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<document id="588C12489BD3B9257F3472A3AC3F5C94" ID-DOI="10.3897/zoologia.35.e21232" ID-ISSN="1984-4689" ID-Zenodo-Dep="13174064" ID-ZooBank="663F033F-E804-444F-A999-9F42993FF545" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722587231085" checkinUser="felipe" docAuthor="Garbelotto, Thereza de Almeida, Kochenborger, Ana Paula Leite &amp; Campos, Luiz Alexandre" docDate="2018" docId="AB1D87D9FF9F0117FF24F2FCFDC36A9C" docLanguage="en" docName="Zoologia.35.e21232.pdf" docOrigin="Zoologia (e 21232) 35" docSource="http://dx.doi.org/10.3897/zoologia.35.e21232" docStyle="DocumentStyle:9D87517C92C8E430512793D922E147D4.1:Zoologia.2017-2020.journal_article" docStyleId="9D87517C92C8E430512793D922E147D4" docStyleName="Zoologia.2017-2020.journal_article" docStyleVersion="1" docTitle="Lineostethus otarus Garbelotto &amp; Kochenborger &amp; Campos 2018, sp. nov." docType="treatment" docUuid="D6A205DB-7ECB-4C93-B813-2515E45876AA" docUuidSource="ZooBank" docVersion="2" lastPageNumber="21" masterDocId="5724FFA1FF8D0103FF96F222FFF66B55" masterDocTitle="Revision of Lineostethus (Heteroptera: Pentatomidae: Discocephalini)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="19" updateTime="1722619988372" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="095BB7E9BD29E452A648D27AD7CE211F">Revision of Lineostethus (Heteroptera: Pentatomidae: Discocephalini)</mods:title>
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<mods:namePart id="C0EFD9A9984ACAFDB7D65636401174AB">Garbelotto, Thereza de Almeida</mods:namePart>
<mods:affiliation id="38432B8B56F263E0AC29735CAC9CB0CF">Departamento de Ciências Biológicas, Universidade do Sul Catarinense. Avenida José Acácio Moreira 787, 88704 - 900 Tubarão, SC, Brazil.</mods:affiliation>
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<mods:namePart id="B7533D4924D8142E1B89024672279FEF">Kochenborger, Ana Paula Leite</mods:namePart>
<mods:affiliation id="34A657744BBF41F0674BAE9CFACF239B">Departamento de Zoologia, Universidade Federal do Rio Grande do Sul. Avenida Bento Gonçalves 9500, 91501 - 970 Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:namePart id="9D2F0CDFEDA31C87CF9151053FFB0BB5">Campos, Luiz Alexandre</mods:namePart>
<mods:affiliation id="E3A847758E2C11E415EB6EE47B0E2952">Departamento de Zoologia, Universidade Federal do Rio Grande do Sul. Avenida Bento Gonçalves 9500, 91501 - 970 Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:date id="F728B5C5260706C5A61273C40CBBA50D">2018</mods:date>
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<paragraph id="230B36CFFF9F0111FF24F2FCFDF36A40" blockId="18.[178,759,222,277]" pageId="18" pageNumber="19">
<heading id="784381A3FF9F0111FF24F2FCFDF36A40" centered="true" fontSize="10" level="2" pageId="18" pageNumber="19" reason="8">
<taxonomicName id="E4B44D4CFF9F0111FF24F2FCFD056BA3" authority="Garbelotto &amp; Kochenborger" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[178,755,222,246]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="otarus" status="sp. nov.">
<emphasis id="11C0EADDFF9F0111FF24F2FCFE676BA0" box="[178,401,222,246]" italics="true" pageId="18" pageNumber="19">Lineostethus otarus</emphasis>
Garbelotto &amp; Kochenborger
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,
<taxonomicNameLabel id="0AF357A6FF9F0111FE33F2DCFDF36A40" box="[421,517,254,277]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<paragraph id="230B36CFFF9F0111FF09F30EFCFC6A15" blockId="18.[159,778,300,346]" box="[159,778,300,320]" pageId="18" pageNumber="19">
http://zoobank.org/
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</paragraph>
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<subSubSection id="6BAE6544FF9F0117FE15F364FDC36A9C" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="description">
<paragraph id="230B36CFFF9F0111FE15F364FDD06A0F" blockId="18.[159,778,300,346]" box="[387,550,326,346]" pageId="18" pageNumber="19">
