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<document id="9DDD40F1AE49B8CBDADCC2D6014A568E" ID-DOI="10.3897/zoologia.35.e21232" ID-ISSN="1984-4689" ID-Zenodo-Dep="13174064" ID-ZooBank="663F033F-E804-444F-A999-9F42993FF545" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722587231085" checkinUser="felipe" docAuthor="Garbelotto, Thereza de Almeida, Kochenborger, Ana Paula Leite &amp; Campos, Luiz Alexandre" docDate="2018" docId="AB1D87D9FF820112FCF0F401FC076D99" docLanguage="en" docName="Zoologia.35.e21232.pdf" docOrigin="Zoologia (e 21232) 35" docSource="http://dx.doi.org/10.3897/zoologia.35.e21232" docStyle="DocumentStyle:9D87517C92C8E430512793D922E147D4.1:Zoologia.2017-2020.journal_article" docStyleId="9D87517C92C8E430512793D922E147D4" docStyleName="Zoologia.2017-2020.journal_article" docStyleVersion="1" docTitle="Lineostethus acuminatus Kochenborger &amp; Garbelotto 2018, sp. nov." docType="treatment" docUuid="3E36196F-22D0-4FC2-8486-8E2BCCDCC36D" docUuidSource="ZooBank" docVersion="2" lastPageNumber="18" masterDocId="5724FFA1FF8D0103FF96F222FFF66B55" masterDocTitle="Revision of Lineostethus (Heteroptera: Pentatomidae: Discocephalini)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="16" updateTime="1722619988372" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B7EC4A13E6167ADD1FB4E857B673FFDC">Revision of Lineostethus (Heteroptera: Pentatomidae: Discocephalini)</mods:title>
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<mods:name id="D6B52F48A7A3740E7D67FCCE8F767881" type="personal">
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<mods:namePart id="60601315568EF549310B857A165E414E">Garbelotto, Thereza de Almeida</mods:namePart>
<mods:affiliation id="368F632018C795A9BD9E04C3E3E4C983">Departamento de Ciências Biológicas, Universidade do Sul Catarinense. Avenida José Acácio Moreira 787, 88704 - 900 Tubarão, SC, Brazil.</mods:affiliation>
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<mods:namePart id="704A66AEFC8A52019503A38F7E51F978">Kochenborger, Ana Paula Leite</mods:namePart>
<mods:affiliation id="585C10CA507275DED0A35FF0310BA2EB">Departamento de Zoologia, Universidade Federal do Rio Grande do Sul. Avenida Bento Gonçalves 9500, 91501 - 970 Porto Alegre, RS, Brazil.</mods:affiliation>
<mods:nameIdentifier id="B61EA957E3E714A0786216A6A3C725FD" type="email">therezagarbelotto@hotmail</mods:nameIdentifier>
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<mods:namePart id="02CC2C8DEC831A4443ABB4E5E066E50A">Campos, Luiz Alexandre</mods:namePart>
<mods:affiliation id="FD90184CF0A6218D965700677651C1A9">Departamento de Zoologia, Universidade Federal do Rio Grande do Sul. Avenida Bento Gonçalves 9500, 91501 - 970 Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:title id="4A65FC71BB2B81FB415173A341A83BCC">Zoologia</mods:title>
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<mods:date id="33794466C2324A45586FBD07AB95B912">2018</mods:date>
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<mods:title id="CA082375E048ACC428408153DA83181B">e 21232</mods:title>
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<mods:number id="B309BA08F9CF035F1D9F46E557B19C3E">2018-11-01</mods:number>
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<mods:number id="075B68D2C7CF295035CFBD9E8FD95C67">35</mods:number>
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<subSubSection id="6BAE6544FF82010CFCF0F401FA626DD3" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="230B36CFFF82010CFCF0F401FB2F6D0E" blockId="15.[870,1368,1571,1627]" pageId="15" pageNumber="16">
