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<document id="0CE231E27B323AD27B2AE0D8E508E2AA" ID-CLB-Dataset="2767" ID-DOI="10.11646/zootaxa.5087.3.2" ID-GBIF-Dataset="43c1b1e7-1797-4ed7-817d-606114b807ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="5826858" ID-ZooBank="6253FEED-A4FB-4832-AE51-91E4F51655C4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641539129469" checkinUser="plazi" docAuthor="Paz, Fernando, Ratcliffe, Brett C. &amp; Figueroa, Luis" docDate="2022" docId="E61687C1FFA0FFD67BB4B9FDFDC219DB" docLanguage="en" docName="zootaxa.5087.3.2.pdf" docOrigin="Zootaxa 5087 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyclocephala ukuku Paz &amp; Ratcliffe 2022, new species" docType="treatment" docUuid="45080894-FAD8-4952-B444-9F4791C49B2A" docUuidSource="ZooBank" docVersion="6" lastPageNumber="430" masterDocId="1A2FFFB9FFA1FFD57B23BF3BFFAE1A5A" masterDocTitle="Three new species of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Amazonian Peru and a checklist of Cyclocephala species in Peru" masterLastPageNumber="440" masterPageNumber="427" pageNumber="428" updateTime="1699296570230" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E2E48AE798981F076C36FB663FBBC02C">Three new species of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Amazonian Peru and a checklist of Cyclocephala species in Peru</mods:title>
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<mods:name id="15CB57E6C39BA5A5826DE5504E929C87" type="personal">
<mods:role id="0ABAA2D32178CD26E869139C03FD482F">
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<mods:namePart id="74F48E11C1C20DEF025F55688418B835">Paz, Fernando</mods:namePart>
<mods:nameIdentifier id="0F20F610FF9CD01E6C1D0415D2CEA417" type="ORCID">0000-0002-6331-4416</mods:nameIdentifier>
<mods:affiliation id="A6A5C8D03E1856E789936D2E44044D9A">Departamento de Entomología, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Apartado Postal 14 - 0434, Lima, Peru. fernandopaz 683 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6331 - 4416</mods:affiliation>
<mods:nameIdentifier id="9589AB778721D1ACA48C4B634099D2E1" type="email">fernandopaz683@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="706B7A53ED3CA85D845D474DD265196A" type="personal">
<mods:role id="F70140447F9FAF9EFCB8D184FABBAC92">
<mods:roleTerm id="31CC99CD5988F8DB49AE3B687B44F711">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6366352D2C28803A856B3B83D3C00E03">Ratcliffe, Brett C.</mods:namePart>
<mods:nameIdentifier id="024932027AB58D83B6A3EFD35343CB39" type="ORCID">0000-0002-0589-8791</mods:nameIdentifier>
<mods:affiliation id="05FE1C716076587B30B6BD647FF023E5">Systematics Research Collections, University of Nebraska State Museum, University of Nebraska, Lincoln, Nebraska, 68588 - 0338, United States of America. bratcliffe 1 @ unl. edu; http: // orcid. org / 0000 - 0002 - 0589 - 8791</mods:affiliation>
<mods:nameIdentifier id="710320CA20D96459410D2207C4617FE8" type="email">bratcliffe1@unl.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="26DADE21F1EC11326F3BAA09462B554F" type="personal">
<mods:role id="CC16EF5E69B61593E7CDC137053672C4">
<mods:roleTerm id="A457DBB4B765EDAE03E803A4E7D03301">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2086A07E593FD0C9421E07E823BC4A92">Figueroa, Luis</mods:namePart>
<mods:affiliation id="15C76FFC50C268CC6F0F59CC88A20060">Departamento de Entomología, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Lima, Peru.</mods:affiliation>
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<mods:typeOfResource id="63E3916BEE725B292EC381B08C9868EC">text</mods:typeOfResource>
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<mods:title id="0681549997737FC16B4BF8403A6246A2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FD505DCCD79941B261DF10E9E48F7D5E">
<mods:date id="DD1B494A965B9B1D0F4AF3171A2F600F">2022</mods:date>