<figureCitation id="BB8F2A4AFF9F0111FE15F364FE4A6A0F" box="[387,444,326,346]" captionStart="Figures 18" captionStartId="2.[159,226,1248,1268]" captionTargetBox="[163,1432,201,1225]" captionTargetId="figure-1@2.[159,1436,197,1228]" captionTargetPageId="2" captionText="Figures 18.Lineostethus, dorsal aspect:(1) L. clypeatus (female); (2) L. tenebricornis (male paratype); (3) L. marginellus (male); (4) L. graziae (female holotype, photo by Mike Narahara CAS); (5) L. sinuosus sp. nov. (male holotype); (6) L. acuminatus sp. nov. (male holotype); (7) L. otarus sp. nov. (male holotype); (8) L. auritus sp. nov. (male holotype). Scale bars: 1mm." pageId="18" pageNumber="19">Figs 7</figureCitation>
,
<figureCitation id="BB8F2A4AFF9F0111FE50F364FDF36A0F" box="[454,517,326,346]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">6269</figureCitation>
,
<figureCitation id="BB8F2A4AFF9F0111FD98F364FDD06A0F" box="[526,550,326,346]" captionStart="Figure 78" captionStartId="22.[159,217,936,956]" captionTargetBox="[164,1434,198,913]" captionTargetId="figure-21@22.[159,1436,197,916]" captionTargetPageId="22" captionText="Figure 78. Distributional map for studied specimens of the species of Lineostethus. Countries with species records are in green." pageId="18" pageNumber="19">78</figureCitation>
</paragraph>
<paragraph id="230B36CFFF9F0111FF4DF351FD45699A" blockId="18.[157,780,371,1735]" pageId="18" pageNumber="19">
Diagnosis.
<taxonomicName id="E4B44D4CFF9F0111FED5F351FE186AD3" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[323,494,371,390]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="otarus" status="sp. nov.">
<emphasis id="11C0EADDFF9F0111FED5F351FE186AD3" box="[323,494,371,390]" italics="true" pageId="18" pageNumber="19">Lineostethus otarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF357A6FF9F0111FE66F356FDCD6AD2" box="[496,571,372,391]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="E4B44D4CFF9F0111FDF1F356FD4A6AD3" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[615,700,372,390]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="auritus" status="sp. nov.">
<emphasis id="11C0EADDFF9F0111FDF1F356FD4A6AD3" box="[615,700,372,390]" italics="true" pageId="18" pageNumber="19">L. auritus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF357A6FF9F0111FD56F356FCFC6AD2" box="[704,778,372,391]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
share similar features in both male and female genitalia: the long foliaceous expansions of the dorsal rim of pygophore; two lines of bristles on the ventral wall, one of them not associated with punctures; and broad gonocoxites VIII.
<taxonomicName id="E4B44D4CFF9F0111FD87F3C9FD4A6AA8" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[529,700,491,510]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="otarus" status="sp. nov.">
<emphasis id="11C0EADDFF9F0111FD87F3C9FD4A6AA8" box="[529,700,491,510]" italics="true" pageId="18" pageNumber="19">Lineostethus otarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF357A6FF9F0111FD56F3C9FCFC6AAB" box="[704,778,491,510]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished by the wider foliaceous expansions of the dorsal rim, hiding the posterolateral angles of pygophore; the sinuous proximal portion of the lateral margin of the expansions; and the projections of the superior layer of ventral rim more elongated. Females of
<taxonomicName id="E4B44D4CFF9F0111FE99F0A3FE9769C6" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[271,353,641,659]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="otarus" status="sp. nov.">
<emphasis id="11C0EADDFF9F0111FE99F0A3FE9769C6" box="[271,353,641,659]" italics="true" pageId="18" pageNumber="19">L. otarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF357A6FF9F0111FEFEF0A2FE4169C6" box="[360,439,640,659]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
are distinguished by the posterior margin of urosternite VII subrectilinear over laterotergites IX, and the sinuous posterior margin of gonocoxites VIII.