<heading id="784381A3FF82010CFCF0F401FB2F6D0E" centered="true" fontSize="10" level="2" pageId="15" pageNumber="16" reason="8">
<taxonomicName id="E4B44D4CFF82010CFCF0F401FB9B6D0E" authority="Kochenborger &amp; Garbelotto" authorityName="Kochenborger &amp; Garbelotto" authorityYear="2018" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="11C0EADDFF82010CFCF0F401FB726D6E" box="[870,1156,1571,1595]" italics="true" pageId="15" pageNumber="16">Lineostethus acuminatus</emphasis>
Kochenborger &amp; Garbelotto
</taxonomicName>
,
<taxonomicNameLabel id="0AF357A6FF82010CFBEFF466FB2F6D0E" box="[1145,1241,1604,1627]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<paragraph id="230B36CFFF82010CFCBFF450FA626DD3" blockId="15.[809,1428,1650,1695]" box="[809,1428,1650,1670]" pageId="15" pageNumber="16">
http://zoobank.org/
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</paragraph>
</subSubSection>
<subSubSection id="6BAE6544FF820112FB9BF4A9FC076D99" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="230B36CFFF82010CFB9BF4A9FB466DCA" blockId="15.[809,1428,1650,1695]" box="[1037,1200,1675,1695]" pageId="15" pageNumber="16">
<figureCitation id="BB8F2A4AFF82010CFB9BF4A9FBB16DCA" box="[1037,1095,1675,1695]" captionStart="Figures 1–8" captionStartId="2.[159,226,1248,1268]" captionTargetBox="[163,1432,201,1225]" captionTargetId="figure-1@2.[159,1436,197,1228]" captionTargetPageId="2" captionText="Figures 1–8.Lineostethus, dorsal aspect:(1) L. clypeatus (female); (2) L. tenebricornis (male paratype); (3) L. marginellus (male); (4) L. graziae (female holotype, photo by Mike Narahara – CAS); (5) L. sinuosus sp. nov. (male holotype); (6) L. acuminatus sp. nov. (male holotype); (7) L. otarus sp. nov. (male holotype); (8) L. auritus sp. nov. (male holotype). Scale bars: 1mm." pageId="15" pageNumber="16">Figs 6</figureCitation>
,
<figureCitation id="BB8F2A4AFF82010CFBC6F4A9FB796DCA" box="[1104,1167,1675,1695]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="15" pageNumber="16">54–61</figureCitation>
,
<figureCitation id="BB8F2A4AFF82010CFB0EF4A9FB466DCA" box="[1176,1200,1675,1695]" captionStart="Figure 78" captionStartId="22.[159,217,936,956]" captionTargetBox="[164,1434,198,913]" captionTargetId="figure-21@22.[159,1436,197,916]" captionTargetPageId="22" captionText="Figure 78. Distributional map for studied specimens of the species of Lineostethus. Countries with species records are in green." pageId="15" pageNumber="16">78</figureCitation>
</paragraph>
<paragraph id="230B36CFFF820112FCF3F49AFE3B6908" blockId="15.[809,1429,1720,1770]" lastBlockId="17.[149,774,198,1741]" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
Diagnosis. Males of
<taxonomicName id="E4B44D4CFF82010CFBA1F49BFB366D9E" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[1079,1216,1721,1739]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="11C0EADDFF82010CFBA1F49BFB366D9E" box="[1079,1216,1721,1739]" italics="true" pageId="15" pageNumber="16">L. acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF357A6FF82010CFB5CF49BFAEA6D99" box="[1226,1308,1721,1740]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
differ from most species by the subtriangular expansions of the dorsal rim, although resembling
<taxonomicName id="E4B44D4CFF9C0112FEF4F2E5FE4E6B8C" authorityName=", Hildebrand and Becker" authorityYear="1982" box="[354,440,199,217]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="11C0EADDFF9C0112FEF4F2E5FE4E6B8C" box="[354,440,199,217]" italics="true" pageId="17" pageNumber="18">L. graziae</emphasis>
</taxonomicName>
.
