173,1414,166,1404]" captionTargetId-0="figure@34.[151,1436,166,1301]" captionTargetId-1="figure@35.[151,1436,181,1986]" captionTargetId-2="figure@36.[151,1436,342,2018]" captionTargetId-3="figure@38.[151,1436,222,1780]" captionTargetId-4="figure@39.[151,1436,166,1764]" captionTargetId-5="figure@40.[151,1397,166,1814]" captionTargetId-6="figure@41.[151,1436,166,1869]" captionTargetId-7="figure@42.[173,1414,166,1404]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionTargetPageId-2="36" captionTargetPageId-3="38" captionTargetPageId-4="39" captionTargetPageId-5="40" captionTargetPageId-6="41" captionTargetPageId-7="42" captionText-0="FIGURES 66–69. Thraulodes telegraphicus. 66–67, last instar larvae with intensive cuticular coloration and poorly visible hypodermal coloration; 68, younger larva with light cuticle and well visible blackish hypodermal markings on abdomen; 69, claw. Abbreviations: artic.p., articulatory portion; rigid p., rigid portion of claw." captionText-1="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." captionText-2="FIGURES 86–93. Thraulodes telegraphicus. 86, median longitudinal section of labrum in usual condition; 87, the same, in spread condition; 88, labrum in spread condition in dorsal view (setal rows shown as dotted lines); 89, transverse section of hind tibia; 90–93, tergalii I, III, VI and VII. Abbreviations: anter.r, anterior transverse row of setae; dent., five denticles separated by six sensilla; poster.r, posterior transverse row of setae; i.f, inner field of stout pointed setae; i-a.h, inner-anterior recurved hairs; i-a.r, inner-anterior row of stout setae; o-a.r, outer-anterior row of stout setae; o.h, outer hairs; o-p.r, outer-posterior row of stout setae; p.r, row of posterior pointed pectinate setae." captionText-3="FIGURES 94–100. Thraulodes telegraphicus. 94–95, right half of subimaginal exuviae of mesonotum of two different specimens; 96, apex of fore wing; 97–98, male imago in lateral and ventral view; 99, abdominal terga and sterna of male imago (black dots are algae under cuticle); 100, female imago. Abbreviations: acz.ms, achromozone of medioscutum; acz.sms, achromozone of submedioscutum; cz.ms, chromozone of medioscutum; cz.sms, chromozone of submedioscutum; mp.s, medioparapsidal suture; ms.cz.s, medioscutal chromozonal suture; sms.cz.s, submedioscutal chromozonal suture." captionText-4="FIGURES 101–105. Thraulodes telegraphicus. 101–102, genitalia of the same specimen; 103, penis of another specimen; 104– 105, part of penis with telopenis forming spear-like roll.Abbreviations: bl, basal lobe; dp, dorsal projection of styliger; gd, gonoduct; gs1, gs2, gs3, gs4, segments of gonostylus; m.gs, gonostylar muscle; m.lat, lateral penial muscle; m.med, median penial muscle; m.s, styligeral muscle; m-v.r, medio-ventral ridge (recurved flap); sp, seminal pump; tp, spear-like roll (telopenis)." captionText-5="FIGURES 106–111. Thraulodes telegraphicus. 106, protogonostyli of last instar larva (hypoderm of gonostyli and styliger and hidden cuticle of protopenis shown by interrupted lines); 107, enlarged cuticle of protopenis and hypoderm of developing penis inside; 108–109, the same, at later stage of development; 110–111, genital of male imago (110, styliger ventrally; 111, penis dorsally at left and ventrally at right). Abbreviation: bl, basal lobe; dp, dorsal projection of styliger; gd, gonoduct; m.gs, gonostylar muscle; m.med, median penial muscle; m.s, styligeral muscle; m-v.r, medio-ventral ridge (recurved flap); sp, seminal pump; tp, spear-like roll (telopenis)." captionText-6="FIGURES 112–120. Thraulodes telegraphicus. 112, genitalia of male imago in lateral view; 113, apex of penis with spear-like roll in dorsal view; 114, genitalia of the same specimen in ventral view; 115, their subimaginal exuviae at the same magnification; 116, hind wing; 117–120, penis of four different specimens. Abbreviations: dp, dorsal projection of styliger; tp, spear-like roll (telopenis)." captionText-7="FIGURE 121. Thraulodes telegraphicus, eggs." figureDoi-0="http://doi.org/10.5281/zenodo.3736917" figureDoi-1="http://doi.org/10.5281/zenodo.3736919" figureDoi-2="http://doi.org/10.5281/zenodo.3736921" figureDoi-3="http://doi.org/10.5281/zenodo.3736923" figureDoi-4="http://doi.org/10.5281/zenodo.3736925" figureDoi-5="http://doi.org/10.5281/zenodo.3736927" figureDoi-6="http://doi.org/10.5281/zenodo.3736929" figureDoi-7="http://doi.org/10.5281/zenodo.3736931" httpUri-0="https://zenodo.org/record/3736917/files/figure.png" httpUri-1="https://zenodo.org/record/3736919/files/figure.png" httpUri-2="https://zenodo.org/record/3736921/files/figure.png" httpUri-3="https://zenodo.org/record/3736923/files/figure.png" httpUri-4="https://zenodo.org/record/3736925/files/figure.png" httpUri-5="https://zenodo.org/record/3736927/files/figure.png" httpUri-6="https://zenodo.org/record/3736929/files/figure.png" httpUri-7="https://zenodo.org/record/3736931/files/figure.png" pageId="33" pageNumber="34">Figs 66–121</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFBC84458CFBFDF1D86BFD0D" pageId="33" pageNumber="34" type="reference_group">
<paragraph id="8B8136B0FFBC84458CFBFDF1DEA2FD8C" blockId="33.[151,1437,534,1984]" box="[151,1436,534,559]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB2E119EFFBC84458CFBFDF1DEA2FD8C" box="[151,1436,534,559]" pageId="33" pageNumber="34">
<treatmentCitation id="0A9F10A1FFBC84458CFBFDF1D9E4FD8D" author="Needham, J. G. &amp; Murphy, H. E." box="[151,730,534,559]" page="41" pageId="33" pageNumber="34" year="1924">
<taxonomicName id="4C3E4D33FFBC84458CFBFDF1D9E4FD8D" ID-CoL="56LL8" authority="Needham &amp; Murphy 1924: 41" authorityName="Needham &amp; Murphy" authorityPageNumber="41" authorityYear="1924" box="[151,730,534,559]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFBC84458CFBFDF1DAA2FD8D" box="[151,412,535,559]" italics="true" pageId="33" pageNumber="34">Thraulodes telegraphicus</emphasis>
<bibRefCitation id="EFAF4B41FFBC84458DCFFDF0D9E4FD8D" author="Needham, J. G. &amp; Murphy, H. E." box="[419,730,534,559]" pageId="33" pageNumber="34" pagination="1 - 79" refId="ref71682" refString="Needham, J. G. &amp; Murphy, H. E. (1924) Neotropical mayflies. Bulletin of the Lloyd Library, Entomological Series 4, 24, 1 - 79." type="journal article" year="1924">Needham &amp; Murphy 1924: 41</bibRefCitation>
</taxonomicName>
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, figs 99, 102 (
<specimenCount id="9D38FD39FFBC84458F03FDF1D8BCFD8D" box="[879,898,535,559]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
and
<specimenCount id="9D38FD39FFBC84458FD9FDF1D8F6FD8C" box="[949,968,535,558]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
imago);
<treatmentCitation id="0A9F10A1FFBC84458849FDF1DFEBFD8D" author="Traver, J. R." box="[1061,1237,534,559]" page="428" pageId="33" pageNumber="34" year="1946">
<bibRefCitation id="EFAF4B41FFBC84458849FDF1DFEBFD8D" author="Traver, J. R." box="[1061,1237,534,559]" pageId="33" pageNumber="34" pagination="418 - 436" refId="ref72450" refString="Traver, J. R. (1946) Notes on Neotropical mayflies. Part I. Family Baetidae, subfamily Leptophlebiinae. Revista de Entomologia, 17, 418 - 436." type="journal article" year="1946">Traver 1946: 428</bibRefCitation>
</treatmentCitation>
(
<typeStatus id="54858812FFBC8445888FFDF0DE7EFD8D" box="[1251,1344,534,559]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
, synony-
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136B0FFBC84458CABFDD0D85EFDEC" blockId="33.[151,1437,534,1984]" box="[199,864,565,590]" pageId="33" pageNumber="34">
my);
<bibRefCitation id="EFAF4B41FFBC84458C91FDD0D910FDEC" author="Traver, J. R. &amp; Edmunds, G. F. Jr." box="[253,558,565,590]" pageId="33" pageNumber="34" pagination="349 - 395" refId="ref72596" refString="Traver, J. R. &amp; Edmunds, G. F. Jr. (1967) A revision of the genus Thraulodes (Ephemeroptera: Leptophlebiidae). Miscellaneous Publications of the Entomological Society of America, 5 (8), 349 - 395." type="journal article" year="1967">Traver &amp; Edmunds 1967: 379</bibRefCitation>
, figs 40, 60 (
<specimenCount id="9D38FD39FFBC84458EDFFDD0D9F8FDEC" box="[691,710,566,590]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
and
<specimenCount id="9D38FD39FFBC84458E9BFDD0D834FDEF" box="[759,778,566,589]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
imago).
</paragraph>
<paragraph id="8B8136B0FFBC84458CFBFDB2DEA2FDCE" blockId="33.[151,1437,534,1984]" box="[151,1436,596,621]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB2E119EFFBC84458CFBFDB2DEA2FDCE" box="[151,1436,596,621]" pageId="33" pageNumber="34">
<treatmentCitation id="0A9F10A1FFBC84458CFBFDB2D99CFDCE" author="Needham, J. G. &amp; Murphy, H. E." box="[151,674,596,621]" page="42" pageId="33" pageNumber="34" year="1924">
<taxonomicName id="4C3E4D33FFBC84458CFBFDB2D99CFDCE" authority="Needham &amp; Murphy 1924: 42" authorityName="Needham &amp; Murphy" authorityPageNumber="42" authorityYear="1924" box="[151,674,596,621]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="plicatus">
<emphasis id="B94AEAA2FFBC84458CFBFDB2DA5CFDCE" box="[151,354,596,620]" italics="true" pageId="33" pageNumber="34">Thraulodes plicatus</emphasis>
<bibRefCitation id="EFAF4B41FFBC84458D05FDB2D99CFDCE" author="Needham, J. G. &amp; Murphy, H. E." box="[361,674,596,621]" pageId="33" pageNumber="34" pagination="1 - 79" refId="ref71682" refString="Needham, J. G. &amp; Murphy, H. E. (1924) Neotropical mayflies. Bulletin of the Lloyd Library, Entomological Series 4, 24, 1 - 79." type="journal article" year="1924">Needham &amp; Murphy 1924: 42</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 100 (
<specimenCount id="9D38FD39FFBC84458F60FDB2D821FDCE" box="[780,799,596,620]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
and
<specimenCount id="9D38FD39FFBC84458F3EFDB3D85BFDCE" box="[850,869,597,620]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
subimago) (
<typeStatus id="54858812FFBC84458F8AFDB2DF7EFDCF" box="[998,1088,596,621]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
redescribed and synonymy estab-
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136B0FFBC84458CABFD94DA8AFD29" blockId="33.[151,1437,534,1984]" box="[199,436,626,651]" pageId="33" pageNumber="34">
lished by
<bibRefCitation id="EFAF4B41FFBC84458D45FD95DA97FD29" author="Traver, J. R." box="[297,425,626,651]" pageId="33" pageNumber="34" pagination="418 - 436" refId="ref72450" refString="Traver, J. R. (1946) Notes on Neotropical mayflies. Part I. Family Baetidae, subfamily Leptophlebiinae. Revista de Entomologia, 17, 418 - 436." type="journal article" year="1946">Traver 1946</bibRefCitation>
).