</paragraph>
<paragraph id="230B36CFFF9F0111FF4DF0F8FD486F24" blockId="18.[157,780,371,1735]" pageId="18" pageNumber="19">Description. General color of the specimens with some variation, in melanic specimens margins of connexival segments with lines of black punctures more evident. Antennae ivory; segments IIIII with longitudinal brown lines dorsally and ventrally; segments IVV usually darker than the previous. Anterolateral margins of pronotum and proximal portion of lateral margin of corium punctured. Anterior margin of scutellum with two to four brown spots, and one ivory spot medially; post-frenal lobe with 1+1 lateral brown maculae. Unpunctured area laterad to the apex of radial vein of coria inconspicuous to equivalent to the width of one eye. Hemelytral membrane at least reaching apex of abdomen, with 6 veins. Ventrally, punctures uniformly distributed, absent on sterna. Femora with sparse brown circular spots on distal half.</paragraph>
<paragraph id="230B36CFFF9F0111FF4DF65EFB506AD9" blockId="18.[157,780,371,1735]" lastBlockId="18.[815,1438,198,1741]" pageId="18" pageNumber="19">
Male genitalia. Foliaceous expansions of dorsal rim subrectangular, elongated, longer than wide, narrowing towards the apex; proximal portion of the lateral margin of expansions sinuous; apex rounded, surpassing the posterior limit of ventral rim by half of the expansion length, and visible dorsally when fixed at the specimen (
<figureCitation id="BB8F2A4AFF9F0111FE19F733FE2E6E71" box="[399,472,1297,1317]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 62</figureCitation>
: fdr). Basal impression of the foliaceous expansions subrectilinear, cutting of dorsal rim subrectangular (
<figureCitation id="BB8F2A4AFF9F0111FEA8F76EFE726E35" box="[318,388,1356,1376]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 62</figureCitation>
: bi, dr). Posterolateral angles acute and dorsally hidden by the dorsal rim expansions, covered by setae (
<figureCitation id="BB8F2A4AFF9F0111FF32F7AAFEE26ECE" box="[164,276,1416,1436]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Figs 6263</figureCitation>
: pa). Margin of the superior layer of ventral rim V-shaped, median excavation drop-shaped (
<figureCitation id="BB8F2A4AFF9F0111FDD9F784FD656EEC" box="[591,659,1446,1466]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 63</figureCitation>
: vr). Margin of the inferior layer of ventral rim with two lines of short bristles: one posterior, subrectilinear and not associated with punctures; and another anterior, concave and associated with punctures (
<figureCitation id="BB8F2A4AFF9F0111FE87F43FFE896D64" box="[273,383,1565,1585]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Figs 6364</figureCitation>
: b). Segment X with a line of setae on middle (
<figureCitation id="BB8F2A4AFF9F0111FF64F419FEC16D1A" box="[242,311,1595,1615]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 62</figureCitation>
: X). Projections of the superior layer of ventral rim forming a right angle with the posterolateral angles, covered by setae; dorsal margin convex and ventral margin subrectlinear, elongated and uniformly wide throughout its length, apex acute and hook-shaped (
<figureCitation id="BB8F2A4AFF9F0111FEC6F491FE656D93" box="[336,403,1715,1735]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 64</figureCitation>
: psl, pa). Parameres parallel to segment X, at least reaching the foliaceous expansions of dorsal rim; head lozenge (
<figureCitation id="BB8F2A4AFF9F0111FC11F2C1FC056BA2" box="[903,1011,227,247]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Figs 6466</figureCitation>
: par, X, fdr). Ventral surface of phallotheca concave (
<figureCitation id="BB8F2A4AFF9F0111FC18F323FC226A40" box="[910,980,257,277]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 67</figureCitation>
: ph, ap). Vesica apex subquadrangular; apical margin with a sclerotized contour, subrectilinear, and surpassing ductus seminis distalis (
<figureCitation id="BB8F2A4AFF9F0111FB87F31FFBA26A05" box="[1041,1108,317,337]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 67</figureCitation>
: v, av, ds). Distal portion of ductus seminis distalis placed in a concavity on the middle of vesical, short and bending down (
<figureCitation id="BB8F2A4AFF9F0111FBA2F35BFB8F6AD9" box="[1076,1145,377,397]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 67</figureCitation>
: ds).