<taxonomicName id="E4B44D4CFF9C0112FE52F2E5FDB26B8C" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[452,580,199,217]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="11C0EADDFF9C0112FE52F2E5FDB26B8C" box="[452,580,199,217]" italics="true" pageId="17" pageNumber="18">L. acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF357A6FF9C0112FDD0F2E4FD676B8C" box="[582,657,198,217]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
expansions are longer and narrower than
<taxonomicName id="E4B44D4CFF9C0112FE5EF2C6FDD56BA3" authorityName=", Hildebrand and Becker" authorityYear="1982" box="[456,547,228,246]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="11C0EADDFF9C0112FE5EF2C6FDD56BA3" box="[456,547,228,246]" italics="true" pageId="17" pageNumber="18">L. graziae</emphasis>
</taxonomicName>
(
<figureCitation id="BB8F2A4AFF9C0112FDA7F2C1FD8F6BA2" box="[561,633,227,247]" captionStart="Figures 37–44" captionStartId="12.[159,226,1544,1564]" captionTargetBox="[333,1267,202,1523]" captionTargetId="figure-165@12.[329,1267,197,1529]" captionTargetPageId="12" captionText="Figures 37–44. Lineostethus graziae: (37–39) male pygophore, dorsal, ventral and posterior views respectively; (40–41) left paramere, dorsal and lateral views; (42) phallus, lateral view; (43) female genital plates; (44) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area;od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 37–41, 43 = 1 mm; 42, 44 = 0.5 mm." pageId="17" pageNumber="18">Figs 37</figureCitation>
,
<figureCitation id="BB8F2A4AFF9C0112FD10F2C6FD576BA2" box="[646,673,228,247]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">54</figureCitation>
: fdr), and the dorsal rim cutting is U-shaped in
<taxonomicName id="E4B44D4CFF9C0112FDB7F320FD5C6A41" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[545,682,258,276]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="11C0EADDFF9C0112FDB7F320FD5C6A41" box="[545,682,258,276]" italics="true" pageId="17" pageNumber="18">L. acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF357A6FF9C0112FD25F320FCF26A40" box="[691,772,258,277]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
while subquadrangular in
<taxonomicName id="E4B44D4CFF9C0112FE36F302FE0D6A67" authorityName=", Hildebrand and Becker" authorityYear="1982" box="[416,507,288,306]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="11C0EADDFF9C0112FE36F302FE0D6A67" box="[416,507,288,306]" italics="true" pageId="17" pageNumber="18">L. graziae</emphasis>
</taxonomicName>
(
<figureCitation id="BB8F2A4AFF9C0112FD91F33DFDB96A66" box="[519,591,287,307]" captionStart="Figures 37–44" captionStartId="12.[159,226,1544,1564]" captionTargetBox="[333,1267,202,1523]" captionTargetId="figure-165@12.[329,1267,197,1529]" captionTargetPageId="12" captionText="Figures 37–44. Lineostethus graziae: (37–39) male pygophore, dorsal, ventral and posterior views respectively; (40–41) left paramere, dorsal and lateral views; (42) phallus, lateral view; (43) female genital plates; (44) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area;od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 37–41, 43 = 1 mm; 42, 44 = 0.5 mm." pageId="17" pageNumber="18">Figs 37</figureCitation>
,
<figureCitation id="BB8F2A4AFF9C0112FDCDF33DFD806A67" box="[603,630,287,306]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">54</figureCitation>
: dr). Also, the projections of the superior layer of the ventral rim in
<taxonomicName id="E4B44D4CFF9C0112FD15F31CFCF46A05" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[643,770,318,336]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="11C0EADDFF9C0112FD15F31CFCF46A05" box="[643,770,318,336]" italics="true" pageId="17" pageNumber="18">L. acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF357A6FF9C0112FF03F37EFF166A3A" box="[149,224,348,367]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
are uniformly wide and the apex is hook-shaped (
<figureCitation id="BB8F2A4AFF9C0112FD2DF379FD086A3B" box="[699,766,347,367]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 55</figureCitation>
: psl). The phallus of
<taxonomicName id="E4B44D4CFF9C0112FEC4F358FE226AD9" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[338,468,378,396]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="11C0EADDFF9C0112FEC4F358FE226AD9" box="[338,468,378,396]" italics="true" pageId="17" pageNumber="18">L. acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF357A6FF9C0112FE4EF358FDD26AD8" box="[472,548,378,397]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
bears the largest vesica and the longest ductus seminis distalis in the genus (
<figureCitation id="BB8F2A4AFF9C0112FD14F3B5FD326AFF" box="[642,708,407,427]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig.59</figureCitation>
: v, ds). The females of the new species are recognized by the truncated apex of laterotergites IX and the posterior margin of segment X concave to subrectilinear (
<figureCitation id="BB8F2A4AFF9C0112FE05F3D2FE206951" box="[403,470,496,516]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 60</figureCitation>
: la9, X); the circular thickening of vaginal intima, the ductus receptaculi distal helicoid, and the pars intermedialis with an apical constriction help to identify this species (
<figureCitation id="BB8F2A4AFF9C0112FE84F068FEA16908" box="[274,343,586,606]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 61</figureCitation>
: vi, ddr, pi).