</paragraph>
<paragraph id="8B8136B0FFBC84458CFBFD71D86BFD0D" blockId="33.[151,1437,534,1984]" box="[151,853,662,687]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB2E119EFFBC84458CFBFD71D86BFD0D" box="[151,853,662,687]" pageId="33" pageNumber="34">
<taxonomicName id="4C3E4D33FFBC84458CFBFD71DA34FD0D" ID-CoL="648C7" authorityName="Ulmer" authorityYear="1920" box="[151,266,663,687]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAA2FFBC84458CFBFD71DA34FD0D" box="[151,266,663,687]" italics="true" pageId="33" pageNumber="34">Thraulodes</emphasis>
</taxonomicName>
sp. 1, sp. 2, sp. 4, sp. 6:
<treatmentCitation id="0A9F10A1FFBC84458E68FD70D9FEFD0D" author="Roback, S. S." box="[516,704,662,687]" page="145" pageId="33" pageNumber="34" year="1966">
<bibRefCitation id="EFAF4B41FFBC84458E68FD70D9FEFD0D" author="Roback, S. S." box="[516,704,662,687]" pageId="33" pageNumber="34" pagination="129 - 199" refId="ref72121" refString="Roback, S. S. (1966) The Catherwood Foundation Peruvia-Amazon expedition. VI- Ephemeroptera nymphs. Monographs of the Academy of Natural Sciences of Philadelphia, No. 14, 129 - 199." type="book chapter" year="1966">Roback 1966: 145</bibRefCitation>
</treatmentCitation>
, pl. 13 (larva).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFBC84458CFBFD5FD990FBB5" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8B8136B0FFBC84458CFBFD5FD990FBB5" blockId="33.[151,1437,534,1984]" pageId="33" pageNumber="34">
<emphasis id="B94AEAA2FFBC84458CFBFD5FDABEFD76" bold="true" box="[151,384,697,724]" pageId="33" pageNumber="34">Material examined.</emphasis>
<materialsCitation id="3B563CEDFFBC84458DE9FD5CDEA2FD55" ID-GBIF-Occurrence="2594509922" collectingDate="2006-01-05" collectingDateMax="2006-01-10" collectingDateMin="2006-01-05" collectorName="N. Kluge" country="Peru" county="Provincia Satipo" elevation="350" latitude="-7.2" location="Rio Sotorani" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" pageId="33" pageNumber="34" specimenCount="8" specimenCount-female="2" specimenCount-larva="4" specimenCount-male="2" stateProvince="Region Junin">
<collectingCountry id="F3297620FFBC84458DE9FD5CDAEFFD76" box="[389,465,698,724]" name="Peru" pageId="33" pageNumber="34">PERU</collectingCountry>
:
<collectingRegion id="49FAF852FFBC84458DB7FD5FD94EFD71" box="[475,624,697,723]" country="Peru" name="Junin" pageId="33" pageNumber="34">Region Junin</collectingRegion>
,
<collectingCounty id="62E04E3CFFBC84458E17FD5FD806FD71" box="[635,824,697,723]" pageId="33" pageNumber="34">Provincia Satipo</collectingCounty>
:
<collectingMunicipality id="6BE5ACCAFFBC84458F2EFD5FDF1DFD71" box="[834,1059,697,723]" pageId="33" pageNumber="34">Rio Satipo in Satipo</collectingMunicipality>
,
<date id="FF801070FFBC84458842FD5FDFB0FD76" box="[1070,1166,697,724]" pageId="33" pageNumber="34" value="2006-01-05">
<collectingDate id="EFC4E998FFBC84458842FD5FDFB0FD76" box="[1070,1166,697,724]" pageId="33" pageNumber="34" value="2006-01-05">5.I.2006</collectingDate>
</date>
, coll.
<collectorName id="26CB5366FFBC844588A3FD5CDE07FD71" box="[1231,1337,697,724]" pageId="33" pageNumber="34">N. Kluge</collectorName>
: 1 L-S
<specimenCount id="9D38FD39FFBC844589E8FD5FDEA6FD71" box="[1412,1432,697,723]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
,
<specimenCount id="9D38FD39FFBC84458CFBFD3BDBCAFD55" box="[151,244,733,759]" pageId="33" pageNumber="34" type="larva">4 larvae</specimenCount>
;
<location id="8EE1606BFFBC84458D6CFD3BDAB1FD55" LSID="urn:lsid:plazi:treatment:039787A6FFBC844F8CFBFE49DF79FF5E:8EE1606BFFBC84458D6CFD3BDAB1FD55" box="[256,399,733,759]" country="Peru" county="Provincia Satipo" latitude="-7.2" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" name="Rio Sotorani" pageId="33" pageNumber="34" stateProvince="Region Junin">Rio Sotorani</location>
near
<location id="8EE1606BFFBC84458DA1FD3BD905FD55" LSID="urn:lsid:plazi:treatment:039787A6FFBC844F8CFBFE49DF79FF5E:8EE1606BFFBC84458DA1FD3BD905FD55" box="[461,571,733,759]" country="Peru" county="Provincia Satipo" latitude="-7.2" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" name="Pichanaki" pageId="33" pageNumber="34" stateProvince="Region Junin">Pichanaki</location>
(
<geoCoordinate id="EE0A5077FFBC84458E20FD3BD994FD5A" box="[588,682,733,760]" degrees="10" direction="south" minutes="57" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="-10.95">10°57’S</geoCoordinate>
,
<geoCoordinate id="EE0A5077FFBC84458EDAFD3BD81EFD5A" box="[694,800,733,760]" degrees="74" direction="west" minutes="52" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="-74.86667">74°52’W</geoCoordinate>
),
<date id="FF801070FFBC84458F59FD3BD880FD55" box="[821,958,733,759]" pageId="33" pageNumber="34" value="2006-01-08" valueMax="2006-01-10" valueMin="2006-01-08">
<collectingDate id="EFC4E998FFBC84458F59FD3BD880FD55" box="[821,958,733,759]" pageId="33" pageNumber="34" value="2006-01-08" valueMax="2006-01-10" valueMin="2006-01-08">8–10.I.2006</collectingDate>
</date>
, coll.
<collectorName id="26CB5366FFBC8445886FFD38DF50FD55" box="[1027,1134,733,760]" pageId="33" pageNumber="34">N. Kluge</collectorName>
: 1 L-S-I
<specimenCount id="9D38FD39FFBC844588B8FD3BDFDCFD55" box="[1236,1250,733,759]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
, 1 S-I
<specimenCount id="9D38FD39FFBC84458945FD3BDE03FD55" box="[1321,1341,733,759]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
, 2 S-I
<specimenCount id="9D38FD39FFBC844589EBFD3BDEA9FD55" box="[1415,1431,733,759]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
,
</materialsCitation>
<materialsCitation id="3B563CEDFFBC84458CFBFCE7DA95FC9D" ID-GBIF-Occurrence="2594509918" collectingDate="2013-09-10" collectingDateMax="2013-09-14" collectingDateMin="2013-09-10" collectorName="N. Kluge &amp; L. Sheyko" country="Peru" county="Provincia Satipo" elevation="350" latitude="-7.2" location="Puerto Prado" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" pageId="33" pageNumber="34" specimenCount="29" specimenCount-female="3" specimenCount-larva="24" specimenCount-male="2" stateProvince="Junin">
<specimenCount id="9D38FD39FFBC84458CFBFCE7DA3DFCB9" box="[151,259,769,795]" pageId="33" pageNumber="34" type="larva">24 larvae</specimenCount>
;
<location id="8EE1606BFFBC84458D62FCE4DAA1FCB9" LSID="urn:lsid:plazi:treatment:039787A6FFBC844F8CFBFE49DF79FF5E:8EE1606BFFBC84458D62FCE4DAA1FCB9" box="[270,415,769,796]" country="Peru" county="Provincia Satipo" latitude="-7.2" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" name="Puerto Prado" pageId="33" pageNumber="34" stateProvince="Junin">Puerto Prado</location>
(
<geoCoordinate id="EE0A5077FFBC84458DDCFCE7D933FCB9" box="[432,525,769,795]" degrees="11" direction="south" minutes="09" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="-11.15">11°09’S</geoCoordinate>
,
<geoCoordinate id="EE0A5077FFBC84458E76FCE7D9BAFCB9" box="[538,644,769,795]" degrees="74" direction="west" minutes="14" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="-74.23333">74°14’W</geoCoordinate>
),
<date id="FF801070FFBC84458EF4FCE7D87BFCB9" box="[664,837,769,795]" pageId="33" pageNumber="34" value="2013-09-10" valueMax="2013-09-14" valueMin="2013-09-10">
<collectingDate id="EFC4E998FFBC84458EF4FCE7D87BFCB9" box="[664,837,769,795]" pageId="33" pageNumber="34" value="2013-09-10" valueMax="2013-09-14" valueMin="2013-09-10">10–14.IX.2013</collectingDate>
</date>
, coll.
<collectorName id="26CB5366FFBC84458FE5FCE4D8D1FCB9" box="[905,1007,769,796]" pageId="33" pageNumber="34">N. Kluge</collectorName>
&amp;
<collectorName id="26CB5366FFBC84458878FCE4DFB2FCB9" box="[1044,1164,769,796]" pageId="33" pageNumber="34">L. Sheyko</collectorName>
: 11 L-S-I
<specimenCount id="9D38FD39FFBC84458896FCE7DE33FCB9" box="[1274,1293,769,795]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
, 5 L-S-I
<specimenCount id="9D38FD39FFBC84458918FCE7DEBCFCB9" box="[1396,1410,769,795]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
, 6 L-S
<specimenCount id="9D38FD39FFBC84458CA5FCC3DBE6FC9D" box="[201,216,805,831]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
, 10 L/S
<specimenCount id="9D38FD39FFBC84458D5EFCC3DA78FC9D" box="[306,326,805,831]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
, 6 L/S
<specimenCount id="9D38FD39FFBC84458DFAFCC3DA98FC9D" box="[406,422,805,831]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
,
</materialsCitation>
<materialsCitation id="3B563CEDFFBC84458DDDFCC3DA59FCC1" ID-GBIF-Occurrence="2594509928" collectingDate="2006-01-06" collectingDateMax="2013-09-14" collectingDateMin="2006-01-05" collectorName="A. Petrov" country="Peru" county="Provincia Satipo" elevation="350" latitude="-7.2" location="Rio Shima" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" pageId="33" pageNumber="34" specimenCount="63" specimenCount-female="11" specimenCount-larva="63" specimenCount-male="9" stateProvince="Junin">
<specimenCount id="9D38FD39FFBC84458DDDFCC3D922FC9D" box="[433,540,805,831]" pageId="33" pageNumber="34" type="larva">63 larvae</specimenCount>
;
<location id="8EE1606BFFBC84458E4BFCC3D9A0FC9D" LSID="urn:lsid:plazi:treatment:039787A6FFBC844F8CFBFE49DF79FF5E:8EE1606BFFBC84458E4BFCC3D9A0FC9D" box="[551,670,805,831]" country="Peru" county="Provincia Satipo" latitude="-7.2" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" name="Rio Shima" pageId="33" pageNumber="34" stateProvince="Junin">Rio Shima</location>
<locationDeviation id="9A332F92FFBC84458EC8FCC3D8B0FC9D" box="[676,910,805,832]" location="Puerto Prado" pageId="33" pageNumber="34">
<quantity id="4CC69B55FFBC84458EC8FCC3D9E3FC9D" box="[676,733,805,832]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="33" pageNumber="34" unit="km" value="5.0">5 km</quantity>
E Puerto Prado
</locationDeviation>
, (
<geoCoordinate id="EE0A5077FFBC84458FCFFCC3DF3EFC9D" box="[931,1024,805,831]" degrees="11" direction="south" minutes="07" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="-11.116667">11°07’S</geoCoordinate>
,
<geoCoordinate id="EE0A5077FFBC84458867FCC3DF4BFC9D" box="[1035,1141,805,831]" degrees="74" direction="west" minutes="13" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="-74.21667">74°13’W</geoCoordinate>
), alt.
<quantity id="4CC69B55FFBC844588DFFCC3DFCBFCE2" box="[1203,1269,805,832]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="33" pageNumber="34" unit="m" value="350.0">
<elevation id="0013D183FFBC844588DFFCC3DFCBFCE2" box="[1203,1269,805,832]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="33" pageNumber="34" unit="m" value="350.0">350m</elevation>
</quantity>
,
<date id="FF801070FFBC8445896CFCC3DE5EFC9D" box="[1280,1376,805,831]" pageId="33" pageNumber="34" value="2006-01-06">
<collectingDate id="EFC4E998FFBC8445896CFCC3DE5EFC9D" box="[1280,1376,805,831]" pageId="33" pageNumber="34" value="2006-01-06">6.I.2006</collectingDate>
</date>
, coll.