</paragraph>
<paragraph id="230B36CFFF9F0111FCFBF3B5FAA76F55" blockId="18.[815,1438,198,1741]" pageId="18" pageNumber="19">
Female genitalia. Posterior margin of urosternite VII sinuous, middle third subrectilinear and lateral thirds convex (
<figureCitation id="BB8F2A4AFF9F0111FC1BF3F0FC2A6AB3" box="[909,988,466,486]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 68</figureCitation>
: VII). Gonocoxites VIII subquadrangular, broad; posterior margin slightly sinuous, subrectilinear over laterotergites IX, with external angles truncated and weakly projecting over laterotergites VIII (
<figureCitation id="BB8F2A4AFF9F0111FB0FF00EFB15696A" box="[1177,1251,556,576]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 68</figureCitation>
: gcVIII). Posterior margin of laterotergites VIII subrectilinear; limit between laterotergites VIII and tergite VIII indistinguishable (
<figureCitation id="BB8F2A4AFF9F0111FAC6F04AFA6E692E" box="[1360,1432,616,636]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 68</figureCitation>
: la8). Posterior margin of gonocoxites IX subrectilinear to convex (
<figureCitation id="BB8F2A4AFF9F0111FC13F081FC3A69E2" box="[901,972,675,695]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 68</figureCitation>
: gcIX). Laterotergites IX digitiform, tumescent at apex; apex varying between rounded and slightly acute (
<figureCitation id="BB8F2A4AFF9F0111FAE0F0E3FCBD69A6" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 68</figureCitation>
: la9). Posterior margin of segment X subrectilinear (
<figureCitation id="BB8F2A4AFF9F0111FAC4F0FDFA6169A6" box="[1362,1431,735,755]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 68</figureCitation>
: X). Thickening of gonapophyses IX kidney-shaped, surpassing the lateral margin of the thickening of vaginal intima (
<figureCitation id="BB8F2A4AFF9F0111FAC4F139FA61687B" box="[1362,1431,795,815]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 69</figureCitation>
: gpIX, tgIX). Thickening of vaginal intima subquadrangular (
<figureCitation id="BB8F2A4AFF9F0111FAEEF11BFCBD683F" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 69</figureCitation>
: vi). Distal portion of ductus receptaculi apically enlarged and more sclerotized (
<figureCitation id="BB8F2A4AFF9F0111FBB7F157FB9B68DD" box="[1057,1133,885,905]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 69</figureCitation>
: ddr); anterior annular crest turned to ductus receptaculi; pars intermedialis membranous at apex, longer than capsula seminalis; posterior annular crest perpendicular to pars intermedialis (
<figureCitation id="BB8F2A4AFF9F0111FB3FF1ECFB0568B4" box="[1193,1267,974,994]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 69</figureCitation>
: pi, cs). Capsula seminalis shallowly constricted near base (
<figureCitation id="BB8F2A4AFF9F0111FB48F1CEFAD368AA" box="[1246,1317,1004,1024]" captionStart="Figures 6269" captionStartId="19.[151,217,1545,1565]" captionTargetBox="[247,1335,203,1519]" captionTargetId="figure-166@19.[241,1338,197,1522]" captionTargetPageId="19" captionText="Figures 6269.Lineostethus otarus sp. nov.: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm." pageId="18" pageNumber="19">Fig. 69</figureCitation>
: cs).