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<mods:number id="CB94F39821C9CB7439837AB276D1C019">2022-01-07</mods:number>
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<mods:number id="D3CD598ADDDB45681C580D706D246F7C">5087</mods:number>
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<mods:classification id="EF8878D95004C1F2806D1A4C03374F70">journal article</mods:classification>
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<treatment id="E61687C1FFA0FFD67BB4B9FDFDC219DB" ID-DOI="http://doi.org/10.5281/zenodo.5826862" ID-GBIF-Taxon="191928392" ID-Zenodo-Dep="5826862" LSID="urn:lsid:zoobank.org:act:45080894-FAD8-4952-B444-9F4791C49B2A" httpUri="http://treatment.plazi.org/id/E61687C1FFA0FFD67BB4B9FDFDC219DB" lastPageId="3" lastPageNumber="430" pageId="1" pageNumber="428">
<subSubSection id="26A5655CFFA0FFD47BB4B9FDFD581CBA" box="[151,758,1734,1761]" pageId="1" pageNumber="428" type="nomenclature">
<paragraph id="6E0036D7FFA0FFD47BB4B9FDFD581CBA" blockId="1.[151,758,1734,1797]" box="[151,758,1734,1761]" pageId="1" pageNumber="428">
<heading id="354881BBFFA0FFD47BB4B9FDFD581CBA" bold="true" box="[151,758,1734,1761]" fontSize="11" level="1" pageId="1" pageNumber="428" reason="1">
<emphasis id="5CCBEAC5FFA0FFD47BB4B9FDFD581CBA" bold="true" box="[151,758,1734,1761]" pageId="1" pageNumber="428">
<taxonomicName id="A9BF4D54FFA0FFD47BB4B9FDFDF71CBA" authority="Paz &amp; Ratcliffe" authorityName="Paz &amp; Ratcliffe" authorityYear="2022" box="[151,601,1734,1761]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="ukuku" status="sp. nov.">
<emphasis id="5CCBEAC5FFA0FFD47BB4B9FDFE211CBB" bold="true" box="[151,399,1734,1761]" italics="true" pageId="1" pageNumber="428">Cyclocephala ukuku</emphasis>
Paz &amp; Ratcliffe
</taxonomicName>
,
<taxonomicNameLabel id="47F857BEFFA0FFD47945B9FCFD581CBA" box="[614,758,1734,1761]" pageId="1" pageNumber="428" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="26A5655CFFA0FFD47BB4B9D0FEC81D5F" box="[151,358,1771,1797]" pageId="1" pageNumber="428" type="description">
<paragraph id="6E0036D7FFA0FFD47BB4B9D0FEC81D5F" blockId="1.[151,758,1734,1797]" box="[151,358,1771,1797]" pageId="1" pageNumber="428">
(
<figureCitation id="F6842A52FFA0FFD47BBCB9D0FEA51D5F" box="[159,267,1771,1797]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1587,1611]" captionTargetBox="[157,1429,185,1559]" captionTargetId="figure-23@2.[152,1435,179,1560]" captionTargetPageId="2" captionText="FIGURES 1–6. Cyclocephala ukuku. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="1" pageNumber="428">Figs. 1–6</figureCitation>
,
<figureCitation id="F6842A52FFA0FFD47A34B9D0FEF01D5F" box="[279,350,1771,1797]" captionStart-0="FIGURE 20" captionStart-1="FIGURE 21" captionStartId-0="7.[151,250,1943,1967]" captionStartId-1="8.[151,250,1521,1545]" captionTargetBox-0="[198,1390,181,1919]" captionTargetBox-1="[155,1432,561,1498]" captionTargetId-0="figure-18@7.[198,1390,181,1920]" captionTargetId-1="figure-204@8.[155,1432,561,1498]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 20. Distribution map of the new species of Peruvian Cyclocephala." captionText-1="FIGURE 21. Aerial view of the habitat of the holotype of Cyclocephala ukuku at Reserva Comunal Machiguenga, Cusco, Peru. Photograph by Juan Grados." figureDoi-0="http://doi.org/10.5281/zenodo.5826870" figureDoi-1="http://doi.org/10.5281/zenodo.5826872" httpUri-0="https://zenodo.org/record/5826870/files/figure.png" httpUri-1="https://zenodo.org/record/5826872/files/figure.png" pageId="1" pageNumber="428">20–21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="26A5655CFFA0FFD47BB4B808FB2D1D87" pageId="1" pageNumber="428" type="materials_examined">
<paragraph id="6E0036D7FFA0FFD47BB4B808FB2D1D87" blockId="1.[151,1437,1842,2013]" pageId="1" pageNumber="428">