</paragraph>
<caption id="77CB6647FF9D0113FF09F7CFFDFE6DA5" pageId="16" pageNumber="17" startId="16.[159,226,1517,1537]" targetBox="[332,1268,199,1489]" targetPageId="16" targetType="figure">
<paragraph id="230B36CFFF9D0113FF09F7CFFDFE6DA5" blockId="16.[157,1439,1516,1776]" pageId="16" pageNumber="17">
Figures 54–61.
<taxonomicName id="E4B44D4CFF9D0113FEA7F7CEFDE66D55" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[305,528,1516,1536]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="11C0EADDFF9D0113FEA7F7CEFDE66D55" box="[305,528,1516,1536]" italics="true" pageId="16" pageNumber="17">Lineostethus acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AF357A6FF9D0113FD80F7CFFD926D54" box="[534,612,1517,1537]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="230B36CFFF9C0112FF42F04AFD45683F" blockId="17.[149,774,198,1741]" pageId="17" pageNumber="18">Description. Antennae ivory; segments II–III with longitudinal brown lines. Anterolateral margins of pronotum and proximal portion of lateral margin of corium punctured in some specimens. Scutellum may present a spot of brown punctures on disc and on post-frenal lobe. Unpunctured area laterad to the apex of radial vein of coria smaller than the width of one eye. Hemelytral membrane reaching the apex of abdomen, with 6–7 veins. Ventrally, punctures sparse and laterally concentrated. Femora with sparse brown circular spots on distal half.</paragraph>
<paragraph id="230B36CFFF9C0112FF42F157FE0F6EAE" blockId="17.[149,774,198,1741]" pageId="17" pageNumber="18">
Male genitalia. Foliaceous expansions of dorsal rim subtriangular; apex of expansions rounded and surpassing the posterior limit of ventral rim; posterolateral angles visible dorsally (
<figureCitation id="BB8F2A4AFF9C0112FF60F1ECFEC968B4" box="[246,319,974,994]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 54</figureCitation>
: fdr, pa). Basal impression of the foliaceous expansions sinuous, cutting of dorsal rim U-shaped (
<figureCitation id="BB8F2A4AFF9C0112FD2FF1CEFCF768AA" box="[697,769,1004,1024]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 54</figureCitation>
: bi, dr). Posterolateral angles rounded (
<figureCitation id="BB8F2A4AFF9C0112FD8EF628FD716F48" box="[536,647,1034,1054]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Figs 55–56</figureCitation>
: pa). Margin of the superior layer of ventral rim sinuous, median excavation drop-shaped (
<figureCitation id="BB8F2A4AFF9C0112FEBCF664FE8E6F0C" box="[298,376,1094,1114]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 55</figureCitation>
: vr). Margin of the inferior layer of ventral rim with one subrectilinear line of short bristles (
<figureCitation id="BB8F2A4AFF9C0112FD4DF641FF2E6FC0" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Figs 55–56</figureCitation>
: b). Segment X with a line of setae on base (
<figureCitation id="BB8F2A4AFF9C0112FD04F6A3FCF66FC0" box="[658,768,1153,1173]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Figs 54–55</figureCitation>
: X). Projections of the superior layer of ventral rim forming a right angle with posterolateral angles, covered by setae; dorsal margin convex and ventral margin concave; narrow at base, apex acute and hook-shaped, median third uniformly enlarged towards the apex (
<figureCitation id="BB8F2A4AFF9C0112FE3CF735FE066E7F" box="[426,496,1303,1323]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 56</figureCitation>
: psl, pa). Parameres parallel or slightly directed to segment X, reaching the foliaceous expansions of dorsal rim; head lozenge (
