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<document id="A941BDB0F2954DA5274C76386C7748EC" ID-CLB-Dataset="51608" ID-DOI="10.11646/zootaxa.4091.1.1" ID-GBIF-Dataset="bdd057d5-b4d6-4b57-940a-d7839f483e25" ID-ISSN="1175-5326" ID-Zenodo-Dep="265332" ID-ZooBank="urn:lsid:zoobank.org:pub:76D13D36-682E-4E91-AC91-693CA9D3D465" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457950950343" checkinUser="plazi" docAuthor="Chris J. Hodgson &amp; Douglas J. Williams" docDate="2016" docId="03F2FF4881350D1E24B6A8D1FC63FBF3" docLanguage="en" docName="15613-5135-1-PB.pdf.imf" docOrigin="Zootaxa 4091 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Antecerococcus ankaratrae Mamet, comb. nov." docType="treatment" docVersion="16" lastPageNumber="33" masterDocId="FFCB8730812B0D3E2421AB7EFF8EFF8E" masterDocTitle="(Hemiptera: Sternorrhyncha, Coccomorpha) with particular reference to species from the Afrotropical, western Palaearctic and western Oriental Regions, with the revival of Antecerococcus Green and description of a new genus and fifteen new species, and with ten new synonomies" masterLastPageNumber="175" masterPageNumber="1" pageNumber="31" updateTime="1698185172771" updateUser="ExternalLinkService">
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<mods:title id="47B88BF2BB77CAB14AD62FCA97613E5A">(Hemiptera: Sternorrhyncha, Coccomorpha) with particular reference to species from the Afrotropical, western Palaearctic and western Oriental Regions, with the revival of Antecerococcus Green and description of a new genus and fifteen new species, and with ten new synonomies</mods:title>
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<mods:namePart id="D6C324CC5F5CAC74CC61355D9BF21488">Chris J. Hodgson</mods:namePart>
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<mods:namePart id="1A61DE0220C7152BEFE0087CEE6A8F16">Douglas J. Williams</mods:namePart>
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<mods:date id="2E908E6CDDD067DB3D529876746899E4">2016</mods:date>
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<treatment id="03F2FF4881350D1E24B6A8D1FC63FBF3" ID-DOI="http://doi.org/10.5281/zenodo.6081542" ID-GBIF-Taxon="117911268" ID-Zenodo-Dep="6081542" LSID="urn:lsid:plazi:treatment:03F2FF4881350D1E24B6A8D1FC63FBF3" httpUri="http://treatment.plazi.org/id/03F2FF4881350D1E24B6A8D1FC63FBF3" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
<subSubSection id="C3411DD581350D2024B6A8D1FDB7FBBE" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BE44E5E81350D2024B6A8D1FD65FC44" blockId="30.[151,747,943,1003]" box="[151,747,943,970]" pageId="30" pageNumber="31">
<heading id="D0ACF93281350D2024B6A8D1FD65FC44" bold="true" box="[151,747,943,970]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<emphasis id="B92F924C81350D2024B6A8D1FD65FC44" bold="true" box="[151,747,943,970]" pageId="30" pageNumber="31">
<taxonomicName id="4C5B35DD81350D2024B6A8D1FDD6FC44" authority="Mamet" authorityName="Mamet" box="[151,600,943,970]" class="Insecta" family="Cerococcidae" genus="Antecerococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="ankaratrae" status="comb. nov.">
<emphasis id="B92F924C81350D2024B6A8D1FE6CFC47" bold="true" box="[151,482,943,969]" italics="true" pageId="30" pageNumber="31">Antecerococcus ankaratrae</emphasis>
(Mamet)
</taxonomicName>
,
<taxonomicNameLabel id="A21C2F3781350D202644A8CEFD65FC44" box="[613,747,944,970]" pageId="30" pageNumber="31">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE44E5E81350D2024B6A8ACFF65FC65" blockId="30.[151,747,943,1003]" box="[151,235,978,1003]" pageId="30" pageNumber="31">