<emphasis id="5CCBEAC5FFA0FFD47BB4B808FEE71D16" bold="true" box="[151,329,1842,1869]" pageId="1" pageNumber="428">Type material:</emphasis>
<materialsCitation id="DED73C8AFFA0FFD47A71B808FAF91D2B" ID-GBIF-Occurrence="3427663320" collectingDate="2010-01-18" collectorName="J. Grados" country="Peru" county="Mundo" elevation="741" location="Machiguenga" municipality="Km" pageId="1" pageNumber="428" specimenCount="741" typeStatus="holotype">
<typeStatus id="B1048875FFA0FFD47A71B808FE141D17" box="[338,442,1843,1869]" pageId="1" pageNumber="428" type="holotype">Holotype</typeStatus>
male labeled: “
<collectingCountry id="16A87647FFA0FFD4794DB808FD151D17" box="[622,699,1843,1869]" name="Peru" pageId="1" pageNumber="428">PERU</collectingCountry>
, CU, RC /
<location id="6B60600CFFA0FFD4781DB808FC771D17" LSID="urn:lsid:plazi:treatment:E61687C1FFA0FFD67BB4B9FDFDC219DB:6B60600CFFA0FFD4781DB808FC771D17" box="[830,985,1843,1869]" country="Peru" county="Mundo" municipality="Km" name="Machiguenga" pageId="1" pageNumber="428">Machiguenga</location>
,
<location id="6B60600CFFA0FFD478C4B808FBEE1D17" LSID="urn:lsid:plazi:treatment:E61687C1FFA0FFD67BB4B9FDFDC219DB:6B60600CFFA0FFD478C4B808FBEE1D17" box="[999,1088,1843,1869]" country="Peru" county="Mundo" municipality="Km" name="Campto" pageId="1" pageNumber="428">Campto</location>
/
<location id="6B60600CFFA0FFD47F7AB808FB391D17" LSID="urn:lsid:plazi:treatment:E61687C1FFA0FFD67BB4B9FDFDC219DB:6B60600CFFA0FFD47F7AB808FB391D17" box="[1113,1175,1843,1869]" country="Peru" county="Mundo" municipality="Km" name="Mapi" pageId="1" pageNumber="428">Mapi</location>
, 35.5
<collectingMunicipality id="8E64ACADFFA0FFD47FFDB808FAA61D17" box="[1246,1288,1843,1869]" pageId="1" pageNumber="428">Km</collectingMunicipality>
<locationDeviation id="7FB22FF5FFA0FFD47E32B808FA321D17" box="[1297,1436,1843,1869]" location="Nuevo" pageId="1" pageNumber="428">O de Nuevo</locationDeviation>
/
<collectingCounty id="87614E5BFFA0FFD47B84B86CFF551D2B" box="[167,251,1879,1905]" pageId="1" pageNumber="428">Mundo</collectingCounty>
,
<specimenCount id="78B9FD5EFFA0FFD47A2BB86CFEF91D2B" box="[264,343,1879,1905]" pageId="1" pageNumber="428" type="generic">
<quantity id="A9479B32FFA0FFD47A2BB86CFEFD1D2B" box="[264,339,1879,1905]" metricMagnitude="2" metricUnit="m" metricValue="7.409999999999999" pageId="1" pageNumber="428" unit="m" value="741.0">
<elevation id="E592D1E4FFA0FFD47A2BB86CFEFD1D2B" box="[264,339,1879,1905]" metricMagnitude="2" metricUnit="m" metricValue="7.409999999999999" pageId="1" pageNumber="428" unit="m" value="741.0">741 m</elevation>
</quantity>
.
</specimenCount>
73° / 28'30,46&quot;/ 11°31'30,24&quot;, 13- /
<date id="1A011017FFA0FFD479D8B86CFCCB1D2B" box="[763,869,1879,1905]" pageId="1" pageNumber="428" value="2010-01-18">
<collectingDate id="0A45E9FFFFA0FFD479D8B86CFCCB1D2B" box="[763,869,1879,1905]" pageId="1" pageNumber="428" value="2010-01-18">18.i.2010</collectingDate>
</date>
<collectorName id="C34A5301FFA0FFD4784EB86CFC751D2B" box="[877,987,1879,1905]" pageId="1" pageNumber="428">J. Grados</collectorName>
” and with our red
<typeStatus id="B1048875FFA0FFD47F91B86CFABA1D2B" box="[1202,1300,1879,1905]" pageId="1" pageNumber="428" type="holotype">holotype</typeStatus>
label
</materialsCitation>
.
<materialsCitation id="DED73C8AFFA0FFD47E42B86CFF4F1D87" ID-GBIF-Occurrence="3427663321" collectingDate="2010-09-27" collectorName="M. Alvarado &amp; J. Peralta" country="Peru" county="Mundo" elevation="1708" latitude="-12.571834" location="Zona" longLatPrecision="1" longitude="-73.087" municipality="Megantoni" pageId="1" pageNumber="428" specimenCount="3" typeStatus="allotype">
<typeStatus id="B1048875FFA0FFD47E42B86CFF691DCF" pageId="1" pageNumber="428" type="allotype">Allotype</typeStatus>
female labeled: “
<collectingCountry id="16A87647FFA0FFD47AB3B840FE731DCF" box="[400,477,1915,1941]" name="Peru" pageId="1" pageNumber="428">PERU</collectingCountry>
, CU.