<collectorName id="26CB5366FFBC84458CFBFCACDA36FCC6" box="[151,264,842,868]" pageId="33" pageNumber="34">A. Petrov</collectorName>
: 2 S(m);
</materialsCitation>
<materialsCitation id="3B563CEDFFBC84458D02FCAFD9CDFC09" ID-GBIF-Occurrence="2594509913" collectingDate="2006-01-13" collectingDateMax="2006-01-25" collectingDateMin="2006-01-13" collectorName="Provincia Leoncio Prado &amp; Cayumba &amp; Puente Perez &amp; N. Kluge" country="Peru" county="Provincia Satipo" elevation="350" latitude="-7.2" location="Rio Chunatahua" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" pageId="33" pageNumber="34" specimenCount="8" specimenCount-female="4" specimenCount-larva="199" specimenCount-male="4" stateProvince="Region Huanuco">
<collectingRegion id="49FAF852FFBC84458D02FCAFD911FCC6" box="[366,559,841,868]" country="Peru" name="Huanuco" pageId="33" pageNumber="34">Region Huanuco</collectingRegion>
,
<collectorName id="26CB5366FFBC84458E56FCAFD86AFCC1" box="[570,852,841,867]" pageId="33" pageNumber="34">Provincia Leoncio Prado</collectorName>
,
<location id="8EE1606BFFBC84458F33FCAFDF2BFCC1" LSID="urn:lsid:plazi:treatment:039787A6FFBC844F8CFBFE49DF79FF5E:8EE1606BFFBC84458F33FCAFDF2BFCC1" box="[863,1045,841,867]" country="Peru" county="Provincia Satipo" latitude="-7.2" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" name="Rio Chunatahua" pageId="33" pageNumber="34" stateProvince="Region Huanuco">Rio Chunatahua</location>
near
<collectorName id="26CB5366FFBC84458838FCAFDF81FCC1" box="[1108,1215,841,867]" pageId="33" pageNumber="34">Cayumba</collectorName>
(
<geoCoordinate id="EE0A5077FFBC844588A3FCAFDE21FCC1" box="[1231,1311,841,867]" degrees="9" direction="south" minutes="30" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="-9.5">9°30’S</geoCoordinate>
,
<geoCoordinate id="EE0A5077FFBC84458940FCAFDEAAFCC6" box="[1324,1428,841,868]" degrees="75" direction="west" minutes="57" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="-75.95">75°57’W</geoCoordinate>
) and
<location id="8EE1606BFFBC84458CA4FC8BDA69FC25" LSID="urn:lsid:plazi:treatment:039787A6FFBC844F8CFBFE49DF79FF5E:8EE1606BFFBC84458CA4FC8BDA69FC25" box="[200,343,877,903]" country="Peru" county="Provincia Satipo" latitude="-7.2" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" name="Rio Cantaria" pageId="33" pageNumber="34" stateProvince="Region Huanuco">Rio Cantaria</location>
near
<collectorName id="26CB5366FFBC84458DFBFC88D917FC2A" box="[407,553,878,904]" pageId="33" pageNumber="34">Puente Perez</collectorName>
(
<geoCoordinate id="EE0A5077FFBC84458E57FC8BD9B5FC25" box="[571,651,877,903]" degrees="9" direction="south" minutes="21" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="-9.35">9°21’S</geoCoordinate>
,
<geoCoordinate id="EE0A5077FFBC84458EF4FC8BD83CFC2A" box="[664,770,877,904]" degrees="75" direction="west" minutes="58" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="-75.96667">75°58’W</geoCoordinate>
),
<date id="FF801070FFBC84458F7BFC8BD891FC2A" box="[791,943,877,904]" pageId="33" pageNumber="34" value="2006-01-13" valueMax="2006-01-25" valueMin="2006-01-13">
<collectingDate id="EFC4E998FFBC84458F7BFC8BD891FC2A" box="[791,943,877,904]" pageId="33" pageNumber="34" value="2006-01-13" valueMax="2006-01-25" valueMin="2006-01-13">13–25.I.2006</collectingDate>
</date>
, coll.,
<collectorName id="26CB5366FFBC84458F97FC88DF59FC25" box="[1019,1127,877,904]" pageId="33" pageNumber="34">N. Kluge</collectorName>
: 20 L-S-I
<specimenCount id="9D38FD39FFBC844588BBFC8BDFD4FC25" box="[1239,1258,877,903]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
, 2 L-S/I
<specimenCount id="9D38FD39FFBC84458920FC8BDE5EFC25" box="[1356,1376,877,903]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
, 7 L- S
<specimenCount id="9D38FD39FFBC84458CCBFC77DB83FC09" box="[167,189,913,939]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
, 27 L-S-I
<specimenCount id="9D38FD39FFBC84458D59FC77DA7DFC09" box="[309,323,913,939]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
, 2 L-S/I
<specimenCount id="9D38FD39FFBC84458DC5FC77DA86FC09" box="[425,440,913,939]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
, 15 L-S
<specimenCount id="9D38FD39FFBC84458E76FC77D917FC09" box="[538,553,913,939]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
, 2 L/S
<specimenCount id="9D38FD39FFBC84458E14FC77D9B2FC09" box="[632,652,913,939]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
, 2 L/S
<specimenCount id="9D38FD39FFBC84458EB3FC77D9D1FC09" box="[735,751,913,939]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
,
</materialsCitation>
<materialsCitation id="3B563CEDFFBC84458E90FC77D958FC6D" ID-GBIF-Occurrence="2594509914" collectingDate="2006-01-05" collectingDateMax="2013-09-14" collectingDateMin="2006-01-05" collectorName="N. Kluge &amp; L. Sheyko &amp; A. Petrov &amp; Provincia Leoncio Prado &amp; Cayumba &amp; Puente Perez &amp; Provincia Ambo &amp; Ambo &amp; Huacar &amp; Provincia &amp; Aguas Termales" country="Peru" county="Provincia Satipo" elevation="350" latitude="-7.2" location="Rio Sotorani" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" pageId="33" pageNumber="34" specimenCount="219" specimenCount-female="11" specimenCount-larva="199" specimenCount-male="9" stateProvince="Junin">
<specimenCount id="9D38FD39FFBC84458E90FC77D856FC09" box="[764,872,913,939]" pageId="33" pageNumber="34" type="larva">74 larvae</specimenCount>
;
<collectingRegion id="49FAF852FFBC84458F18FC77DF0BFC0E" box="[884,1077,913,940]" country="Peru" name="Huanuco" pageId="33" pageNumber="34">Region Huanuco</collectingRegion>
,
<collectorName id="26CB5366FFBC8445882DFC77DFC2FC09" box="[1089,1276,913,939]" pageId="33" pageNumber="34">Provincia Ambo</collectorName>
:
<location id="8EE1606BFFBC84458964FC77DEA2FC0E" LSID="urn:lsid:plazi:treatment:039787A6FFBC844F8CFBFE49DF79FF5E:8EE1606BFFBC84458964FC77DEA2FC0E" box="[1288,1436,913,940]" country="Peru" county="Provincia Satipo" latitude="-7.2" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" name="river Huertas" pageId="33" pageNumber="34" stateProvince="Junin">river Huertas</location>
near
<collectorName id="26CB5366FFBC84458CA1FC53DA28FC6D" box="[205,278,949,975]" pageId="33" pageNumber="34">Ambo</collectorName>
,
<date id="FF801070FFBC84458D4CFC53DA8BFC6D" box="[288,437,949,975]" pageId="33" pageNumber="34" value="2013-08-28">
<collectingDate id="EFC4E998FFBC84458D4CFC53DA8BFC6D" box="[288,437,949,975]" pageId="33" pageNumber="34" value="2013-08-28">28.VIII.2013</collectingDate>
</date>
, coll.
<collectorName id="26CB5366FFBC84458D94FC50D95DFC6D" box="[504,611,949,976]" pageId="33" pageNumber="34">N. Kluge</collectorName>
:
</materialsCitation>
<materialsCitation id="3B563CEDFFBC84458E01FC53DE44FC6D" ID-GBIF-Occurrence="2594509910" box="[621,1402,949,976]" collectingDate="2006-01-05" collectingDateMax="2013-09-14" collectingDateMin="2006-01-05" collectorName="N. Kluge &amp; L. Sheyko &amp; A. Petrov &amp; Provincia Leoncio Prado &amp; Cayumba &amp; Puente Perez &amp; Provincia Ambo &amp; Ambo &amp; Huacar &amp; Provincia &amp; Aguas Termales" country="Peru" county="Provincia Satipo" elevation="350" latitude="-7.2" location="Rio Sotorani" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" pageId="33" pageNumber="34" specimenCount="219" specimenCount-female="11" specimenCount-larva="199" specimenCount-male="9" stateProvince="Junin">
<specimenCount id="9D38FD39FFBC84458E01FC53D9F7FC6D" box="[621,713,949,975]" pageId="33" pageNumber="34" type="larva">2 larvae</specimenCount>
;
<collectorName id="26CB5366FFBC84458EB8FC50D81BFC72" box="[724,805,950,976]" pageId="33" pageNumber="34">Huacar</collectorName>
(
<geoCoordinate id="EE0A5077FFBC84458F59FC53D8BBFC6D" box="[821,901,949,975]" degrees="10" direction="south" minutes="9" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="-10.15">10°9’S</geoCoordinate>
,
<geoCoordinate id="EE0A5077FFBC84458FFDFC53D8C4FC72" box="[913,1018,949,976]" degrees="76" direction="west" minutes="15" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="-76.25">76°15’W</geoCoordinate>
),
<date id="FF801070FFBC84458863FC53DF9EFC72" box="[1039,1184,949,976]" pageId="33" pageNumber="34" value="2013-09-01" valueMax="2013-09-05" valueMin="2013-09-01">
<collectingDate id="EFC4E998FFBC84458863FC53DF9EFC72" box="[1039,1184,949,976]" pageId="33" pageNumber="34" value="2013-09-01" valueMax="2013-09-05" valueMin="2013-09-01">1–5.IX.2013</collectingDate>
</date>
: 1 L-S-I
<specimenCount id="9D38FD39FFBC84458969FC53DE2DFC6D" box="[1285,1299,949,975]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
, 1 L-S
<specimenCount id="9D38FD39FFBC84458909FC53DE4AFC6D" box="[1381,1396,949,975]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
,
</materialsCitation>
<materialsCitation id="3B563CEDFFBC844589ECFC53D990FBB5" ID-GBIF-Occurrence="2594509933" collectingDate="2013-07-26" collectingDateMax="2013-08-04" collectingDateMin="2013-07-26" collectorName="Provincia &amp; Aguas Termales &amp; N. Kluge &amp; L. Sheyko" country="Peru" county="Provincia Satipo" elevation="350" latitude="-7.2" location="Region Loreto" longLatPrecision="1303" longitude="-74.95" municipality="Rio Satipo in Satipo" pageId="33" pageNumber="34" specimenCount="33" specimenCount-female="11" specimenCount-larva="32" specimenCount-male="1" stateProvince="Region Loreto">
<specimenCount id="9D38FD39FFBC844589ECFC53DBE1FC51" pageId="33" pageNumber="34" type="larva">32 larvae</specimenCount>
;
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,
<collectorName id="26CB5366FFBC84458DCDFC3FD932FC51" box="[417,524,985,1011]" pageId="33" pageNumber="34">Provincia</collectorName>
<collectingRegion id="49FAF852FFBC84458E79FC3FD951FC51" box="[533,623,985,1011]" country="Peru" name="Ucayali" pageId="33" pageNumber="34">Ucayali</collectingRegion>
,
<collectorName id="26CB5366FFBC84458E16FC3CD80DFC51" box="[634,819,985,1012]" pageId="33" pageNumber="34">Aguas Termales</collectorName>
,
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NNE Contamana
</locationDeviation>
(
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,
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),
<date id="FF801070FFBC84458928FC3FDA20FBB5" pageId="33" pageNumber="34" value="2013-07-26" valueMax="2013-08-04" valueMin="2013-07-26">
<collectingDate id="EFC4E998FFBC84458928FC3FDA20FBB5" pageId="33" pageNumber="34" value="2013-07-26" valueMax="2013-08-04" valueMin="2013-07-26">26.VII– 4.VIII.2013</collectingDate>
</date>
, coll.
<collectorName id="26CB5366FFBC84458D0DFC18DAF6FBB5" box="[353,456,1021,1048]" pageId="33" pageNumber="34">N. Kluge</collectorName>
&amp;
<collectorName id="26CB5366FFBC84458D80FC18D95DFBB5" box="[492,611,1021,1048]" pageId="33" pageNumber="34">L. Sheyko</collectorName>
: 1 S
<specimenCount id="9D38FD39FFBC84458EFFFC1BD994FBB5" box="[659,682,1021,1047]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFBC844E8CABFBC7DA80F99B" lastPageId="42" lastPageNumber="43" pageId="33" pageNumber="34" type="description">
<paragraph id="8B8136B0FFBC84458CABFBC7D947FAB1" blockId="33.[151,1437,534,1984]" pageId="33" pageNumber="34">
<emphasis id="B94AEAA2FFBC84458CABFBC7DA84FB9E" bold="true" box="[199,442,1057,1084]" pageId="33" pageNumber="34">
Descriptions.