(
<figureCitation id="136052DB81350D2024BEA8ACFF6CFC65" box="[159,226,978,1003]" captionStart="FIGURE 7" captionStartId="31.[151,250,1908,1930]" captionTargetBox="[223,1404,268,1823]" captionTargetId="figure@31.[163,1417,196,1872]" captionTargetPageId="31" captionText="FIGURE 7. Antecerococcus ankaratrae (Mamet). Adult female." httpUri="https://zenodo.org/record/265338/files/figure.png" pageId="30" pageNumber="31">Fig. 7</figureCitation>
)
</paragraph>
<paragraph id="8BE44E5E81350D2024B6AF64FDB7FBBE" blockId="30.[151,569,1050,1072]" box="[151,569,1050,1072]" pageId="30" pageNumber="31">
<taxonomicName id="4C5B35DD81350D2024B6AF64FDBBFBBE" authority="Mamet 1954: 43" authorityName="Mamet" authorityPageNumber="43" authorityYear="1954" box="[151,565,1050,1072]" class="Insecta" family="Cerococcidae" genus="Cerococcus" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="ankaratrae">
<emphasis id="B92F924C81350D2024B6AF64FE0BFBBE" box="[151,389,1050,1072]" italics="true" pageId="30" pageNumber="31">Cerococcus ankaratrae</emphasis>
Mamet 1954: 43
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3411DD581350D2024B6AF20FF5EF800" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BE44E5E81350D2024B6AF20FC1BFB6C" blockId="30.[151,1437,1117,2006]" pageId="30" pageNumber="31">
<emphasis id="B92F924C81350D2024B6AF20FD8FFBF8" bold="true" box="[151,513,1117,1142]" pageId="30" pageNumber="31">
<typeStatus id="54E0F0FC81350D2024B6AF20FF5CFBFB" box="[151,210,1118,1141]" pageId="30" pageNumber="31">Type</typeStatus>
details.
<collectingCountry id="F34C0ECE81350D202515AF23FD8FFBF8" box="[308,513,1117,1142]" name="Madagascar" pageId="30" pageNumber="31">MADAGASCAR</collectingCountry>
</emphasis>
, Ankaratra, Manjakatompo,
<quantity id="4CA3E3BB81350D202767AF20FC1DFBF8" box="[838,915,1118,1142]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="30" pageNumber="31" unit="m" value="2000.0">2000m</quantity>
alt., on
<taxonomicName id="4C5B35DD81350D2027CFAF20FBB2FBF8" box="[1006,1084,1118,1142]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="B92F924C81350D2027CFAF20FBB2FBF8" box="[1006,1084,1118,1142]" italics="true" pageId="30" pageNumber="31">Croton</emphasis>
</taxonomicName>
sp.,
<date id="FFE5689E81350D202053AF20FB6CFBF8" box="[1138,1250,1118,1142]" pageId="30" pageNumber="31" value="1950-05-24">24.v.1950</date>
, R. Mamet &amp; R. Paulian.
<emphasis id="B92F924C81350D202521AFFFFE0DFB14" bold="true" box="[256,387,1153,1178]" pageId="30" pageNumber="31">Depository</emphasis>
:
<emphasis id="B92F924C81350D2025BBAFFCFDFFFB17" bold="true" box="[410,625,1154,1177]" pageId="30" pageNumber="31">
MNHN:
<typeStatus id="54E0F0FC81350D20262BAFFCFDFFFB17" box="[522,625,1154,1177]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
</emphasis>
adf (MNHN 14876-1);
<typeStatus id="54E0F0FC81350D2027BDAFFCFB9CFB17" box="[924,1042,1154,1177]" pageId="30" pageNumber="31" type="paratype">
<emphasis id="B92F924C81350D2027BDAFFCFB9CFB17" bold="true" box="[924,1042,1154,1177]" pageId="30" pageNumber="31">paratypes</emphasis>
</typeStatus>
, same data as
<typeStatus id="54E0F0FC81350D2020F5AFFFFAB7FB14" box="[1236,1337,1153,1178]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
, 7/8adff (MNHN 14876-2 (1 adf), MNHN 14876-3 (1 adf), MNHN 14876-4 (1 adf), MNHN 14876-5 (1 adf), MNHN 14876-6 (1 adf), MNHN 14876-7 (1 adf), MNHN 14876-8 (2 adff)).