<location id="6B60600CFFA0FFD47902B840FDF51DCF" LSID="urn:lsid:plazi:treatment:E61687C1FFA0FFD67BB4B9FDFDC219DB:6B60600CFFA0FFD47902B840FDF51DCF" box="[545,603,1915,1941]" country="Peru" latitude="-12.571834" longLatPrecision="1" longitude="-73.087" municipality="Megantoni" name="Zona" pageId="1" pageNumber="428">Zona</location>
de / amortiguamiento /
<collectingMunicipality id="8E64ACADFFA0FFD47853B840FC471DCF" box="[880,1001,1915,1941]" pageId="1" pageNumber="428">Megantoni</collectingMunicipality>
<geoCoordinate id="0B8B5010FFA0FFD478D1B840FB2B1DCF" box="[1010,1157,1915,1941]" degrees="12" direction="south" minutes="34" orientation="latitude" pageId="1" pageNumber="428" precision="1" seconds="18.6" value="-12.571834">12°34'18.6&quot;S</geoCoordinate>
/
<geoCoordinate id="0B8B5010FFA0FFD47FB4B841FA981DCF" box="[1175,1334,1914,1941]" degrees="73" direction="west" minutes="5" orientation="longitude" pageId="1" pageNumber="428" precision="1" seconds="13.18" value="-73.087">73°5'13.18&quot;W</geoCoordinate>
<quantity id="A9479B32FFA0FFD47E1CB840FA361DCF" box="[1343,1432,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="1.708" pageId="1" pageNumber="428" unit="m" value="1708.0">
<elevation id="E592D1E4FFA0FFD47E1CB840FA361DCF" box="[1343,1432,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="1.708" pageId="1" pageNumber="428" unit="m" value="1708.0">1708 m</elevation>
</quantity>
. /
<date id="1A011017FFA0FFD47B84B8A4FEB01DE3" box="[167,286,1951,1977]" pageId="1" pageNumber="428" value="2010-09-27">
<collectingDate id="0A45E9FFFFA0FFD47B84B8A4FEB01DE3" box="[167,286,1951,1977]" pageId="1" pageNumber="428" value="2010-09-27">27.ix.2010</collectingDate>
</date>
<collectorName id="C34A5301FFA0FFD47A05B8A4FE1A1DE3" box="[294,436,1951,1977]" pageId="1" pageNumber="428">M. Alvarado</collectorName>
y /
<collectorName id="C34A5301FFA0FFD47AC2B8A4FDE21DE3" box="[481,588,1951,1977]" pageId="1" pageNumber="428">J. Peralta</collectorName>
” and with our red
<typeStatus id="B1048875FFA0FFD4783DB8A4FCD61DE3" box="[798,888,1951,1977]" pageId="1" pageNumber="428" type="allotype">allotype</typeStatus>
label. Two
<typeStatus id="B1048875FFA0FFD478DDB8A4FBF01DE3" box="[1022,1118,1951,1977]" pageId="1" pageNumber="428" type="paratype">paratype</typeStatus>
males with same data as allotype
</materialsCitation>
.
<materialsCitation id="DED73C8AFFA0FFD47BC8B8F8FCF31D87" ID-GBIF-Occurrence="3427663329" box="[235,861,1987,2013]" collectionCode="MUSM" country="Peru" pageId="1" pageNumber="428" specimenCount="1" typeStatus="holotype">
<typeStatus id="B1048875FFA0FFD47BC8B8F8FEF81D87" box="[235,342,1987,2013]" pageId="1" pageNumber="428" type="holotype">Holotype</typeStatus>
,
<typeStatus id="B1048875FFA0FFD47A42B8F8FE101D87" box="[353,446,1987,2013]" pageId="1" pageNumber="428" type="allotype">allotype</typeStatus>
, and
<specimenCount id="78B9FD5EFFA0FFD47ADAB8F8FDC01D87" box="[505,622,1987,2013]" pageId="1" pageNumber="428" type="generic" typeStatus="paratype">1 paratype</specimenCount>
deposited in
<collectionCode id="08AEAE12FFA0FFD47826B8F8FCF31D87" box="[773,861,1987,2013]" pageId="1" pageNumber="428">MUSM</collectionCode>
</materialsCitation>
,
<materialsCitation id="DED73C8AFFA0FFD4784BB8F8FBD11D87" ID-GBIF-Occurrence="3427663322" box="[872,1151,1987,2013]" collectionCode="BCRC" country="Peru" pageId="1" pageNumber="428" specimenCount="1" typeStatus="holotype">
and
<specimenCount id="78B9FD5EFFA0FFD478BBB8F8FBA31D87" box="[920,1037,1987,2013]" pageId="1" pageNumber="428" type="generic" typeStatus="paratype">1 paratype</specimenCount>
in
<collectionCode id="08AEAE12FFA0FFD47F12B8F8FBD11D87" box="[1073,1151,1987,2013]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="1" pageNumber="428">BCRC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="3AC0665FFFA3FFD77BB4B908FCFC1C34" ID-DOI="http://doi.org/10.5281/zenodo.5826860" ID-Zenodo-Dep="5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="2" pageNumber="429" startId="2.[151,264,1587,1611]" targetBox="[157,1429,185,1559]" targetPageId="2">
<paragraph id="6E0036D7FFA3FFD77BB4B908FCFC1C34" blockId="2.[151,1436,1587,1647]" pageId="2" pageNumber="429">
<emphasis id="5CCBEAC5FFA3FFD77BB4B908FE921C11" bold="true" box="[151,316,1587,1611]" pageId="2" pageNumber="429">FIGURES 1–6.</emphasis>
<taxonomicName id="A9BF4D54FFA3FFD77A67B908FDBA1C11" box="[324,532,1587,1611]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="species" species="ukuku">
<emphasis id="5CCBEAC5FFA3FFD77A67B908FDBA1C11" box="[324,532,1587,1611]" italics="true" pageId="2" pageNumber="429">Cyclocephala ukuku</emphasis>
</taxonomicName>
. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm.