<emphasis id="B94AEAA2FFBC84458D02FBC4DA84FB9E" bold="true" box="[366,442,1058,1084]" italics="true" pageId="33" pageNumber="34">Larva.</emphasis>
</emphasis>
CUTICULAR COLORATION (
<figureCitation id="13052A35FFBC84458F59FBC7D884FB99" box="[821,954,1057,1083]" captionStart="FIGURES 66–69" captionStartId="34.[151,264,1320,1345]" captionTargetBox="[151,1436,166,1301]" captionTargetId="figure@34.[151,1436,166,1301]" captionTargetPageId="34" captionText="FIGURES 66–69. Thraulodes telegraphicus. 66–67, last instar larvae with intensive cuticular coloration and poorly visible hypodermal coloration; 68, younger larva with light cuticle and well visible blackish hypodermal markings on abdomen; 69, claw. Abbreviations: artic.p., articulatory portion; rigid p., rigid portion of claw." figureDoi="http://doi.org/10.5281/zenodo.3736917" httpUri="https://zenodo.org/record/3736917/files/figure.png" pageId="33" pageNumber="34">Figs 66–67</figureCitation>
,
<figureCitation id="13052A35FFBC84458FA6FBC7DF2BFB99" box="[970,1045,1057,1083]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="33" pageNumber="34">70–82</figureCitation>
). Dorsal side of head, thorax and abdomen with highly variable coloration. Head either medially brown and laterally light (
<figureCitation id="13052A35FFBC84458811FBA3DFEEFBC2" box="[1149,1232,1093,1120]" captionStart="FIGURES 66–69" captionStartId="34.[151,264,1320,1345]" captionTargetBox="[151,1436,166,1301]" captionTargetId="figure@34.[151,1436,166,1301]" captionTargetPageId="34" captionText="FIGURES 66–69. Thraulodes telegraphicus. 66–67, last instar larvae with intensive cuticular coloration and poorly visible hypodermal coloration; 68, younger larva with light cuticle and well visible blackish hypodermal markings on abdomen; 69, claw. Abbreviations: artic.p., articulatory portion; rigid p., rigid portion of claw." figureDoi="http://doi.org/10.5281/zenodo.3736917" httpUri="https://zenodo.org/record/3736917/files/figure.png" pageId="33" pageNumber="34">Fig. 66</figureCitation>
), or mostly brown (
<figureCitation id="13052A35FFBC84458CCCFB8FDBCAFB26" box="[160,244,1129,1156]" captionStart="FIGURES 66–69" captionStartId="34.[151,264,1320,1345]" captionTargetBox="[151,1436,166,1301]" captionTargetId="figure@34.[151,1436,166,1301]" captionTargetPageId="34" captionText="FIGURES 66–69. Thraulodes telegraphicus. 66–67, last instar larvae with intensive cuticular coloration and poorly visible hypodermal coloration; 68, younger larva with light cuticle and well visible blackish hypodermal markings on abdomen; 69, claw. Abbreviations: artic.p., articulatory portion; rigid p., rigid portion of claw." figureDoi="http://doi.org/10.5281/zenodo.3736917" httpUri="https://zenodo.org/record/3736917/files/figure.png" pageId="33" pageNumber="34">Fig. 67</figureCitation>
); pronotum either mostly light with contrasting brown maculae on these or that margins (
<figureCitation id="13052A35FFBC8445888EFB8FDE07FB21" box="[1250,1337,1129,1155]" captionStart="FIGURES 66–69" captionStartId="34.[151,264,1320,1345]" captionTargetBox="[151,1436,166,1301]" captionTargetId="figure@34.[151,1436,166,1301]" captionTargetPageId="34" captionText="FIGURES 66–69. Thraulodes telegraphicus. 66–67, last instar larvae with intensive cuticular coloration and poorly visible hypodermal coloration; 68, younger larva with light cuticle and well visible blackish hypodermal markings on abdomen; 69, claw. Abbreviations: artic.p., articulatory portion; rigid p., rigid portion of claw." figureDoi="http://doi.org/10.5281/zenodo.3736917" httpUri="https://zenodo.org/record/3736917/files/figure.png" pageId="33" pageNumber="34">Figs 66</figureCitation>
,
<figureCitation id="13052A35FFBC8445892AFB8FDEB1FB21" box="[1350,1423,1129,1155]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="33" pageNumber="34">70–73</figureCitation>
), or mostly brown with light lateral margins (
<figureCitation id="13052A35FFBC84458E10FB6BD9ECFB05" box="[636,722,1165,1191]" captionStart="FIGURES 66–69" captionStartId="34.[151,264,1320,1345]" captionTargetBox="[151,1436,166,1301]" captionTargetId="figure@34.[151,1436,166,1301]" captionTargetPageId="34" captionText="FIGURES 66–69. Thraulodes telegraphicus. 66–67, last instar larvae with intensive cuticular coloration and poorly visible hypodermal coloration; 68, younger larva with light cuticle and well visible blackish hypodermal markings on abdomen; 69, claw. Abbreviations: artic.p., articulatory portion; rigid p., rigid portion of claw." figureDoi="http://doi.org/10.5281/zenodo.3736917" httpUri="https://zenodo.org/record/3736917/files/figure.png" pageId="33" pageNumber="34">Figs 67</figureCitation>
,
<figureCitation id="13052A35FFBC84458EB2FB6BD817FB0A" box="[734,809,1165,1192]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="33" pageNumber="34">74–75</figureCitation>
); mesonotum mostly brown, either with several separate blanks, or crossed by wide blanks (
<figureCitation id="13052A35FFBC84458E71FB57D94FFB6E" box="[541,625,1201,1228]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="33" pageNumber="34">Fig. 77</figureCitation>
); abdominal terga either brown with blanks on lateral parts, or terga I–III and VI–VII brown and terga IV–V and VIII–X contrastingly light (
<figureCitation id="13052A35FFBC84458F12FB33D8EAFB4D" box="[894,980,1237,1263]" captionStart="FIGURES 66–69" captionStartId="34.[151,264,1320,1345]" captionTargetBox="[151,1436,166,1301]" captionTargetId="figure@34.[151,1436,166,1301]" captionTargetPageId="34" captionText="FIGURES 66–69. Thraulodes telegraphicus. 66–67, last instar larvae with intensive cuticular coloration and poorly visible hypodermal coloration; 68, younger larva with light cuticle and well visible blackish hypodermal markings on abdomen; 69, claw. Abbreviations: artic.p., articulatory portion; rigid p., rigid portion of claw." figureDoi="http://doi.org/10.5281/zenodo.3736917" httpUri="https://zenodo.org/record/3736917/files/figure.png" pageId="33" pageNumber="34">Figs 66</figureCitation>
,
<figureCitation id="13052A35FFBC84458FB3FB33D8C0FB4D" box="[991,1022,1237,1263]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="33" pageNumber="34">81</figureCitation>
). Femora with brown and light areas, tibiae and tarsi unicolor light (
<figureCitation id="13052A35FFBC84458D86FB1FD953FAB1" box="[490,621,1273,1299]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="33" pageNumber="34">Figs 77–80</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFBC84458CABFAFBDE20FADD" blockId="33.[151,1437,534,1984]" pageId="33" pageNumber="34">
HYPODERMAL COLORATION (
<figureCitation id="13052A35FFBC84458E3BFAFBD990FA9A" box="[599,686,1309,1336]" captionStart="FIGURES 66–69" captionStartId="34.[151,264,1320,1345]" captionTargetBox="[151,1436,166,1301]" captionTargetId="figure@34.[151,1436,166,1301]" captionTargetPageId="34" captionText="FIGURES 66–69. Thraulodes telegraphicus. 66–67, last instar larvae with intensive cuticular coloration and poorly visible hypodermal coloration; 68, younger larva with light cuticle and well visible blackish hypodermal markings on abdomen; 69, claw. Abbreviations: artic.p., articulatory portion; rigid p., rigid portion of claw." figureDoi="http://doi.org/10.5281/zenodo.3736917" httpUri="https://zenodo.org/record/3736917/files/figure.png" pageId="33" pageNumber="34">Fig. 68</figureCitation>
). Abdominal terga, besides cuticular coloration (see above) often with paired transverse or round blackish spots corresponding to midway gray spots of imago. Tergalii either light grayish, or light brownish-reddish, or colorless; tracheae often black, sometimes light gray or colorless.
</paragraph>
<paragraph id="8B8136B0FFBC84458CABFA6FD82BF9B2" blockId="33.[151,1437,534,1984]" pageId="33" pageNumber="34">
SHAPE AND SETATION. Clypeus widened distally; labrum 1.2–1.5 times wider than clypeus (
<figureCitation id="13052A35FFBC84458967FA6FDE5FFA06" box="[1291,1377,1417,1444]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="33" pageNumber="34">Fig. 76</figureCitation>
). Labrum widest at 1/3 length from base; initial fore margin (often turned ventrally) without median emargination, slightly convex, with all 5 denticles wide; anterior transverse setal row regular, as wide as 3 or 5 denticles (
<figureCitation id="13052A35FFBC84458950FA37DEB1FA49" box="[1340,1423,1489,1515]" captionStart="FIGURES 86–93" captionStartId="37.[151,264,188,213]" captionTargetId="figure@36.[151,1436,342,2018]" captionTargetPageId="36" captionText="FIGURES 86–93. Thraulodes telegraphicus. 86, median longitudinal section of labrum in usual condition; 87, the same, in spread condition; 88, labrum in spread condition in dorsal view (setal rows shown as dotted lines); 89, transverse section of hind tibia; 90–93, tergalii I, III, VI and VII. Abbreviations: anter.r, anterior transverse row of setae; dent., five denticles separated by six sensilla; poster.r, posterior transverse row of setae; i.f, inner field of stout pointed setae; i-a.h, inner-anterior recurved hairs; i-a.r, inner-anterior row of stout setae; o-a.r, outer-anterior row of stout setae; o.h, outer hairs; o-p.r, outer-posterior row of stout setae; p.r, row of posterior pointed pectinate setae." figureDoi="http://doi.org/10.5281/zenodo.3736921" httpUri="https://zenodo.org/record/3736921/files/figure.png" pageId="33" pageNumber="34">Fig. 88</figureCitation>
). Maxilla with 14–17 pectinate setae in apical-ventral row.
</paragraph>
<paragraph id="8B8136B0FFBC84458CABF9FCDF0CF9F5" blockId="33.[151,1437,534,1984]" pageId="33" pageNumber="34">
Femora: Stout setae on anterior surface parallel-sided or widened distally, apically blunt (
<figureCitation id="13052A35FFBC844588CFF9FFDFC8F991" box="[1187,1270,1561,1587]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="33" pageNumber="34">Fig. 83</figureCitation>
). Irregular row of hairs near inner margin sparse on fore femur, dense on middle and hind femora.
</paragraph>
<paragraph id="8B8136B0FFBC84458CABF984D94AF94A" blockId="33.[151,1437,534,1984]" pageId="33" pageNumber="34">
Fore tibia (
<figureCitation id="13052A35FFBC84458D2FF987DAA6F9D9" box="[323,408,1633,1659]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="33" pageNumber="34">Fig. 84</figureCitation>
): outer hairs form two irregular rows; inner-anterior row of recurved hairs absent; inner-anterior row of stout setae represented by few (2–4) blunt stout setae near tibia base; inner field of stout pointed setae dense (i.e. setae longer than distances between them), consists of pointed bipectinate and smooth setae, situated irregularly (about 3 setae in cross section).
</paragraph>
<paragraph id="8B8136B0FFBC84458CABF917DA09F839" blockId="33.[151,1437,534,1984]" pageId="33" pageNumber="34">
Hind tibia (
<figureCitation id="13052A35FFBC84458D22F917DA98F8AE" box="[334,422,1777,1804]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="33" pageNumber="34">Fig. 85</figureCitation>
): outer-anterior row of stout setae consists of two clearly different
<typeStatus id="54858812FFBC844588D9F914DFCEF8AE" box="[1205,1264,1778,1804]" pageId="33" pageNumber="34">types</typeStatus>
of stout setae: numerous short rounded ones and several long spoon-like ones; outer-posterior row of stout setae consists of stout blunt setae of various lengths; hairs located between these rows, numerous and form more than one row (besides row of hairs posteriad of outer-posterior row of stout setae); stout setae of inner-anterior row short, widened distally and truncated.
</paragraph>
<paragraph id="8B8136B0FFBC84458CABF843DE15F81D" blockId="33.[151,1437,534,1984]" box="[199,1323,1957,1984]" pageId="33" pageNumber="34">Claws with 6–10 denticles on rigid portion, with or without minute denticles on articulatory portion.</paragraph>
<caption id="DF416638FFBF84468CFBFACED881FA2B" ID-DOI="http://doi.org/10.5281/zenodo.3736917" ID-Zenodo-Dep="3736917" httpUri="https://zenodo.org/record/3736917/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,264,1320,1345]" targetBox="[151,1436,166,1301]" targetPageId="34">
<paragraph id="8B8136B0FFBF84468CFBFACED881FA2B" blockId="34.[151,1437,1320,1417]" pageId="34" pageNumber="35">
<emphasis id="B94AEAA2FFBF84468CFBFACEDA69FAE3" bold="true" box="[151,343,1320,1345]" pageId="34" pageNumber="35">FIGURES 66–69.</emphasis>
<taxonomicName id="4C3E4D33FFBF84468D33FACFD95BFAE3" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[351,613,1321,1345]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFBF84468D33FACFD95BFAE3" box="[351,613,1321,1345]" italics="true" pageId="34" pageNumber="35">Thraulodes telegraphicus</emphasis>
</taxonomicName>
. 66–67, last instar larvae with intensive cuticular coloration and poorly visible hypodermal coloration; 68, younger larva with light cuticle and well visible blackish hypodermal markings on abdomen; 69, claw. Abbreviations: artic.p., articulatory portion; rigid p., rigid portion of claw.