</paragraph>
<paragraph id="230B36CFFF9F0111FCFBF629FC2F6E1D" blockId="18.[815,1438,198,1741]" pageId="18" pageNumber="19">Measurements. Male (n = 5): head length, 1.75 ± 0.06 (1.681.82); width, 3.13 ± 0.06 (3.043.20); length of antennal segments: I, 0.48 ± 0.02 (0.460.50); II, 0.73 ± 0.03 (0.700.78); III, 0.80 ± 0.01 (0.800.82); IV, 0.97 ± 0.09 (0.901.10); V, 1.06 ± 0.07 (0.981.12); length of labial segments: I, 0.76 ± 0.04 (0.720.80); II, 1.63 ± 0.11 (1.521.82); III, 0.72 ± 0.07 (0.660.84); IV, 0.53 ± 0.02 (0.500.54); pronotum length, 1.79 ± 0.07 (1.781.86); width, 4.41 ± 0.33 (4.204.98); scutellum length, 3.46 ± 0.15 (3.283.68); width, 2.74 ± 0.07 (2.672.84); total length, 8.16 ± 0.25 (7.808.40); abdominal width, 4.82 ± 0.10 (4.704.95).</paragraph>
<paragraph id="230B36CFFF9F0111FCFBF771FA626D27" blockId="18.[815,1438,198,1741]" pageId="18" pageNumber="19">Female (n = 5): head length, 1.81 ± 0.08 (1.721.92); width, 3.22 ± 0.11 (3.103.36); length of antennal segments: I, 0.49 ± 0.02 (0.480.52); II, 0.77 ± 0.05 (0.700.80); III, 0.82 ± 0.09 (0.720.92); IV, 0.97 ± 0.05 (0.921.02); V, 1.16 ± 0.04 (1.101.12); length of labial segments: I, 0.77 ± 0.10 (0.600.86); II, 1.40 ± 0.45 (0.601.70); III, 0.71 ± 0.08 (0.620.84); IV, 0.52 ± 0.04 (0.460.56); pronotum length, 1.82 ± 0.09 (1.821.94); width, 4.35 ± 0.21 (4.104.68); scutellum length, 3.54 ± 0.18 (3.283.68); width, 2.86 ± 0.14 (2.703.07); total length, 8.58 ± 0.30 (8.209.00); abdominal width, 4.94 ± 0.30 (4.555.36).</paragraph>
<paragraph id="230B36CFFF9F0117FCFBF45CFD476A04" blockId="18.[815,1438,198,1741]" lastBlockId="20.[157,779,198,457]" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
Type material.
<materialsCitation id="93DC3C92FF9F0111FB9DF45FFB4F6D99" collectingDate="1982-08-15" collectionCode="AMNH" collectorName="C. W. &amp; L. O'Brien &amp; G. Wibmer" country="Mexico" location="Morelia" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Michoacan de Ocampo" typeStatus="holotype">
<typeStatus id="FC0F886DFF9F0111FB9DF45FFB9F6DC4" box="[1035,1129,1661,1681]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="5BA3765FFF9F0111FB55F45CFAE86DC4" box="[1219,1310,1662,1681]" name="Mexico" pageId="18" pageNumber="19">MEXICO</collectingCountry>
,
<emphasis id="11C0EADDFF9F0111FAB8F45FFA606DC5" box="[1326,1430,1661,1680]" italics="true" pageId="18" pageNumber="19">
<collectingRegion id="E170F82DFF9F0111FAB8F45FFA606DC5" box="[1326,1430,1661,1680]" country="Mexico" name="Michoacan de Ocampo" pageId="18" pageNumber="19">Michoacán</collectingRegion>
</emphasis>
: (Hwy. 15,
<quantity id="E44C9B2AFF9F0111FC34F4B9FC176DFB" box="[930,993,1691,1710]" metricMagnitude="4" metricUnit="m" metricValue="3.3796224" pageId="18" pageNumber="19" unit="mi" value="21.0">21 mi</quantity>
E
<location id="266B6014FF9F0111FB95F4B9FBA06DFB" LSID="urn:lsid:plazi:treatment:AB1D87D9FF9F0117FF24F2FCFDC36A9C:266B6014FF9F0111FB95F4B9FBA06DFB" box="[1027,1110,1691,1710]" country="Mexico" name="Morelia" pageId="18" pageNumber="19" stateProvince="Michoacan de Ocampo">Morelia</location>
, 7000),
<date id="570A100FFF9F0111FB21F4B9FAB66DFB" box="[1207,1344,1691,1710]" pageId="18" pageNumber="19" value="1982-08-15">