</paragraph>
</caption>
<subSubSection id="26A5655CFFA3FFD67BE4B9A6FE0E1BF7" lastPageId="3" lastPageNumber="430" pageId="2" pageNumber="429" type="description">
<paragraph id="6E0036D7FFA3FFD67BE4B9A6FB311A8F" blockId="2.[151,1437,1693,2044]" lastBlockId="3.[151,1437,151,897]" lastPageId="3" lastPageNumber="430" pageId="2" pageNumber="429">
<emphasis id="5CCBEAC5FFA3FFD77BE4B9A6FE7A1CED" bold="true" box="[199,468,1693,1719]" pageId="2" pageNumber="429">
Description.
<typeStatus id="B1048875FFA3FFD77A43B9A6FE611CED" box="[352,463,1693,1719]" pageId="2" pageNumber="429" type="holotype">Holotype</typeStatus>
,
</emphasis>
male. (
<figureCitation id="F6842A52FFA3FFD77909B9A6FDC21CED" box="[554,620,1693,1719]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1587,1611]" captionTargetBox="[157,1429,185,1559]" captionTargetId="figure-23@2.[152,1435,179,1560]" captionTargetPageId="2" captionText="FIGURES 1–6. Cyclocephala ukuku. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="2" pageNumber="429">Fig. 1</figureCitation>
). Length
<quantity id="A9479B32FFA3FFD779FFB9A6FCEA1CE2" box="[732,836,1693,1720]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="2" pageNumber="429" unit="mm" value="18.5">18.5 mm</quantity>
; width
<quantity id="A9479B32FFA3FFD778BBB9A6FC5C1CE2" box="[920,1010,1693,1720]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="2" pageNumber="429" unit="mm" value="9.1">9.1 mm</quantity>
. Color dark reddish brown, shiny, except for black tarsomeres and black on posterior half of frons.
<emphasis id="5CCBEAC5FFA3FFD7787DB9FAFC091C81" bold="true" box="[862,935,1729,1755]" pageId="2" pageNumber="429">Head:</emphasis>
Frons and clypeus with punctures moderate in size and density, setigerous; setae minute, tawny. Clypeal apex broadly subtruncate, slightly reflexed. Interocular width equals 3.1 transverse eye diameters. Antenna with 10 antennomeres, club subequal in length to antennomeres 2–7.
<emphasis id="5CCBEAC5FFA3FFD77BF3B816FEFD1D1D" bold="true" box="[208,339,1837,1863]" pageId="2" pageNumber="429">Pronotum:</emphasis>
Surface similar to that of frons; setae short, tawny. Base lacking marginal bead.
<emphasis id="5CCBEAC5FFA3FFD77FCAB816FA901D1D" bold="true" box="[1257,1342,1837,1863]" pageId="2" pageNumber="429">Elytra:</emphasis>
Surface weakly punctate-striate, transversely wrinkled, punctures slightly larger and denser than those of pronotum, setigerous; setae short, tawny.
<emphasis id="5CCBEAC5FFA3FFD77A9EB84EFD971DD5" bold="true" box="[445,569,1909,1935]" pageId="2" pageNumber="429">Pygidium:</emphasis>
Surface with large, dense, setigerous punctures; setae long, tawny. In lateral view, surface weakly convex.