</paragraph>
</caption>
<paragraph id="8B8136B0FFBF84468CABFA3DD950F9FD" blockId="34.[151,1437,1499,1988]" pageId="34" pageNumber="35">
Tergalii (
<figureCitation id="13052A35FFBF84468D40FA3DDA91FA57" box="[300,431,1499,1525]" captionStart="FIGURES 86–93" captionStartId="37.[151,264,188,213]" captionTargetId="figure@36.[151,1436,342,2018]" captionTargetPageId="36" captionText="FIGURES 86–93. Thraulodes telegraphicus. 86, median longitudinal section of labrum in usual condition; 87, the same, in spread condition; 88, labrum in spread condition in dorsal view (setal rows shown as dotted lines); 89, transverse section of hind tibia; 90–93, tergalii I, III, VI and VII. Abbreviations: anter.r, anterior transverse row of setae; dent., five denticles separated by six sensilla; poster.r, posterior transverse row of setae; i.f, inner field of stout pointed setae; i-a.h, inner-anterior recurved hairs; i-a.r, inner-anterior row of stout setae; o-a.r, outer-anterior row of stout setae; o.h, outer hairs; o-p.r, outer-posterior row of stout setae; p.r, row of posterior pointed pectinate setae." figureDoi="http://doi.org/10.5281/zenodo.3736921" httpUri="https://zenodo.org/record/3736921/files/figure.png" pageId="34" pageNumber="35">Figs 90–93</figureCitation>
): wide; on both lamellae main trachea with many branches directed toward costal and anal margins; dorsal lamella with costal margin most convex in proximal part and anal margin most convex in distal part, gradually narrowed toward apex, with slender apical filament; ventral lamella widest near base, gradually narrowed toward apex, with slender apical filament.
</paragraph>
<paragraph id="8B8136B0FFBF84468CABF98FDBEBF968" blockId="34.[151,1437,1499,1988]" pageId="34" pageNumber="35">
Male genitalia in last larval instar (
<figureCitation id="13052A35FFBF84468E23F98FD9D1F926" box="[591,751,1641,1668]" captionStart="FIGURES 106–111" captionStartId="40.[151,264,1834,1859]" captionTargetBox="[159,1390,175,1807]" captionTargetId="figure@40.[151,1397,166,1814]" captionTargetPageId="40" captionText="FIGURES 106–111. Thraulodes telegraphicus. 106, protogonostyli of last instar larva (hypoderm of gonostyli and styliger and hidden cuticle of protopenis shown by interrupted lines); 107, enlarged cuticle of protopenis and hypoderm of developing penis inside; 108–109, the same, at later stage of development; 110–111, genital of male imago (110, styliger ventrally; 111, penis dorsally at left and ventrally at right). Abbreviation: bl, basal lobe; dp, dorsal projection of styliger; gd, gonoduct; m.gs, gonostylar muscle; m.med, median penial muscle; m.s, styligeral muscle; m-v.r, medio-ventral ridge (recurved flap); sp, seminal pump; tp, spear-like roll (telopenis)." figureDoi="http://doi.org/10.5281/zenodo.3736927" httpUri="https://zenodo.org/record/3736927/files/figure.png" pageId="34" pageNumber="35">Figs 106–109</figureCitation>
): protogonostyli separated one from another by deep emargination. Each protopenes lobe with gonopore-bearing process located near median margin; gonopore opened caudally.
</paragraph>
<paragraph id="8B8136B0FFBF84468CABF934DE54F861" blockId="34.[151,1437,1499,1988]" pageId="34" pageNumber="35">
<emphasis id="B94AEAA2FFBF84468CABF934DA02F94F" bold="true" box="[199,316,1746,1773]" italics="true" pageId="34" pageNumber="35">Subimago</emphasis>
(
<figureCitation id="13052A35FFBF84468D20F935DAF1F94F" box="[332,463,1746,1773]" captionStart="FIGURES 94–100" captionStartId="38.[151,264,1799,1824]" captionTargetBox="[151,1436,222,1780]" captionTargetId="figure@38.[151,1436,222,1780]" captionTargetPageId="38" captionText="FIGURES 94–100. Thraulodes telegraphicus. 94–95, right half of subimaginal exuviae of mesonotum of two different specimens; 96, apex of fore wing; 97–98, male imago in lateral and ventral view; 99, abdominal terga and sterna of male imago (black dots are algae under cuticle); 100, female imago. Abbreviations: acz.ms, achromozone of medioscutum; acz.sms, achromozone of submedioscutum; cz.ms, chromozone of medioscutum; cz.sms, chromozone of submedioscutum; mp.s, medioparapsidal suture; ms.cz.s, medioscutal chromozonal suture; sms.cz.s, submedioscutal chromozonal suture." figureDoi="http://doi.org/10.5281/zenodo.3736923" httpUri="https://zenodo.org/record/3736923/files/figure.png" pageId="34" pageNumber="35">Figs 94–95</figureCitation>
,
<figureCitation id="13052A35FFBF84468DB0F934D937F94F" box="[476,521,1746,1773]" captionStart="FIGURES 112–120" captionStartId="41.[151,264,1889,1914]" captionTargetBox="[151,1423,170,1862]" captionTargetId="figure@41.[151,1436,166,1869]" captionTargetPageId="41" captionText="FIGURES 112–120. Thraulodes telegraphicus. 112, genitalia of male imago in lateral view; 113, apex of penis with spear-like roll in dorsal view; 114, genitalia of the same specimen in ventral view; 115, their subimaginal exuviae at the same magnification; 116, hind wing; 117–120, penis of four different specimens. Abbreviations: dp, dorsal projection of styliger; tp, spear-like roll (telopenis)." figureDoi="http://doi.org/10.5281/zenodo.3736929" httpUri="https://zenodo.org/record/3736929/files/figure.png" pageId="34" pageNumber="35">115</figureCitation>
). CUTICULAR COLORATION. Cuticle mostly colorless. Flagellum of antenna brown. Pronotum colorless. Mesonotum with colorless and brown areas; antelateroparapsidal suture contrastingly colored with dark brown, lateroparapsidal suture and some other areas lighter brown; chromozone of medioscutum light brown, chromozone of submedioscutum contrastingly lighter (
<figureCitation id="13052A35FFBF84468FF8F8DBDF26F8FA" box="[916,1048,1853,1880]" captionStart="FIGURES 94–100" captionStartId="38.[151,264,1799,1824]" captionTargetBox="[151,1436,222,1780]" captionTargetId="figure@38.[151,1436,222,1780]" captionTargetPageId="38" captionText="FIGURES 94–100. Thraulodes telegraphicus. 94–95, right half of subimaginal exuviae of mesonotum of two different specimens; 96, apex of fore wing; 97–98, male imago in lateral and ventral view; 99, abdominal terga and sterna of male imago (black dots are algae under cuticle); 100, female imago. Abbreviations: acz.ms, achromozone of medioscutum; acz.sms, achromozone of submedioscutum; cz.ms, chromozone of medioscutum; cz.sms, chromozone of submedioscutum; mp.s, medioparapsidal suture; ms.cz.s, medioscutal chromozonal suture; sms.cz.s, submedioscutal chromozonal suture." figureDoi="http://doi.org/10.5281/zenodo.3736923" httpUri="https://zenodo.org/record/3736923/files/figure.png" pageId="34" pageNumber="35">Figs 94–95</figureCitation>
), sometimes colorless. Fore femur mostly colorless, more or less colored by brown on outer margin and on apex; middle and hind femora colorless; on each leg tibia and tarsus light brown. Cuticle of wings colorless, microtrichia light brownish.Abdominal terga mostly colorless, diffusively shed by brown medially; sterna colorless; gonostyli colorless; caudalii light brownish.
</paragraph>
<caption id="DF416638FFB984408CFBFF5ADFF3FEE2" ID-DOI="http://doi.org/10.5281/zenodo.3736919" ID-Zenodo-Dep="3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="36" pageNumber="37" targetBox="[151,1436,181,1986]" targetPageId="35">
<paragraph id="8B8136B0FFB984408CFBFF5ADFF3FEE2" blockId="36.[151,1437,188,321]" pageId="36" pageNumber="37">
<emphasis id="B94AEAA2FFB984408CFBFF5ADA6BFF77" bold="true" box="[151,341,188,213]" pageId="36" pageNumber="37">FIGURES 70–85.</emphasis>
<taxonomicName id="4C3E4D33FFB984408D37FF5AD961FF76" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[347,607,188,212]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFB984408D37FF5AD961FF76" box="[347,607,188,212]" italics="true" pageId="36" pageNumber="37">Thraulodes telegraphicus</emphasis>
</taxonomicName>
, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae.
</paragraph>
</caption>
<caption id="DF416638FFB884418CFBFF5AD9A8FE2A" ID-DOI="http://doi.org/10.5281/zenodo.3736921" ID-Zenodo-Dep="3736921" httpUri="https://zenodo.org/record/3736921/files/figure.png" pageId="37" pageNumber="38" targetBox="[158,1434,354,2012]" targetPageId="36">
<paragraph id="8B8136B0FFB884418CFBFF5AD9A8FE2A" blockId="37.[151,1437,188,393]" pageId="37" pageNumber="38">
<emphasis id="B94AEAA2FFB884418CFBFF5ADA68FF77" bold="true" box="[151,342,188,213]" pageId="37" pageNumber="38">FIGURES 86–93.</emphasis>
<taxonomicName id="4C3E4D33FFB884418D32FF5AD95AFF76" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[350,612,188,212]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFB884418D32FF5AD95AFF76" box="[350,612,188,212]" italics="true" pageId="37" pageNumber="38">Thraulodes telegraphicus</emphasis>
</taxonomicName>
. 86, median longitudinal section of labrum in usual condition; 87, the same, in spread condition; 88, labrum in spread condition in dorsal view (setal rows shown as dotted lines); 89, transverse section of hind tibia; 90–93, tergalii I, III, VI and VII. Abbreviations: anter.r, anterior transverse row of setae; dent., five denticles separated by six sensilla; poster.r, posterior transverse row of setae; i.f, inner field of stout pointed setae; i-a.h, inner-anterior recurved hairs; i-a.r, inner-anterior row of stout setae; o-a.r, outer-anterior row of stout setae; o.h, outer hairs; o-p.r, outer-posterior row of stout setae; p.r, row of posterior pointed pectinate setae.