<collectingDate id="474EE9E7FF9F0111FB21F4B9FAB66DFB" box="[1207,1344,1691,1710]" pageId="18" pageNumber="19" value="1982-08-15">15-VIII-1982</collectingDate>
</date>
,
<collectorName id="8E415319FF9F0111FAD8F4B9FC626D99" pageId="18" pageNumber="19">C.W. &amp; L.OBrien</collectorName>
&amp;
<collectorName id="8E415319FF9F0111FC2FF49BFBD26D99" box="[953,1060,1721,1740]" pageId="18" pageNumber="19">G.Wibmer</collectorName>
leg. (
<collectionCode id="45A5AE0AFF9F0111FBF2F49BFB446D99" box="[1124,1202,1721,1740]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="19">AMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="93DC3C92FF9F0117FB50F498FCF06A40" collectingDate="1982-08-15" collectionCode="UFRG, DARC" collectorName="C. W. &amp; L. O'Brien &amp; G. B. Marshall" country="Mexico" lastPageId="20" lastPageNumber="21" location="Hwy" pageId="18" pageNumber="19" specimenCount="42" specimenCount-female="26" specimenCount-male="16" stateProvince="Michoacan de Ocampo" typeStatus="paratype">
<typeStatus id="FC0F886DFF9F0111FB50F498FADA6D98" box="[1222,1324,1722,1741]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="5BA3765FFF9F0111FAAFF49BFA6E6D99" box="[1337,1432,1721,1740]" name="Mexico" pageId="18" pageNumber="19">MEXICO</collectingCountry>
,
<emphasis id="11C0EADDFF990117FF09F2E4FEF26B8C" box="[159,260,198,217]" italics="true" pageId="20" pageNumber="21">
<collectingRegion id="E170F82DFF990117FF09F2E4FEF26B8C" box="[159,260,198,217]" country="Mexico" name="Michoacan de Ocampo" pageId="20" pageNumber="21">Michoacán</collectingRegion>
</emphasis>
: (
<location id="266B6014FF990117FE8FF2E4FEBE6B8F" LSID="urn:lsid:plazi:treatment:AB1D87D9FF9F0117FF24F2FCFDC36A9C:266B6014FF990117FE8FF2E4FEBE6B8F" box="[281,328,198,218]" country="Mexico" name="Hwy" pageId="20" pageNumber="21" stateProvince="Michoacan de Ocampo">Hwy</location>
15,
<locationDeviation id="32B92FEDFF990117FEFBF2E4FDE06B8C" box="[365,534,198,217]" location="Morelia" pageId="20" pageNumber="21">
<quantity id="E44C9B2AFF990117FEFBF2E4FE5C6B8C" box="[365,426,198,217]" metricMagnitude="4" metricUnit="m" metricValue="3.3796224" pageId="20" pageNumber="21" unit="mi" value="21.0">21 mi</quantity>
E Morelia
</locationDeviation>
, 7000),
<specimenCount id="35B2FD46FF990117FDF8F2E4FD4A6B8C" box="[622,700,198,217]" count="8" pageId="20" pageNumber="21" type="male">8 males</specimenCount>
and
<specimenCount id="35B2FD46FF990117FD64F2E4FF1A6BA2" count="13" pageId="20" pageNumber="21" type="female">13 females</specimenCount>
,
<date id="570A100FFF990117FF63F2C6FE836BA2" box="[245,373,228,247]" pageId="20" pageNumber="21" value="1982-08-15">
<collectingDate id="474EE9E7FF990117FF63F2C6FE836BA2" box="[245,373,228,247]" pageId="20" pageNumber="21" value="1982-08-15">15-VIII-1982</collectingDate>
</date>
,
<collectorName id="8E415319FF990117FEE8F2C6FDDE6BA2" box="[382,552,228,247]" pageId="20" pageNumber="21">C.W. &amp; L.OBrien</collectorName>
&amp;
<collectorName id="8E415319FF990117FDD2F2C6FD3F6BA2" box="[580,713,228,247]" pageId="20" pageNumber="21">G.B. Marshall</collectorName>
leg. (
<specimenCount id="35B2FD46FF990117FC96F2C6FF386A40" count="1" pageId="20" pageNumber="21" type="male">1 male</specimenCount>