<emphasis id="5CCBEAC5FFA3FFD77ACBB8A2FD841DE9" bold="true" box="[488,554,1945,1971]" pageId="2" pageNumber="429">Legs:</emphasis>
Protibia slender, tridentate, basal tooth reduced to a rounded prominence. Protarsomere 5 enlarged, slightly bent, venter concave; medial claw large, strongly bent, apex split (
<figureCitation id="F6842A52FFA3FFD77FEAB886FAA31D82" box="[1225,1293,1981,2008]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1587,1611]" captionTargetBox="[157,1429,185,1559]" captionTargetId="figure-23@2.[152,1435,179,1560]" captionTargetPageId="2" captionText="FIGURES 1–6. Cyclocephala ukuku. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="2" pageNumber="429">Fig. 5</figureCitation>
). Metatarsus slightly shorter than metatibia.
<emphasis id="5CCBEAC5FFA3FFD77ACCB8DAFDE61DA1" bold="true" box="[495,584,2017,2043]" pageId="2" pageNumber="429">Venter:</emphasis>
Prosternal process long, columnar, apex obliquely flattened into transversely oval disc with transverse, elevated “button” on anterior half.
<emphasis id="5CCBEAC5FFA2FFD67815BFACFC6E1AEB" bold="true" box="[822,960,151,177]" pageId="3" pageNumber="430">Parameres:</emphasis>
In caudal view, form elongate, subparallel, apices rounded (
<figureCitation id="F6842A52FFA2FFD67A6CBF80FE3A1A8F" box="[335,404,187,213]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1587,1611]" captionTargetBox="[157,1429,185,1559]" captionTargetId="figure-23@2.[152,1435,179,1560]" captionTargetPageId="2" captionText="FIGURES 1–6. Cyclocephala ukuku. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="3" pageNumber="430">Fig. 3</figureCitation>
). In lateral view, basal piece subequal in length to parameres (
<figureCitation id="F6842A52FFA2FFD67F6EBF80FB3C1A8F" box="[1101,1170,187,213]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1587,1611]" captionTargetBox="[157,1429,185,1559]" captionTargetId="figure-23@2.[152,1435,179,1560]" captionTargetPageId="2" captionText="FIGURES 1–6. Cyclocephala ukuku. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="3" pageNumber="430">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="6E0036D7FFA2FFD67BE4BFE5FC961B3F" blockId="3.[151,1437,151,897]" pageId="3" pageNumber="430">
<emphasis id="5CCBEAC5FFA2FFD67BE4BFE5FE9D1AA2" bold="true" box="[199,307,222,248]" pageId="3" pageNumber="430">
<typeStatus id="B1048875FFA2FFD67BE4BFE5FE801AA2" box="[199,302,222,248]" pageId="3" pageNumber="430" type="allotype">Allotype</typeStatus>
,
</emphasis>
female (
<figureCitation id="F6842A52FFA2FFD67AB5BFE4FE721AA3" box="[406,476,223,249]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1587,1611]" captionTargetBox="[157,1429,185,1559]" captionTargetId="figure-23@2.[152,1435,179,1560]" captionTargetPageId="2" captionText="FIGURES 1–6. Cyclocephala ukuku. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="3" pageNumber="430">Fig. 2</figureCitation>
). Length
<quantity id="A9479B32FFA2FFD6796BBFE4FD011AA3" box="[584,687,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="3" pageNumber="430" unit="mm" value="19.3">19.3 mm</quantity>
; width across humeri 9.0 mm. As
<typeStatus id="B1048875FFA2FFD67F0CBFE4FB3F1AA3" box="[1071,1169,223,249]" pageId="3" pageNumber="430" type="holotype">holotype</typeStatus>
except in the following respects:
<emphasis id="5CCBEAC5FFA2FFD67A22BE38FEF91B47" bold="true" box="[257,343,259,285]" pageId="3" pageNumber="430">Elytra:</emphasis>
Epipleuron (ventral view) gradually tapering beginning at level of abdominal sternite 2 (
<figureCitation id="F6842A52FFA2FFD67E6ABE38FA201B44" box="[1353,1422,259,286]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1587,1611]" captionTargetBox="[157,1429,185,1559]" captionTargetId="figure-23@2.[152,1435,179,1560]" captionTargetPageId="2" captionText="FIGURES 1–6. Cyclocephala ukuku. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="3" pageNumber="430">Fig. 6</figureCitation>
). In dorsal view, lateral margin simple, evenly rounded.
<emphasis id="5CCBEAC5FFA2FFD6782EBE1DFC2E1B1A" bold="true" box="[781,896,294,320]" pageId="3" pageNumber="430">Pygidium</emphasis>
: In lateral view, surface weakly convex.
<emphasis id="5CCBEAC5FFA2FFD67E78BE1CFA321B1B" bold="true" box="[1371,1436,295,321]" pageId="3" pageNumber="430">Legs:</emphasis>
Protibia distinctly tridentate. Protarsus simple, not enlarged.