</paragraph>
</caption>
<paragraph id="8B8136B0FFB884418CABFE3DDA1CFD9F" blockId="37.[151,1437,474,2013]" pageId="37" pageNumber="38">
TEXTURE. On tarsi of all legs, 1st tarsomere with microtrichia (as tibia), 2nd–5th tarsomeres coved by blunt microlepides; pointed microlepides present near apical margins of 2nd–4th tarsomeres of middle and hind legs (as in
<figureCitation id="13052A35FFB884418CD8FDC5DA2BFD9F" box="[180,277,547,573]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="37" pageNumber="38">Fig. 473</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFB884418CABFDA0DA05FD24" blockId="37.[151,1437,474,2013]" pageId="37" pageNumber="38">
<emphasis id="B94AEAA2FFB884418CABFDA0DA6AFDC3" bold="true" box="[199,340,582,609]" italics="true" pageId="37" pageNumber="38">Male imago</emphasis>
(
<figureCitation id="13052A35FFB884418D09FDA1DAD7FDC3" box="[357,489,583,609]" captionStart="FIGURES 94–100" captionStartId="38.[151,264,1799,1824]" captionTargetBox="[151,1436,222,1780]" captionTargetId="figure@38.[151,1436,222,1780]" captionTargetPageId="38" captionText="FIGURES 94–100. Thraulodes telegraphicus. 94–95, right half of subimaginal exuviae of mesonotum of two different specimens; 96, apex of fore wing; 97–98, male imago in lateral and ventral view; 99, abdominal terga and sterna of male imago (black dots are algae under cuticle); 100, female imago. Abbreviations: acz.ms, achromozone of medioscutum; acz.sms, achromozone of submedioscutum; cz.ms, chromozone of medioscutum; cz.sms, chromozone of submedioscutum; mp.s, medioparapsidal suture; ms.cz.s, medioscutal chromozonal suture; sms.cz.s, submedioscutal chromozonal suture." figureDoi="http://doi.org/10.5281/zenodo.3736923" httpUri="https://zenodo.org/record/3736923/files/figure.png" pageId="37" pageNumber="38">Figs 96–99</figureCitation>
,
<figureCitation id="13052A35FFB884418D9AFDA0D965FDC3" box="[502,603,582,609]" captionStart="FIGURES 101–105" captionStartId="39.[151,264,1784,1809]" captionTargetBox="[151,1436,166,1764]" captionTargetId="figure@39.[151,1436,166,1764]" captionTargetPageId="39" captionText="FIGURES 101–105. Thraulodes telegraphicus. 101–102, genitalia of the same specimen; 103, penis of another specimen; 104– 105, part of penis with telopenis forming spear-like roll.Abbreviations: bl, basal lobe; dp, dorsal projection of styliger; gd, gonoduct; gs1, gs2, gs3, gs4, segments of gonostylus; m.gs, gonostylar muscle; m.lat, lateral penial muscle; m.med, median penial muscle; m.s, styligeral muscle; m-v.r, medio-ventral ridge (recurved flap); sp, seminal pump; tp, spear-like roll (telopenis)." figureDoi="http://doi.org/10.5281/zenodo.3736925" httpUri="https://zenodo.org/record/3736925/files/figure.png" pageId="37" pageNumber="38">101–105</figureCitation>
,
<figureCitation id="13052A35FFB884418E05FDA1D9F1FDC3" box="[617,719,583,609]" captionStart-0="FIGURES 106–111" captionStart-1="FIGURES 112–120" captionStartId-0="40.[151,264,1834,1859]" captionStartId-1="41.[151,264,1889,1914]" captionTargetBox-0="[159,1390,175,1807]" captionTargetBox-1="[151,1423,170,1862]" captionTargetId-0="figure@40.[151,1397,166,1814]" captionTargetId-1="figure@41.[151,1436,166,1869]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURES 106–111. Thraulodes telegraphicus. 106, protogonostyli of last instar larva (hypoderm of gonostyli and styliger and hidden cuticle of protopenis shown by interrupted lines); 107, enlarged cuticle of protopenis and hypoderm of developing penis inside; 108–109, the same, at later stage of development; 110–111, genital of male imago (110, styliger ventrally; 111, penis dorsally at left and ventrally at right). Abbreviation: bl, basal lobe; dp, dorsal projection of styliger; gd, gonoduct; m.gs, gonostylar muscle; m.med, median penial muscle; m.s, styligeral muscle; m-v.r, medio-ventral ridge (recurved flap); sp, seminal pump; tp, spear-like roll (telopenis)." captionText-1="FIGURES 112–120. Thraulodes telegraphicus. 112, genitalia of male imago in lateral view; 113, apex of penis with spear-like roll in dorsal view; 114, genitalia of the same specimen in ventral view; 115, their subimaginal exuviae at the same magnification; 116, hind wing; 117–120, penis of four different specimens. Abbreviations: dp, dorsal projection of styliger; tp, spear-like roll (telopenis)." figureDoi-0="http://doi.org/10.5281/zenodo.3736927" figureDoi-1="http://doi.org/10.5281/zenodo.3736929" httpUri-0="https://zenodo.org/record/3736927/files/figure.png" httpUri-1="https://zenodo.org/record/3736929/files/figure.png" pageId="37" pageNumber="38">110–120</figureCitation>
). Described by
<bibRefCitation id="EFAF4B41FFB884418FE8FDA1DFF8FDC3" author="Needham, J. G. &amp; Murphy, H. E." box="[900,1222,583,609]" pageId="37" pageNumber="38" pagination="1 - 79" refId="ref71682" refString="Needham, J. G. &amp; Murphy, H. E. (1924) Neotropical mayflies. Bulletin of the Lloyd Library, Entomological Series 4, 24, 1 - 79." type="journal article" year="1924">Needham &amp; Murphy (1924)</bibRefCitation>
and
<bibRefCitation id="EFAF4B41FFB88441896DFDA1DA08FD24" author="Traver, J. R. &amp; Edmunds, G. F. Jr." pageId="37" pageNumber="38" pagination="349 - 395" refId="ref72596" refString="Traver, J. R. &amp; Edmunds, G. F. Jr. (1967) A revision of the genus Thraulodes (Ephemeroptera: Leptophlebiidae). Miscellaneous Publications of the Entomological Society of America, 5 (8), 349 - 395." type="journal article" year="1967">Traver &amp; Edmunds (1967)</bibRefCitation>
.
</paragraph>
<paragraph id="8B8136B0FFB884418CABFD69DF24FD6F" blockId="37.[151,1437,474,2013]" pageId="37" pageNumber="38">Head brown. Antenna with scapus ocher with brown macula; pedicellus proximally ocher, distally brown; flagellum brownish with ocher base. Dorsal eyes contiguous medially, dull orange.</paragraph>
<paragraph id="8B8136B0FFB884418CABFD31DA06FCFF" blockId="37.[151,1437,474,2013]" pageId="37" pageNumber="38">
Prothorax ocher with pair of dark brown oblique stripes on pleura. Mesonotum and metanotum ocher or/and brown. Lateral sides of pterothorax with brown and ocher areas. Ventral side of pterothorax with mesothoracic epi- sterna, basisternum, furcasternal protuberances and furcasternal median impression either brown with ocher (
<figureCitation id="13052A35FFB884418903FCF9DB88FCFF" captionStart="FIGURES 94–100" captionStartId="38.[151,264,1799,1824]" captionTargetBox="[151,1436,222,1780]" captionTargetId="figure@38.[151,1436,222,1780]" captionTargetPageId="38" captionText="FIGURES 94–100. Thraulodes telegraphicus. 94–95, right half of subimaginal exuviae of mesonotum of two different specimens; 96, apex of fore wing; 97–98, male imago in lateral and ventral view; 99, abdominal terga and sterna of male imago (black dots are algae under cuticle); 100, female imago. Abbreviations: acz.ms, achromozone of medioscutum; acz.sms, achromozone of submedioscutum; cz.ms, chromozone of medioscutum; cz.sms, chromozone of submedioscutum; mp.s, medioparapsidal suture; ms.cz.s, medioscutal chromozonal suture; sms.cz.s, submedioscutal chromozonal suture." figureDoi="http://doi.org/10.5281/zenodo.3736923" httpUri="https://zenodo.org/record/3736923/files/figure.png" pageId="37" pageNumber="38">Fig. 98</figureCitation>
), or brown.
</paragraph>
<paragraph id="8B8136B0FFB884418CABFC81DA93FB03" blockId="37.[151,1437,474,2013]" pageId="37" pageNumber="38">Cuticle of legs mostly colorless; on fore leg apex of femur and base of tibia with cuticle brown. Legs with following hypodermal pigmentation on whitish-ocher background: Fore femur brown with diffusive spot(s) or interrupted band on proximal half and with multicolored apical band occupying distal 0.4 of femur; apical band brownish-yellowish, proximally bordered by brown, on inner and outer sides with brown longitudinal stripes. Middle and hind femora with multicolored pre-apical band within apical 0.4 of femur; pre-apical band brownish-yellowish, proximally bordered by brown, on inner and outer sides with brown longitudinal stripes; apical 1.0 of femur nonpigmented. Tibia of each leg apically dark brown. In selected individual, relation of fore femur to fore wing length 75:325; proportions femur/tibia/tarsomeres on fore leg 75:107:2:26:20:18:10; on middle leg 70:80:4:4:3:3:9; on hind leg 85:90:4:4:3:3:9.</paragraph>
<paragraph id="8B8136B0FFB884418CABFB4DD8CEFADB" blockId="37.[151,1437,474,2013]" pageId="37" pageNumber="38">
Fore wing with longitudinal and cross veins ocher; costal braces and bases of Sc and R bordered by brown. Costal cross veins proximad of bulla either absent, of few, colorless and very thin, so that visible only under high magnification (as in
<figureCitation id="13052A35FFB884418DE6FB15DAD1FAAF" box="[394,495,1267,1293]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="37" pageNumber="38">Fig. 142</figureCitation>
); occasionally proximal part of costal field with several well-visible cross veins. Pterostigmatic cross veins moderately dense, oblique, arched, non-branched (
<figureCitation id="13052A35FFB884418F98FAF1DF7AFA90" box="[1012,1092,1303,1330]" captionStart="FIGURES 94–100" captionStartId="38.[151,264,1799,1824]" captionTargetBox="[151,1436,222,1780]" captionTargetId="figure@38.[151,1436,222,1780]" captionTargetPageId="38" captionText="FIGURES 94–100. Thraulodes telegraphicus. 94–95, right half of subimaginal exuviae of mesonotum of two different specimens; 96, apex of fore wing; 97–98, male imago in lateral and ventral view; 99, abdominal terga and sterna of male imago (black dots are algae under cuticle); 100, female imago. Abbreviations: acz.ms, achromozone of medioscutum; acz.sms, achromozone of submedioscutum; cz.ms, chromozone of medioscutum; cz.sms, chromozone of submedioscutum; mp.s, medioparapsidal suture; ms.cz.s, medioscutal chromozonal suture; sms.cz.s, submedioscutal chromozonal suture." figureDoi="http://doi.org/10.5281/zenodo.3736923" httpUri="https://zenodo.org/record/3736923/files/figure.png" pageId="37" pageNumber="38">Fig. 96</figureCitation>
) or occasionally anastomozed. Hind wing with veins colorless, with brown macula on costal brace. Proximal coloration of fore and hind wings appears during larval development and present both in subimago and imago.
</paragraph>
<paragraph id="8B8136B0FFB884418CABFA65DA8FF93B" blockId="37.[151,1437,474,2013]" pageId="37" pageNumber="38">Abdominal hypodermal coloration: Tergum I ocher-brown. Terga II–VI mostly light ocher, translucent, with or without brown band on posterior margin, medially more or less tinged with darker ocher; terga III–VI or III–VI with pair of smaller or larger gray midway spots; pleura with gray stigmatic dots. Terga VII–X with orange or ocherbrownish coloration: tergum VII mostly colored with latero-posterior areas whitish, tergum VIII colored anteriorlymedially and whitish posteriorly-laterally, terga XI–X mostly colored; lateral margins of terga VIII and IX with blackish-brown longitudinal stripes. Abdominal sterna I–IX light ocher; posterolateral angles of sterna VII and VIII with pair of blackish-brown spots. Caudalii with segments alternated as following: black; white; white proximally and black distally; white.</paragraph>
<paragraph id="8B8136B0FFB884418CABF945D893F837" blockId="37.[151,1437,474,2013]" pageId="37" pageNumber="38">
Genitalia (
<figureCitation id="13052A35FFB884418D51F945DAE3F91F" box="[317,477,1699,1725]" captionStart-0="FIGURES 101–105" captionStart-1="FIGURES 106–111" captionStart-2="FIGURES 112–120" captionStartId-0="39.[151,264,1784,1809]" captionStartId-1="40.[151,264,1834,1859]" captionStartId-2="41.[151,264,1889,1914]" captionTargetBox-0="[151,1436,166,1764]" captionTargetBox-1="[159,1390,175,1807]" captionTargetBox-2="[151,1423,170,1862]" captionTargetId-0="figure@39.[151,1436,166,1764]" captionTargetId-1="figure@40.[151,1397,166,1814]" captionTargetId-2="figure@41.[151,1436,166,1869]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="41" captionText-0="FIGURES 101–105. Thraulodes telegraphicus. 101–102, genitalia of the same specimen; 103, penis of another specimen; 104– 105, part of penis with telopenis forming spear-like roll.Abbreviations: bl, basal lobe; dp, dorsal projection of styliger; gd, gonoduct; gs1, gs2, gs3, gs4, segments of gonostylus; m.gs, gonostylar muscle; m.lat, lateral penial muscle; m.med, median penial muscle; m.s, styligeral muscle; m-v.r, medio-ventral ridge (recurved flap); sp, seminal pump; tp, spear-like roll (telopenis)." captionText-1="FIGURES 106–111. Thraulodes telegraphicus. 106, protogonostyli of last instar larva (hypoderm of gonostyli and styliger and hidden cuticle of protopenis shown by interrupted lines); 107, enlarged cuticle of protopenis and hypoderm of developing penis inside; 108–109, the same, at later stage of development; 110–111, genital of male imago (110, styliger ventrally; 111, penis dorsally at left and ventrally at right). Abbreviation: bl, basal lobe; dp, dorsal projection of styliger; gd, gonoduct; m.gs, gonostylar muscle; m.med, median penial muscle; m.s, styligeral muscle; m-v.r, medio-ventral ridge (recurved flap); sp, seminal pump; tp, spear-like roll (telopenis)." captionText-2="FIGURES 112–120. Thraulodes telegraphicus. 112, genitalia of male imago in lateral view; 113, apex of penis with spear-like roll in dorsal view; 114, genitalia of the same specimen in ventral view; 115, their subimaginal exuviae at the same magnification; 116, hind wing; 117–120, penis of four different specimens. Abbreviations: dp, dorsal projection of styliger; tp, spear-like roll (telopenis)." figureDoi-0="http://doi.org/10.5281/zenodo.3736925" figureDoi-1="http://doi.org/10.5281/zenodo.3736927" figureDoi-2="http://doi.org/10.5281/zenodo.3736929" httpUri-0="https://zenodo.org/record/3736925/files/figure.png" httpUri-1="https://zenodo.org/record/3736927/files/figure.png" httpUri-2="https://zenodo.org/record/3736929/files/figure.png" pageId="37" pageNumber="38">Figs 101–120</figureCitation>
): Styliger ocher. Gonostyli either entirely ocher, or with long segment distally brownish (
<figureCitation id="13052A35FFB884418CABF921DA16F943" box="[199,296,1735,1761]" captionStart="FIGURES 101–105" captionStartId="39.[151,264,1784,1809]" captionTargetBox="[151,1436,166,1764]" captionTargetId="figure@39.[151,1436,166,1764]" captionTargetPageId="39" captionText="FIGURES 101–105. Thraulodes telegraphicus. 101–102, genitalia of the same specimen; 103, penis of another specimen; 104– 105, part of penis with telopenis forming spear-like roll.Abbreviations: bl, basal lobe; dp, dorsal projection of styliger; gd, gonoduct; gs1, gs2, gs3, gs4, segments of gonostylus; m.gs, gonostylar muscle; m.lat, lateral penial muscle; m.med, median penial muscle; m.s, styligeral muscle; m-v.r, medio-ventral ridge (recurved flap); sp, seminal pump; tp, spear-like roll (telopenis)." figureDoi="http://doi.org/10.5281/zenodo.3736925" httpUri="https://zenodo.org/record/3736925/files/figure.png" pageId="37" pageNumber="38">Fig. 102</figureCitation>
). Penis either entirely light ocher, or partly with brown cuticular or/and hypodermal coloration. Dorsal extension of styliger small and prominent. Penis lobes long and divergent; each lobe gradually widened apically, without lateral pouch; in other respects shape of penis lobes variable. Apico-lateral area usually forming well-expressed ear, occasionally without ear (
<figureCitation id="13052A35FFB884418E2FF8D5D99AF8EF" box="[579,676,1843,1869]" captionStart="FIGURES 112–120" captionStartId="41.[151,264,1889,1914]" captionTargetBox="[151,1423,170,1862]" captionTargetId="figure@41.[151,1436,166,1869]" captionTargetPageId="41" captionText="FIGURES 112–120. Thraulodes telegraphicus. 112, genitalia of male imago in lateral view; 113, apex of penis with spear-like roll in dorsal view; 114, genitalia of the same specimen in ventral view; 115, their subimaginal exuviae at the same magnification; 116, hind wing; 117–120, penis of four different specimens. Abbreviations: dp, dorsal projection of styliger; tp, spear-like roll (telopenis)." figureDoi="http://doi.org/10.5281/zenodo.3736929" httpUri="https://zenodo.org/record/3736929/files/figure.png" pageId="37" pageNumber="38">Fig. 119</figureCitation>
). Telopenes in form of spear-like rolls, more or less straight, arising on dorsal side of penes and directed medially or medially-caudally (not inclined dorsally or ventrally), with groove opened dorsally-laterally; length of spear-like rolls varies individually.