</paragraph>
<paragraph id="8BE44E5E81350D2024E6AF93FD1DFAA4" blockId="30.[151,1437,1117,2006]" pageId="30" pageNumber="31">
<emphasis id="B92F924C81350D2024E6AF93FE1AFA88" bold="true" box="[199,404,1261,1286]" pageId="30" pageNumber="31">Material studied.</emphasis>
<typeStatus id="54E0F0FC81350D2025BBAF90FD89FA8B" box="[410,519,1262,1285]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F34C0ECE81350D202637AF90FD6EFA88" box="[534,736,1262,1286]" name="Madagascar" pageId="30" pageNumber="31">MADAGASCAR</collectingCountry>
, Manjakatompo, on
<taxonomicName id="4C5B35DD81350D2027E4AF90FB9DFA88" box="[965,1043,1262,1286]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="B92F924C81350D2027E4AF90FB9DFA88" box="[965,1043,1262,1286]" italics="true" pageId="30" pageNumber="31">Croton</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C5B35DD81350D202069AF93FB7AFA88" box="[1096,1268,1261,1286]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
),
<date id="FFE5689E81350D202126AF90FAF9FA88" box="[1287,1399,1262,1286]" pageId="30" pageNumber="31" value="1950-05-24">24.v.1950</date>
, R. Mamet &amp; R. Paulian (MNHN): 3/3adff (fg).
</paragraph>
<paragraph id="8BE44E5E81350D2024E6AE48FD3EFAC0" blockId="30.[151,1437,1117,2006]" box="[199,688,1333,1358]" pageId="30" pageNumber="31">Note: description taken from all specimens.</paragraph>
<paragraph id="8BE44E5E81350D2024E6AE24FB23FAFC" blockId="30.[151,1437,1117,2006]" box="[199,1197,1369,1394]" pageId="30" pageNumber="31">
<emphasis id="B92F924C81350D2024E6AE24FE29FAFC" bold="true" box="[199,423,1369,1394]" pageId="30" pageNumber="31">Mounted material.</emphasis>
Body roundly pear-shaped,
<quantity id="4CA3E3BB81350D2026C5AE24FCE5FAFF" box="[740,875,1370,1394]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="30" pageNumber="31" unit="mm" value="1.25" valueMax="1.4" valueMin="1.1">1.11.4 mm</quantity>
long, and 0.81.0 mm wide.
</paragraph>
<paragraph id="8BE44E5E81350D2024E6AE00FF5EF800" blockId="30.[151,1437,1117,2006]" pageId="30" pageNumber="31">
<emphasis id="B92F924C81350D2024E6AE00FEA0FA1B" bold="true" box="[199,302,1406,1429]" pageId="30" pageNumber="31">Dorsum.</emphasis>
Eight-shaped pores of 3 sizes: (i) a large pore, each 15
<date id="FFE5689E81350D20278DAE00FBA4FA18" box="[940,1066,1406,1430]" pageId="30" pageNumber="31" value="1917-10-11" valueMax="1917-10-12" valueMin="1917-10-11">17 x 1112</date>
µm, in a sparse lace-like pattern over all of dorsum but absent from posterior to cribriform plates apart from 4 pores on each side of posterior abdominal segments; (ii) an intermediate-sized pore sparse throughout rest of dorsum, somewhat variable in size (6.510.0 x 4.06.5µm), tending to be smaller in spaces between lace-like pattern and also on more posterior abdominal segments; and (iii) a small pore, each
<date id="FFE5689E81350D2026F3AD70FD76F9AB" box="[722,760,1550,1573]" pageId="30" pageNumber="31">5 x</date>
3 µm, singly in apex of each stigmatic pore band. Simple pores, each 2.02.5 µm wide; sparse. Cribriform plates small, mainly roundish, each 1015 µm wide, with a broad margin and moderate-sized micropores; present in submedial group of 2 plates on each side of abdominal segment IV. Dorsal setae few, each setose, and mainly 5 µm long. Tubular ducts with outer ducts about 25 µm long and about 3 µm wide, subequal in size to those on venter; abundant throughout. Anal lobes mainly membranous, but with sclerotized inner margins, each sclerotization with striations or reticulations; each lobe about 80 µm long with a long apical seta 180195 µm long; more apical fleshy setae on each dorsal surface short, 1416 µm long; more basal fleshy setae each 2329 µm long; ventral setose seta near apex each 1117 µm long; medioventral or outer margin setae each about 1114 µm long; each lobe with 1 small 8-shaped pore. Median anal plate quite elongate and bluntly triangular, 3840 µm long, 4045 µm wide at base. Anal ring with 4 pairs of setae, each 6575 µm long.