</paragraph>
<paragraph id="6E0036D7FFA2FFD67BE4BE55FE0E1BF7" blockId="3.[151,1437,151,897]" pageId="3" pageNumber="430">
<emphasis id="5CCBEAC5FFA2FFD67BE4BE55FE911BD2" bold="true" box="[199,319,366,392]" pageId="3" pageNumber="430">Variation.</emphasis>
Males (
<specimenCount id="78B9FD5EFFA2FFD67ABBBE54FDB01BD3" box="[408,542,367,393]" pageId="3" pageNumber="430" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
). Length 18.0–
<quantity id="A9479B32FFA2FFD679E7BE55FC871BD3" box="[708,809,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="3" pageNumber="430" unit="mm" value="19.5">19.5 mm</quantity>
; width 8.5–9.0 mm. The
<typeStatus id="B1048875FFA2FFD67F1EBE54FB061BD3" box="[1085,1192,367,393]" pageId="3" pageNumber="430" type="paratype">paratypes</typeStatus>
do not differ from the
<typeStatus id="B1048875FFA2FFD67BB4BEA8FF571BF7" box="[151,249,403,429]" pageId="3" pageNumber="430" type="holotype">holotype</typeStatus>
except in size.
</paragraph>
</subSubSection>
<subSubSection id="26A5655CFFA2FFD67BE4BE8CFDC91843" pageId="3" pageNumber="430" type="etymology">
<paragraph id="6E0036D7FFA2FFD67BE4BE8CFDC91843" blockId="3.[151,1437,151,897]" pageId="3" pageNumber="430">
<emphasis id="5CCBEAC5FFA2FFD67BE4BE8CFEE01B8B" bold="true" box="[199,334,439,465]" pageId="3" pageNumber="430">Etymology.</emphasis>
<taxonomicName id="A9BF4D54FFA2FFD67A76BE8CFD941B8B" box="[341,570,439,465]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="ukuku">
<emphasis id="5CCBEAC5FFA2FFD67A76BE8CFD941B8B" box="[341,570,439,465]" italics="true" pageId="3" pageNumber="430">Cyclocephala ukuku</emphasis>
</taxonomicName>
is from Peruvian mythology and refers to a being from the mountains that is half bear and half human. The creature is known for being a dancer and guardian of the snowy mountains. This name should be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="26A5655CFFA2FFD67BE4BD19FCBE18DF" pageId="3" pageNumber="430" type="distribution">
<paragraph id="6E0036D7FFA2FFD67BE4BD19FBD4183B" blockId="3.[151,1437,151,897]" pageId="3" pageNumber="430">
<emphasis id="5CCBEAC5FFA2FFD67BE4BD19FECF1866" bold="true" box="[199,353,546,572]" pageId="3" pageNumber="430">Distribution.</emphasis>
<specimenCount id="78B9FD5EFFA2FFD67A4BBD18FE5D1867" box="[360,499,547,573]" pageId="3" pageNumber="430" type="generic">4 specimens</specimenCount>
examined (
<figureCitation id="F6842A52FFA2FFD6795BBD18FD631867" box="[632,717,547,573]" captionStart="FIGURE 20" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[198,1390,181,1919]" captionTargetId="figure-18@7.[198,1390,181,1920]" captionTargetPageId="7" captionText="FIGURE 20. Distribution map of the new species of Peruvian Cyclocephala." figureDoi="http://doi.org/10.5281/zenodo.5826870" httpUri="https://zenodo.org/record/5826870/files/figure.png" pageId="3" pageNumber="430">Fig. 20</figureCitation>
).
<collectingCountry id="16A87647FFA2FFD679C2BD18FCB61860" box="[737,792,547,573]" name="Peru" pageId="3" pageNumber="430">PERU</collectingCountry>
(4):
<collectingRegion id="AC7BF835FFA2FFD67873BD18FC031867" box="[848,941,547,573]" country="Peru" name="Cusco" pageId="3" pageNumber="430">CUSCO</collectingRegion>
(4): Reserva Comunal Machiguenga, Campamento Mapi,
<quantity id="A9479B32FFA2FFD67A6BBD7DFE0A183B" box="[328,420,582,609]" metricMagnitude="4" metricUnit="m" metricValue="3.55" pageId="3" pageNumber="430" unit="km" value="35.5">35.5 km</quantity>
W Nuevo Mundo (1); Zona de Amortiguamiento Megantoni (3).
</paragraph>
<paragraph id="6E0036D7FFA2FFD67BE4BD50FCBE18DF" blockId="3.[151,1437,151,897]" box="[199,784,618,645]" pageId="3" pageNumber="430">
<emphasis id="5CCBEAC5FFA2FFD67BE4BD50FE7918DE" bold="true" box="[199,471,618,645]" pageId="3" pageNumber="430">Temporal distribution.</emphasis>
January (1), September (3).