</paragraph>
<paragraph id="8B8136B0FFB884418CABF878DF30F87F" blockId="37.[151,1437,474,2013]" pageId="37" pageNumber="38">
<emphasis id="B94AEAA2FFB884418CABF878DA4EF81B" bold="true" box="[199,368,1950,1977]" italics="true" pageId="37" pageNumber="38">Female imago</emphasis>
(
<figureCitation id="13052A35FFB884418DEDF879DAD8F81B" box="[385,486,1951,1977]" captionStart="FIGURES 94–100" captionStartId="38.[151,264,1799,1824]" captionTargetBox="[151,1436,222,1780]" captionTargetId="figure@38.[151,1436,222,1780]" captionTargetPageId="38" captionText="FIGURES 94–100. Thraulodes telegraphicus. 94–95, right half of subimaginal exuviae of mesonotum of two different specimens; 96, apex of fore wing; 97–98, male imago in lateral and ventral view; 99, abdominal terga and sterna of male imago (black dots are algae under cuticle); 100, female imago. Abbreviations: acz.ms, achromozone of medioscutum; acz.sms, achromozone of submedioscutum; cz.ms, chromozone of medioscutum; cz.sms, chromozone of submedioscutum; mp.s, medioparapsidal suture; ms.cz.s, medioscutal chromozonal suture; sms.cz.s, submedioscutal chromozonal suture." figureDoi="http://doi.org/10.5281/zenodo.3736923" httpUri="https://zenodo.org/record/3736923/files/figure.png" pageId="37" pageNumber="38">Fig. 100</figureCitation>
). Coloration of head, thorax, legs, wings and caudalii as in male. Abdominal terga II–VI ocher with brownish; coloration of other abdominal segments as in male.
</paragraph>
<paragraph id="8B8136B0FFBB84428CABFF7FD889FF75" blockId="38.[151,1437,153,215]" pageId="38" pageNumber="39">
<emphasis id="B94AEAA2FFBB84428CABFF7FDA3FFF11" bold="true" box="[199,257,153,179]" italics="true" pageId="38" pageNumber="39">Eggs</emphasis>
(
<figureCitation id="13052A35FFBB84428D7CFF7FDA4FFF11" box="[272,369,153,179]" captionStart="FIGURE 121" captionStartId="42.[151,250,1423,1448]" captionTargetBox="[173,1414,166,1404]" captionTargetId="figure@42.[173,1414,166,1404]" captionTargetPageId="42" captionText="FIGURE 121. Thraulodes telegraphicus, eggs." figureDoi="http://doi.org/10.5281/zenodo.3736931" httpUri="https://zenodo.org/record/3736931/files/figure.png" pageId="38" pageNumber="39">Fig. 121</figureCitation>
). Mostly barrel-shaped. Each KTC surrounded by smooth, ring-like cover. Few small protuberances located close to these rings; other chorion without protuberances.
</paragraph>
<caption id="DF416638FFBB84428CFBF8E1DF5BF80D" ID-DOI="http://doi.org/10.5281/zenodo.3736923" ID-Zenodo-Dep="3736923" httpUri="https://zenodo.org/record/3736923/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,264,1799,1824]" targetBox="[151,1436,222,1780]" targetPageId="38">
<paragraph id="8B8136B0FFBB84428CFBF8E1DF5BF80D" blockId="38.[151,1437,1799,1968]" pageId="38" pageNumber="39">
<emphasis id="B94AEAA2FFBB84428CFBF8E1DA5FF882" bold="true" box="[151,353,1799,1824]" pageId="38" pageNumber="39">FIGURES 94–100.</emphasis>
<taxonomicName id="4C3E4D33FFBB84428D04F8EED952F882" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[360,620,1800,1824]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFBB84428D04F8EED952F882" box="[360,620,1800,1824]" italics="true" pageId="38" pageNumber="39">Thraulodes telegraphicus</emphasis>
</taxonomicName>
. 94–95, right half of subimaginal exuviae of mesonotum of two different specimens; 96, apex of fore wing; 97–98, male imago in lateral and ventral view; 99, abdominal terga and sterna of male imago (black dots are algae under cuticle); 100, female imago. Abbreviations: acz.ms, achromozone of medioscutum; acz.sms, achromozone of submedioscutum; cz.ms, chromozone of medioscutum; cz.sms, chromozone of submedioscutum; mp.s, medioparapsidal suture; ms.cz.s, medioscutal chromozonal suture; sms.cz.s, submedioscutal chromozonal suture.
</paragraph>
</caption>
<caption id="DF416638FFBA84438CFBF91EDE4CF8DF" ID-DOI="http://doi.org/10.5281/zenodo.3736925" ID-Zenodo-Dep="3736925" httpUri="https://zenodo.org/record/3736925/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,264,1784,1809]" targetBox="[151,1436,166,1764]" targetPageId="39">
<paragraph id="8B8136B0FFBA84438CFBF91EDE4CF8DF" blockId="39.[151,1437,1784,1917]" pageId="39" pageNumber="40">
<emphasis id="B94AEAA2FFBA84438CFBF91EDA53F8B3" bold="true" box="[151,365,1784,1809]" pageId="39" pageNumber="40">FIGURES 101–105.</emphasis>
<taxonomicName id="4C3E4D33FFBA84438D1EF91FD94AF8B3" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[370,628,1785,1809]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFBA84438D1EF91FD94AF8B3" box="[370,628,1785,1809]" italics="true" pageId="39" pageNumber="40">Thraulodes telegraphicus</emphasis>
</taxonomicName>
. 101–102, genitalia of the same specimen; 103, penis of another specimen; 104– 105, part of penis with telopenis forming spear-like roll.Abbreviations: bl, basal lobe; dp, dorsal projection of styliger; gd, gonoduct; gs1, gs2, gs3, gs4, segments of gonostylus; m.gs, gonostylar muscle; m.lat, lateral penial muscle; m.med, median penial muscle; m.s, styligeral muscle; m-v.r, medio-ventral ridge (recurved flap); sp, seminal pump; tp, spear-like roll (telopenis).
</paragraph>
</caption>
<caption id="DF416638FFB5844C8CFBF8CCD93BF855" ID-DOI="http://doi.org/10.5281/zenodo.3736927" ID-Zenodo-Dep="3736927" httpUri="https://zenodo.org/record/3736927/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,264,1834,1859]" targetBox="[159,1390,175,1807]" targetPageId="40">
<paragraph id="8B8136B0FFB5844C8CFBF8CCD93BF855" blockId="40.[151,1437,1834,2039]" pageId="40" pageNumber="41">
<emphasis id="B94AEAA2FFB5844C8CFBF8CCDA50F8E1" bold="true" box="[151,366,1834,1859]" pageId="40" pageNumber="41">FIGURES 106–111.</emphasis>
<taxonomicName id="4C3E4D33FFB5844C8D1AF8CCD942F8E0" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[374,636,1834,1858]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFB5844C8D1AF8CCD942F8E0" box="[374,636,1834,1858]" italics="true" pageId="40" pageNumber="41">Thraulodes telegraphicus</emphasis>
</taxonomicName>
. 106, protogonostyli of last instar larva (hypoderm of gonostyli and styliger and hidden cuticle of protopenis shown by interrupted lines); 107, enlarged cuticle of protopenis and hypoderm of developing penis inside; 108–109, the same, at later stage of development; 110–111, genital of male imago (110, styliger ventrally; 111, penis dorsally at left and ventrally at right). Abbreviation: bl, basal lobe; dp, dorsal projection of styliger; gd, gonoduct; m.gs, gonostylar muscle; m.med, median penial muscle; m.s, styligeral muscle; m-v.r, medio-ventral ridge (recurved flap); sp, seminal pump; tp, spear-like roll (telopenis).
</paragraph>
</caption>
<caption id="DF416638FFB4844D8CFBF887DA0AF844" ID-DOI="http://doi.org/10.5281/zenodo.3736929" ID-Zenodo-Dep="3736929" httpUri="https://zenodo.org/record/3736929/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,264,1889,1914]" targetBox="[151,1423,170,1862]" targetPageId="41">
<paragraph id="8B8136B0FFB4844D8CFBF887DA0AF844" blockId="41.[151,1437,1889,2022]" pageId="41" pageNumber="42">
<emphasis id="B94AEAA2FFB4844D8CFBF887DA52F8D8" bold="true" box="[151,364,1889,1914]" pageId="41" pageNumber="42">FIGURES 112–120.</emphasis>
<taxonomicName id="4C3E4D33FFB4844D8D1EF887D94BF8DB" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[370,629,1889,1913]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFB4844D8D1EF887D94BF8DB" box="[370,629,1889,1913]" italics="true" pageId="41" pageNumber="42">Thraulodes telegraphicus</emphasis>
</taxonomicName>
. 112, genitalia of male imago in lateral view; 113, apex of penis with spear-like roll in dorsal view; 114, genitalia of the same specimen in ventral view; 115, their subimaginal exuviae at the same magnification; 116, hind wing; 117–120, penis of four different specimens. Abbreviations: dp, dorsal projection of styliger; tp, spear-like roll (telopenis).
</paragraph>
</caption>
<caption id="DF416638FFB7844E8CFBFA69D947FA0B" ID-DOI="http://doi.org/10.5281/zenodo.3736931" ID-Zenodo-Dep="3736931" box="[151,633,1423,1449]" httpUri="https://zenodo.org/record/3736931/files/figure.png" pageId="42" pageNumber="43" startId="42.[151,250,1423,1448]" targetBox="[173,1414,166,1404]" targetPageId="42">
<paragraph id="8B8136B0FFB7844E8CFBFA69D947FA0B" blockId="42.[151,633,1423,1449]" box="[151,633,1423,1449]" pageId="42" pageNumber="43">
<emphasis id="B94AEAA2FFB7844E8CFBFA69DA13FA0A" bold="true" box="[151,301,1423,1448]" pageId="42" pageNumber="43">FIGURE 121.</emphasis>
<taxonomicName id="4C3E4D33FFB7844E8D5FFA76D908FA0A" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[307,566,1424,1448]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFB7844E8D5FFA76D908FA0A" box="[307,566,1424,1448]" italics="true" pageId="42" pageNumber="43">Thraulodes telegraphicus</emphasis>
</taxonomicName>
, eggs.