</paragraph>
</subSubSection>
<subSubSection id="C3411DD581350D1E24E6ACE4FD63FD0B" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
<paragraph id="8BE44E5E81350D1E24E6ACE4FD63FD0B" blockId="30.[151,1437,1117,2006]" lastBlockId="32.[151,1437,151,1149]" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
<emphasis id="B92F924C81350D2024E6ACE4FE96F83F" bold="true" box="[199,280,1946,1969]" pageId="30" pageNumber="31">Venter.</emphasis>
Eight-shaped pores similar to intermediate-sized pores on dorsum, in a sparse, fairly narrow, submarginal band and also in sparse transverse bands about 1 pore wide across abdominal segments; some on inner margin of submarginal band asymetrical. Simple pores similar to those on dorsum but very sparse. Small bilocular pores, each about 5 µm at widest, frequent medially on head and thorax. Spiracular disc-pores small, each 35 µm wide, mainly with 5 loculi, in single, unbifurcated bands mainly 25 pores wide but widening slightly at apex on dorsum; with
<quantity id="4CA3E3BB810B0D1E2516AA7BFE13FE93" box="[311,413,260,285]" metricMagnitude="0" metricUnit="m" metricValue="1.6509999999999998" metricValueMax="1.905" metricValueMin="1.3969999999999998" pageId="32" pageNumber="33" unit="in" value="65.0" valueMax="75.0" valueMin="55.0">5575 in</quantity>
each band; each band with 02 small 8-shaped pores in apical group; no quinquelocular disc-pores found associated with each antenna (see under Comment below). Small convex closed pores absent. Multilocular disc-pores present in sparse transverse rows 12 pores wide, as follows: abdominal segment VIII with 26 on each side; VII with 46 on each side of vulva; VI with 13 submarginally + 1417 medially;
<date id="FFE5689E810B0D1E2172AA0EFA12FE06" box="[1363,1436,368,392]" pageId="32" pageNumber="33">V 24</date>
submarginally + 1522 medially;
<date id="FFE5689E810B0D1E262EAAEBFDD3FE22" box="[527,605,404,428]" pageId="32" pageNumber="33">IV 36</date>
submarginally + 1721 medially;
<date id="FFE5689E810B0D1E27FDAAEBFBADFE22" box="[988,1059,404,428]" pageId="32" pageNumber="33">III 4-8</date>
submarginally + 2431 medially;
<date id="FFE5689E810B0D1E24B6AAC6FF69FE5E" box="[151,231,440,464]" pageId="32" pageNumber="33" value="2011-02-05">II 511</date>
submarginally + 1523 medially; metathorax with a group of 915 laterad to each metathoracic leg stub but 0 medially, and occasionally with 1 laterad to a mesothoracic leg. Tubular ducts similar to those on dorsum, but those posteriorly in abdomen tending to be narrower. Ventral setae all setose and short; preanal setae each about 6570 µm long; companion setae short. Leg stubs small. Antennae unsegmented, each 2535 µm long, with 7 or 8 mainly fleshy setae, without either a cone-like pointed apex or a distinct setal cavity. Clypeolabral shield 105110 µm long. Spiracular peritremes each 2528 µm wide.
</paragraph>
</subSubSection>
<caption id="DF241ED681340D2124B6AC0AFCA9F804" box="[151,807,1908,1930]" httpUri="https://zenodo.org/record/265338/files/figure.png" pageId="31" pageNumber="32" targetBox="[223,1404,268,1823]" targetPageId="31">
<paragraph id="8BE44E5E81340D2124B6AC0AFCA9F804" blockId="31.[151,807,1908,1930]" box="[151,807,1908,1930]" pageId="31" pageNumber="32">
<emphasis id="B92F924C81340D2124B6AC0AFE9CF804" bold="true" box="[151,274,1908,1930]" pageId="31" pageNumber="32">FIGURE 7.</emphasis>
<taxonomicName id="4C5B35DD81340D212538AC0AFD1FF804" authority="Mamet" authorityName="Mamet" box="[281,657,1908,1930]" class="Insecta" family="Cerococcidae" genus="Antecerococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="ankaratrae">
<emphasis id="B92F924C81340D212538AC0AFDA1F804" box="[281,559,1908,1930]" italics="true" pageId="31" pageNumber="32">Antecerococcus ankaratrae</emphasis>
(Mamet)
</taxonomicName>
. Adult female.