</paragraph>
</subSubSection>
<subSubSection id="26A5655CFFA2FFD67BE4BDB5FDC219DB" pageId="3" pageNumber="430" type="diagnosis">
<paragraph id="6E0036D7FFA2FFD67BE4BDB5FF1B1960" blockId="3.[151,1437,151,897]" pageId="3" pageNumber="430">
<emphasis id="5CCBEAC5FFA2FFD67BE4BDB5FEEC18F2" bold="true" box="[199,322,654,680]" pageId="3" pageNumber="430">Diagnosis.</emphasis>
<taxonomicName id="A9BF4D54FFA2FFD67A6ABDB4FD8018F3" box="[329,558,655,681]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="ukuku">
<emphasis id="5CCBEAC5FFA2FFD67A6ABDB4FD8018F3" box="[329,558,655,681]" italics="true" pageId="3" pageNumber="430">Cyclocephala ukuku</emphasis>
</taxonomicName>
is characterized by a distinctive, shiny, dark reddish-brown color (
<figureCitation id="F6842A52FFA2FFD67E02BDB4FA2018F3" box="[1313,1422,655,681]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1587,1611]" captionTargetBox="[157,1429,185,1559]" captionTargetId="figure-23@2.[152,1435,179,1560]" captionTargetPageId="2" captionText="FIGURES 1–6. Cyclocephala ukuku. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="3" pageNumber="430">Figs. 1–2</figureCitation>
); presence of small, abundant setae on the head, pronotum, and elytra; broadly subtruncate clypeus with a weakly emarginate apex; basal bead of the pronotum lacking; tridentate protibia; male large protarsal claw split at its apex (
<figureCitation id="F6842A52FFA2FFD67B83BDC0FF4B194F" box="[160,229,762,789]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1587,1611]" captionTargetBox="[157,1429,185,1559]" captionTargetId="figure-23@2.[152,1435,179,1560]" captionTargetPageId="2" captionText="FIGURES 1–6. Cyclocephala ukuku. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="3" pageNumber="430">Fig. 5</figureCitation>
); and uniquely shaped parameres (
<figureCitation id="F6842A52FFA2FFD6794EBDC0FD75194F" box="[621,731,763,789]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1587,1611]" captionTargetBox="[157,1429,185,1559]" captionTargetId="figure-23@2.[152,1435,179,1560]" captionTargetPageId="2" captionText="FIGURES 1–6. Cyclocephala ukuku. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="3" pageNumber="430">Figs. 3–4</figureCitation>
). The female epipleuron is simple and gradually tapering (
<figureCitation id="F6842A52FFA2FFD67E4CBDC0FF091960" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1587,1611]" captionTargetBox="[157,1429,185,1559]" captionTargetId="figure-23@2.[152,1435,179,1560]" captionTargetPageId="2" captionText="FIGURES 1–6. Cyclocephala ukuku. 1, Male holotype; 2, female allotype; 3–4, parameres of holotype; 5, protarsal claw of holotype; 6, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826860" httpUri="https://zenodo.org/record/5826860/files/figure.png" pageId="3" pageNumber="430">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="6E0036D7FFA2FFD67BE4BC78FDC219DB" blockId="3.[151,1437,151,897]" pageId="3" pageNumber="430">
<emphasis id="5CCBEAC5FFA2FFD67BE4BC78FE281906" bold="true" box="[199,390,834,861]" pageId="3" pageNumber="430">Natural history.</emphasis>
Specimens were collected at elevations of
<quantity id="A9479B32FFA2FFD6784FBC78FC1D1907" box="[876,947,835,861]" metricMagnitude="2" metricUnit="m" metricValue="7.409999999999999" pageId="3" pageNumber="430" unit="m" value="741.0">741 m</quantity>
and
<quantity id="A9479B32FFA2FFD678C8BC78FBEE1907" box="[1003,1088,835,861]" metricMagnitude="3" metricUnit="m" metricValue="1.708" pageId="3" pageNumber="430" unit="m" value="1708.0">1708 m</quantity>
in pristine forest (
<figureCitation id="F6842A52FFA2FFD67E33BC78FACD1907" box="[1296,1379,835,861]" captionStart="FIGURE 21" captionStartId="8.[151,250,1521,1545]" captionTargetBox="[155,1432,561,1498]" captionTargetId="figure-204@8.[155,1432,561,1498]" captionTargetPageId="8" captionText="FIGURE 21. Aerial view of the habitat of the holotype of Cyclocephala ukuku at Reserva Comunal Machiguenga, Cusco, Peru. Photograph by Juan Grados." figureDoi="http://doi.org/10.5281/zenodo.5826872" httpUri="https://zenodo.org/record/5826872/files/figure.png" pageId="3" pageNumber="430">Fig. 21</figureCitation>
), but nothing else is known of their life history.
</paragraph>
</subSubSection>
</treatment>
</document>
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