</paragraph>
</caption>
<paragraph id="8B8136B0FFB7844E8CABFA1CDA80F99B" blockId="42.[151,1437,1530,2025]" pageId="42" pageNumber="43">
<emphasis id="B94AEAA2FFB7844E8CABFA1CDA70F9B7" bold="true" box="[199,334,1530,1557]" pageId="42" pageNumber="43">Dimension.</emphasis>
Fore wing length (and approximate body length)
<quantity id="4CC69B55FFB7844E8FFDFA1CDF25F9B7" box="[913,1051,1530,1557]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.5" metricValueMin="7.5" pageId="42" pageNumber="43" unit="mm" value="8.5" valueMax="9.5" valueMin="7.5">7.5–9.5 mm</quantity>
(this study); 10–11 (according to
<bibRefCitation id="EFAF4B41FFB7844E8CFBF9F9DA8FF99B" author="Traver, J. R. &amp; Edmunds, G. F. Jr." box="[151,433,1566,1593]" pageId="42" pageNumber="43" pagination="349 - 395" refId="ref72596" refString="Traver, J. R. &amp; Edmunds, G. F. Jr. (1967) A revision of the genus Thraulodes (Ephemeroptera: Leptophlebiidae). Miscellaneous Publications of the Entomological Society of America, 5 (8), 349 - 395." type="journal article" year="1967">Traver &amp; Edmunds 1967</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFB7844E8CABF9A4D861F9FF" box="[199,863,1602,1629]" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8B8136B0FFB7844E8CABF9A4D861F9FF" blockId="42.[151,1437,1530,2025]" box="[199,863,1602,1629]" pageId="42" pageNumber="43">
<emphasis id="B94AEAA2FFB7844E8CABF9A4DA5FF9FF" bold="true" box="[199,353,1602,1629]" pageId="42" pageNumber="43">Distribution.</emphasis>
<collectingCountry id="F3297620FFB7844E8D0BF9A5DAA4F9FF" box="[359,410,1603,1629]" name="Peru" pageId="42" pageNumber="43">Peru</collectingCountry>
(
<bibRefCitation id="EFAF4B41FFB7844E8DC5F9A4D9E8F9FF" author="Needham, J. G. &amp; Murphy, H. E." box="[425,726,1602,1629]" pageId="42" pageNumber="43" pagination="1 - 79" refId="ref71682" refString="Needham, J. G. &amp; Murphy, H. E. (1924) Neotropical mayflies. Bulletin of the Lloyd Library, Entomological Series 4, 24, 1 - 79." type="journal article" year="1924">Needham &amp; Murphy 1924</bibRefCitation>
; this study).
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFB7844F8CABF980DF79FF5E" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8B8136B0FFB7844E8CABF980DE31F94E" blockId="42.[151,1437,1530,2025]" pageId="42" pageNumber="43">
<emphasis id="B94AEAA2FFB7844E8CABF980DA75F923" bold="true" box="[199,331,1638,1665]" pageId="42" pageNumber="43">Synonymy.</emphasis>
<taxonomicName id="4C3E4D33FFB7844E8D3EF981D929F923" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[338,535,1639,1665]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFB7844E8D3EF981D929F923" box="[338,535,1639,1665]" italics="true" pageId="42" pageNumber="43">Th. telegraphicus</emphasis>
</taxonomicName>
is one of the most common mayfly species in low altitudes of eastern slopes of Andes (regions
<collectingRegion id="49FAF852FFB7844E8D22F96CDAB5F906" box="[334,395,1674,1700]" country="Peru" name="Junin" pageId="42" pageNumber="43">Junin</collectingRegion>
and
<collectingRegion id="49FAF852FFB7844E8DAAF96DD915F907" box="[454,555,1675,1701]" country="Peru" name="Huanuco" pageId="42" pageNumber="43">Huanuco</collectingRegion>
of
<collectingCountry id="F3297620FFB7844E8E3AF96DD9B0F907" box="[598,654,1675,1701]" name="Peru" pageId="42" pageNumber="43">Peru</collectingCountry>
). Originally (
<bibRefCitation id="EFAF4B41FFB7844E8F4BF96CDF63F907" author="Needham, J. G. &amp; Murphy, H. E." box="[807,1117,1674,1701]" pageId="42" pageNumber="43" pagination="1 - 79" refId="ref71682" refString="Needham, J. G. &amp; Murphy, H. E. (1924) Neotropical mayflies. Bulletin of the Lloyd Library, Entomological Series 4, 24, 1 - 79." type="journal article" year="1924">Needham &amp; Murphy 1924</bibRefCitation>
), imago of this species was described from the Region of
<collectingRegion id="49FAF852FFB7844E8D8FF948D91EF96A" box="[483,544,1710,1736]" country="Peru" name="Junin" pageId="42" pageNumber="43">Junin</collectingRegion>
under the name
<taxonomicName id="4C3E4D33FFB7844E8EB4F949D8A5F96B" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[728,923,1711,1737]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFB7844E8EB4F949D8A5F96B" box="[728,923,1711,1737]" italics="true" pageId="42" pageNumber="43">Th. telegraphicus</emphasis>
</taxonomicName>
, and subimago—under the name
<taxonomicName id="4C3E4D33FFB7844E897EF949DEA8F96B" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[1298,1430,1711,1737]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="plicatus">
<emphasis id="B94AEAA2FFB7844E897EF949DEA8F96B" box="[1298,1430,1711,1737]" italics="true" pageId="42" pageNumber="43">Th. plicatus</emphasis>
</taxonomicName>
. Synonymy of these two names was established by
<bibRefCitation id="EFAF4B41FFB7844E8EA2F935D852F94F" author="Traver, J. R." box="[718,876,1746,1773]" pageId="42" pageNumber="43" pagination="418 - 436" refId="ref72450" refString="Traver, J. R. (1946) Notes on Neotropical mayflies. Part I. Family Baetidae, subfamily Leptophlebiinae. Revista de Entomologia, 17, 418 - 436." type="journal article" year="1946">Traver (1946)</bibRefCitation>
, who reexamined the both
<typeStatus id="54858812FFB7844E88F7F934DE32F94E" box="[1179,1292,1746,1772]" pageId="42" pageNumber="43" type="holotype">holotypes</typeStatus>
.
</paragraph>
<paragraph id="8B8136B0FFB7844F8CABF911DF79FF5E" blockId="42.[151,1437,1530,2025]" lastBlockId="43.[151,1436,153,252]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
Larvae were described by
<bibRefCitation id="EFAF4B41FFB7844E8D9AF910D9A0F8B3" author="Roback, S. S." box="[502,670,1782,1809]" pageId="42" pageNumber="43" pagination="129 - 199" refId="ref72121" refString="Roback, S. S. (1966) The Catherwood Foundation Peruvia-Amazon expedition. VI- Ephemeroptera nymphs. Monographs of the Academy of Natural Sciences of Philadelphia, No. 14, 129 - 199." type="book chapter" year="1966">Roback (1966)</bibRefCitation>
from Region of
<collectingRegion id="49FAF852FFB7844E8F0CF911D8FBF8B3" box="[864,965,1783,1809]" country="Peru" name="Huanuco" pageId="42" pageNumber="43">Huanuco</collectingRegion>
without species names. Probably, among six larval forms distinguished by Roback, the forms reported as
<taxonomicName id="4C3E4D33FFB7844E8F1FF8FDD8CDF897" authorityName="Ulmer" authorityYear="1920" box="[883,1011,1819,1845]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAA2FFB7844E8F1FF8FDD8CDF897" box="[883,1011,1819,1845]" italics="true" pageId="42" pageNumber="43">Thraulodes</emphasis>
</taxonomicName>
sp. 1,
<emphasis id="B94AEAA2FFB7844E882CF8FDDF60F897" box="[1088,1118,1819,1845]" italics="true" pageId="42" pageNumber="43">Th</emphasis>
. sp. 2,
<emphasis id="B94AEAA2FFB7844E88DFF8FDDFEFF897" box="[1203,1233,1819,1845]" italics="true" pageId="42" pageNumber="43">Th</emphasis>
. sp. 4 and
<emphasis id="B94AEAA2FFB7844E8923F8FDDE53F897" box="[1359,1389,1819,1845]" italics="true" pageId="42" pageNumber="43">Th</emphasis>
. sp. 6 belong to
<taxonomicName id="4C3E4D33FFB7844E8D72F8D9DADCF8FB" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[286,482,1855,1881]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFB7844E8D72F8D9DADCF8FB" box="[286,482,1855,1881]" italics="true" pageId="42" pageNumber="43">Th. telegraphicus</emphasis>
</taxonomicName>
. The forms «2» and «6» have contrastingly light pronotum and contrasting dark and light areas on mesonotum and abdomen (as in
<figureCitation id="13052A35FFB7844E8EF0F884D9CEF8DE" box="[668,752,1890,1916]" captionStart="FIGURES 66–69" captionStartId="34.[151,264,1320,1345]" captionTargetBox="[151,1436,166,1301]" captionTargetId="figure@34.[151,1436,166,1301]" captionTargetPageId="34" captionText="FIGURES 66–69. Thraulodes telegraphicus. 66–67, last instar larvae with intensive cuticular coloration and poorly visible hypodermal coloration; 68, younger larva with light cuticle and well visible blackish hypodermal markings on abdomen; 69, claw. Abbreviations: artic.p., articulatory portion; rigid p., rigid portion of claw." figureDoi="http://doi.org/10.5281/zenodo.3736917" httpUri="https://zenodo.org/record/3736917/files/figure.png" pageId="42" pageNumber="43">Figs 66</figureCitation>
,
<figureCitation id="13052A35FFB7844E8E90F885D853F8DF" box="[764,877,1891,1917]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="42" pageNumber="43">70–73, 81</figureCitation>
), while the forms «1» and «4» have more uniform brown coloration (as in
<figureCitation id="13052A35FFB7844E8DC5F860D93EF802" box="[425,512,1926,1952]" captionStart="FIGURES 66–69" captionStartId="34.[151,264,1320,1345]" captionTargetBox="[151,1436,166,1301]" captionTargetId="figure@34.[151,1436,166,1301]" captionTargetPageId="34" captionText="FIGURES 66–69. Thraulodes telegraphicus. 66–67, last instar larvae with intensive cuticular coloration and poorly visible hypodermal coloration; 68, younger larva with light cuticle and well visible blackish hypodermal markings on abdomen; 69, claw. Abbreviations: artic.p., articulatory portion; rigid p., rigid portion of claw." figureDoi="http://doi.org/10.5281/zenodo.3736917" httpUri="https://zenodo.org/record/3736917/files/figure.png" pageId="42" pageNumber="43">Figs 67</figureCitation>
,
<figureCitation id="13052A35FFB7844E8E62F860D964F803" box="[526,602,1926,1953]" captionStart="FIGURES 70–85" captionStartId="36.[151,264,188,213]" captionTargetId="figure@35.[151,1436,181,1986]" captionTargetPageId="35" captionText="FIGURES 70–85. Thraulodes telegraphicus, larval exuviae (70–82 at the same magnification): 70–75, right half of pronotum and mesonotum; 76, head with labrum and mandibles; 77–78 and 79–80, fore and hind legs of two specimens; 81–82, abdominal terga; 83, anterior side of hind femur; 84, anterior side of fore tibia; 85, anterior side of hind tibia. Abbreviations: i-a.r, inneranterior row of stout setae; o-a.r, outer-anterior row of stout setae; o-p.r, outer-posterior row of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736919" httpUri="https://zenodo.org/record/3736919/files/figure.png" pageId="42" pageNumber="43">75, 82</figureCitation>
). The forms «1» and «2» have tergalii with tracheal branches by sides of main trachea, while the forms «4» and «6» are said to lack these branches. Among material examined by me, all larvae have side trachea, but in some larvae from Huacar the side trachea are colorless and can be overlooked. Two other larval forms described by Roback, are different from
<taxonomicName id="4C3E4D33FFB6844F8F40FF7FD8CEFF11" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[812,1008,153,179]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFB6844F8F40FF7FD8CEFF11" box="[812,1008,153,179]" italics="true" pageId="43" pageNumber="44">Th. telegraphicus</emphasis>
</taxonomicName>
. The form «3» (describes as one larva from Puente Perez) differs from
<taxonomicName id="4C3E4D33FFB6844F8E65FF5BD9F0FF75" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[521,718,189,215]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="telegraphicus">
<emphasis id="B94AEAA2FFB6844F8E65FF5BD9F0FF75" box="[521,718,189,215]" italics="true" pageId="43" pageNumber="44">Th. telegraphicus</emphasis>
</taxonomicName>
by acute small stout femoral setae (
<bibRefCitation id="EFAF4B41FFB6844F8804FF5BDE3AFF75" author="Roback, S. S." box="[1128,1284,189,215]" pageId="43" pageNumber="44" pagination="129 - 199" refId="ref72121" refString="Roback, S. S. (1966) The Catherwood Foundation Peruvia-Amazon expedition. VI- Ephemeroptera nymphs. Monographs of the Academy of Natural Sciences of Philadelphia, No. 14, 129 - 199." type="book chapter" year="1966">Roback 1966</bibRefCitation>
: fig. 184); its systematic position remains unknown. The form «5» is
<taxonomicName id="4C3E4D33FFB6844F8F6AFF04D888FF59" authorityName="Flowers" authorityYear="2009" box="[774,950,225,251]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="quevedoensis">
<emphasis id="B94AEAA2FFB6844F8F6AFF04D888FF59" box="[774,950,225,251]" italics="true" pageId="43" pageNumber="44">T. quevedoensis</emphasis>
</taxonomicName>
(see below).
</paragraph>
</subSubSection>
</treatment>
</document>
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