<collectionCode id="45A5AE0AFF990117FF43F320FEE46A40" box="[213,274,258,277]" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="20" pageNumber="21">UFRG</collectionCode>
;
<specimenCount id="35B2FD46FF990117FE8BF320FE846A40" box="[285,370,258,277]" count="1" pageId="20" pageNumber="21" type="female">1 female</specimenCount>
AMNH;
<specimenCount id="35B2FD46FF990117FE5BF320FDEF6A40" box="[461,537,258,277]" count="7" pageId="20" pageNumber="21" type="male">7 males</specimenCount>
and
<specimenCount id="35B2FD46FF990117FDDBF320FD4E6A40" box="[589,696,258,277]" count="12" pageId="20" pageNumber="21" type="female">12 females</specimenCount>
<collectionCode id="45A5AE0AFF990117FD29F320FCF66A40" box="[703,768,258,277]" pageId="20" pageNumber="21">DARC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="93DC3C92FF990117FF36F302FD5B6A04" collectingDate="1983-05-14" collectionCode="UFRG" collectorName="C. W. &amp; L. O'Brien &amp; G. B. Marshall" country="Mexico" location="Hwy" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" stateProvince="Puebla" typeStatus="paratype">
<emphasis id="11C0EADDFF990117FF36F302FF2B6A66" box="[160,221,288,307]" italics="true" pageId="20" pageNumber="21">
<collectingRegion id="E170F82DFF990117FF36F302FF2B6A66" box="[160,221,288,307]" country="Mexico" name="Puebla" pageId="20" pageNumber="21">Puebla</collectingRegion>
</emphasis>
: (
<location id="266B6014FF990117FF78F33DFEEB6A66" LSID="urn:lsid:plazi:treatment:AB1D87D9FF9F0117FF24F2FCFDC36A9C:266B6014FF990117FF78F33DFEEB6A66" box="[238,285,287,307]" country="Mexico" name="Hwy" pageId="20" pageNumber="21" stateProvince="Puebla">Hwy</location>
<locationDeviation id="32B92FEDFF990117FEB7F302FE066A66" box="[289,496,288,307]" location="Azumbilla" pageId="20" pageNumber="21">
<quantity id="E44C9B2AFF990117FEB7F302FEAF6A66" box="[289,345,288,307]" metricMagnitude="4" metricUnit="m" metricValue="2.8968192" pageId="20" pageNumber="21" unit="mi" value="18.0">18 mi</quantity>
NW Azumbilla
</locationDeviation>
, 7700),
<specimenCount id="35B2FD46FF990117FDD4F302FD616A66" box="[578,663,288,307]" count="1" pageId="20" pageNumber="21" type="female">1 female</specimenCount>
,
<date id="570A100FFF990117FD09F302FCF06A66" box="[671,774,288,307]" pageId="20" pageNumber="21" value="1983-05-14">
<collectingDate id="474EE9E7FF990117FD09F302FCF06A66" box="[671,774,288,307]" pageId="20" pageNumber="21" value="1983-05-14">14-V-1983</collectingDate>
</date>
, 1100,
<collectorName id="8E415319FF990117FF76F31CFE7B6A05" box="[224,397,317,337]" pageId="20" pageNumber="21">C.W. &amp; L.OBrien</collectorName>
&amp;
<collectorName id="8E415319FF990117FE3CF31CFDC76A05" box="[426,561,317,337]" pageId="20" pageNumber="21">G.B. Marshall</collectorName>
leg. (
<collectionCode id="45A5AE0AFF990117FDF0F31CFD506A04" box="[614,678,318,337]" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="20" pageNumber="21">UFRG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="77CB6647FF9E0110FF01F42BFA606DBB" pageId="19" pageNumber="20" startId="19.[151,217,1545,1565]" targetBox="[247,1335,203,1519]" targetPageId="19" targetType="figure">
<paragraph id="230B36CFFF9E0110FF01F42BFA606DBB" blockId="19.[149,1432,1544,1774]" pageId="19" pageNumber="20">
Figures 6269.