</paragraph>
</caption>
<subSubSection id="C3411DD5810B0D1E24E6A9F1FC63FBF3" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8BE44E5E810B0D1E24E6A9F1FF55FCB9" blockId="32.[151,1437,151,1149]" pageId="32" pageNumber="33">
<emphasis id="B92F924C810B0D1E24E6A9F1FEB3FD26" bold="true" box="[199,317,655,680]" pageId="32" pageNumber="33">Comment</emphasis>
. The above description is similar to that of Lambdin and Kostarab (1977) except that they: (i) recorded one or two quinquelocular disc-pores near each antenna (not found here and not noted by Mamet, 1954), and (ii) they show no setose setae on the ventral apex of each anal lobe. Mamet (1954) illustrated the ventral 8- shaped pores as being throughout the venter but he clearly mistook the small bilocular pores for normal 8-shaped pores.
</paragraph>
<paragraph id="8BE44E5E810B0D1E24E6A83AFD31FBBB" blockId="32.[151,1437,151,1149]" pageId="32" pageNumber="33">
Adult females of
<taxonomicName id="4C5B35DD810B0D1E25ADA83AFDA7FCD3" box="[396,553,836,861]" class="Insecta" family="Cerococcidae" genus="Antecerococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ankaratrae">
<emphasis id="B92F924C810B0D1E25ADA83AFDA7FCD3" box="[396,553,836,861]" italics="true" pageId="32" pageNumber="33">A. ankaratrae</emphasis>
</taxonomicName>
are characterised by the following combination of character-states: (i) dorsum with three sizes of 8-shaped pore; (ii) larger 8-shaped pores on dorsum in a lace-like pattern; (iii) single small 8- shaped pores present in apex of each stigmatic band; (iv) four large 8-shaped pores present on each side of posterior abdominal segments; (v) only two cribriform plates present on each side of abdominal segment IV; (vi) median anal plate rather elongate; (vii) posterior stigmatic pore bands not bifurcate; (viii) leg stubs small; (ix) multilocular disc-pores in sparse bands across all abdominal segments and submarginally on metathorax, and (x) antennae without a cone-like apex or setal cavity.
</paragraph>
<paragraph id="8BE44E5E810B0D1E24E6AF41FC63FBF3" blockId="32.[151,1437,151,1149]" pageId="32" pageNumber="33">
The adult female of
<taxonomicName id="4C5B35DD810B0D1E258FAF3EFDC5FBD6" box="[430,587,1087,1112]" class="Insecta" family="Cerococcidae" genus="Antecerococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ankaratrae">
<emphasis id="B92F924C810B0D1E258FAF3EFDC5FBD6" box="[430,587,1087,1112]" italics="true" pageId="32" pageNumber="33">A. ankaratrae</emphasis>
</taxonomicName>
falls within Group A in the key to species of
<taxonomicName id="4C5B35DD810B0D1E2075AF3EFA8AFBD6" box="[1108,1284,1088,1112]" class="Insecta" family="Cerococcidae" genus="Antecerococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B92F924C810B0D1E2075AF3EFA8AFBD6" box="[1108,1284,1088,1112]" italics="true" pageId="32" pageNumber="33">Antecerococcus</emphasis>
</taxonomicName>
and keys out close to other species from
<collectingCountry id="F34C0ECE810B0D1E25EBAF1AFDDFFBF3" box="[458,593,1124,1149]" name="Madagascar" pageId="32" pageNumber="33">Madagascar</collectingCountry>
and to
<taxonomicName id="4C5B35DD810B0D1E2685AF1AFCB7FBF3" box="[676,825,1124,1149]" class="Insecta" family="Eriococcidae" genus="Eriococcus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="B92F924C810B0D1E2685AF1AFCB7FBF3" box="[676,825,1124,1149]" italics="true" pageId="32" pageNumber="33">A. paradoxus</emphasis>
</taxonomicName>
from
<collectingCountry id="F34C0ECE810B0D1E275FAF1AFC67FBF3" box="[894,1001,1124,1149]" name="Australia" pageId="32" pageNumber="33">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>