<taxonomicName id="E4B44D4CFF9E0110FEB7F42AFE3D6D49" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[289,459,1544,1564]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="otarus" status="sp. nov.">
<emphasis id="11C0EADDFF9E0110FEB7F42AFE3D6D49" box="[289,459,1544,1564]" italics="true" pageId="19" pageNumber="20">Lineostethus otarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF357A6FF9E0110FE46F42BFDED6D48" box="[464,539,1545,1565]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
: (6264) male pygophore, dorsal, ventral and posterior views respectively; (6566) left paramere, dorsal and lateral views; (67) phallus, lateral view; 68, female genital plates; (69) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area;pa, posterolateral angle; par,paramere;ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 6266, 68 = 1 mm; 67, 69 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="230B36CFFF990117FF4DF379FCFD6A3B" blockId="20.[157,779,198,457]" box="[219,779,347,367]" pageId="20" pageNumber="21">
Distribution (
<figureCitation id="BB8F2A4AFF990117FEC9F379FE546A3B" box="[351,418,347,367]" captionStart="Figure 78" captionStartId="22.[159,217,936,956]" captionTargetBox="[164,1434,198,913]" captionTargetId="figure-21@22.[159,1436,197,916]" captionTargetPageId="22" captionText="Figure 78. Distributional map for studied specimens of the species of Lineostethus. Countries with species records are in green." pageId="20" pageNumber="21">Fig. 78</figureCitation>
).
<collectingCountry id="5BA3765FFF990117FE27F379FE0B6A3B" box="[433,509,347,366]" name="Mexico" pageId="20" pageNumber="21">México</collectingCountry>
:
<emphasis id="11C0EADDFF990117FD95F379FD9E6A3B" box="[515,616,347,366]" italics="true" pageId="20" pageNumber="21">
<collectingRegion id="E170F82DFF990117FD95F379FD9E6A3B" box="[515,616,347,366]" country="Mexico" name="Michoacan de Ocampo" pageId="20" pageNumber="21">Michoacán</collectingRegion>
</emphasis>
,
<emphasis id="11C0EADDFF990117FDE2F379FD446A3B" box="[628,690,347,366]" italics="true" pageId="20" pageNumber="21">
<collectingRegion id="E170F82DFF990117FDE2F379FD446A3B" box="[628,690,347,366]" country="Mexico" name="Puebla" pageId="20" pageNumber="21">Puebla</collectingRegion>
</emphasis>
(
<figureCitation id="BB8F2A4AFF990117FD2AF379FCF66A3B" box="[700,768,347,367]" captionStart="Figure 78" captionStartId="22.[159,217,936,956]" captionTargetBox="[164,1434,198,913]" captionTargetId="figure-21@22.[159,1436,197,916]" captionTargetPageId="22" captionText="Figure 78. Distributional map for studied specimens of the species of Lineostethus. Countries with species records are in green." pageId="20" pageNumber="21">Fig. 78</figureCitation>
).
</paragraph>
<paragraph id="230B36CFFF990117FF4DF35BFDC36A9C" blockId="20.[157,779,198,457]" pageId="20" pageNumber="21">
Etymology. The epithet refers to the shape of the foliaceous expansions of dorsal rim of pygophore; adjective
<taxonomicName id="E4B44D4CFF990117FDE0F3BAFD596AFF" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[630,687,408,426]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="otarus">
<emphasis id="11C0EADDFF990117FDE0F3BAFD596AFF" box="[630,687,408,426]" italics="true" pageId="20" pageNumber="21">otarus</emphasis>
</taxonomicName>
(
<emphasis id="11C0EADDFF990117FD2AF3BAFD386AFF" box="[700,718,408,426]" italics="true" pageId="20" pageNumber="21">-a</emphasis>
,
<emphasis id="11C0EADDFF990117FD4EF3BAFD0B6AFF" box="[728,765,408,426]" italics="true" pageId="20" pageNumber="21">-um</emphasis>
), latinized from Greek
<emphasis id="11C0EADDFF990117FEE4F394FE5C6A9D" box="[370,426,438,456]" italics="true" pageId="20" pageNumber="21">otaros</emphasis>
= large-eared.
</paragraph>
</subSubSection>
</treatment>
</document>