<figureCitation id="BB8F2A4AFF9C0112FDD6F770FD476E33" box="[576,689,1362,1382]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Figs 56–58</figureCitation>
: par, X, fdr). Ventral surface of phallotheca concave (
<figureCitation id="BB8F2A4AFF9C0112FDF4F752FD5C6ED1" box="[610,682,1392,1412]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 59</figureCitation>
: ph, ap). Vesica apex rounded; apical margin concave extending to mid of ductus seminis distalis (
<figureCitation id="BB8F2A4AFF9C0112FE35F78EFE1C6EEA" box="[419,490,1452,1472]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 59</figureCitation>
: v, av, ds). Distal portion of ductus seminis distalis sinuous, about 2/3 longer than vesica, narrowing posteriorly (
<figureCitation id="BB8F2A4AFF9C0112FE14F7CAFE3C6EAE" box="[386,458,1512,1532]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 59</figureCitation>
: ds).
</paragraph>
<paragraph id="230B36CFFF9C0112FF42F424FB8C6951" blockId="17.[149,774,198,1741]" lastBlockId="17.[807,1430,198,1740]" pageId="17" pageNumber="18">
Female genitalia. Posterior margin of urosternite VII concave (
<figureCitation id="BB8F2A4AFF9C0112FF60F401FECB6D62" box="[246,317,1571,1591]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 60</figureCitation>
: VII). Gonocoxites VIII subtriangular; sutural and posterior margins subrectilinear (
<figureCitation id="BB8F2A4AFF9C0112FD9BF463FDA56D00" box="[525,595,1601,1621]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 60</figureCitation>
: gcVIII). Posterior margin of laterotergites VIII slightly convex; limit between laterotergites VIII and tergite VIII indistinguishable (
<figureCitation id="BB8F2A4AFF9C0112FD20F45FFD096DC5" box="[694,767,1661,1681]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 60</figureCitation>
: la8). Posterior margin of gonocoxites IX subrectilinear (
<figureCitation id="BB8F2A4AFF9C0112FD2DF4B9FD096DFB" box="[699,767,1691,1711]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 60</figureCitation>
: gcIX). Laterotergites IX digitiform and truncate apex; mesial and lateral margins convex (
<figureCitation id="BB8F2A4AFF9C0112FBDDF2E4FB656B8C" box="[1099,1171,198,218]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 60</figureCitation>
: la9). Posterior margin of segment X concave to rectilinear (
<figureCitation id="BB8F2A4AFF9C0112FB1AF2C1FB236BA2" box="[1164,1237,227,247]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 60</figureCitation>
: X). Thickening of gonapophyses IX subrectangular, not surpassing the lateral margin of the thickening of vaginal intima (
<figureCitation id="BB8F2A4AFF9C0112FB43F33DFAEF6A66" box="[1237,1305,287,307]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 61</figureCitation>
: gpIX, tgIX). Thickening of vaginal intima circular and flat (
<figureCitation id="BB8F2A4AFF9C0112FB7CF31FFADB6A05" box="[1258,1325,317,337]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 61</figureCitation>
: vi). Distal portion of ductus receptaculi helicoid (
<figureCitation id="BB8F2A4AFF9C0112FB21F379FB096A3B" box="[1207,1279,347,367]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 61</figureCitation>
: ddr); anterior annular crest turned to ductus receptaculi; pars intermedialis with a shallow constriction on its apex and subequal in length to capsula seminalis; posterior annular crest perpendicular to pars intermedialis (
<figureCitation id="BB8F2A4AFF9C0112FC7FF3F0FBD86AB3" box="[1001,1070,466,486]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 61</figureCitation>
: pi, cs). Capsula seminalis shallowly constricted near base (
<figureCitation id="BB8F2A4AFF9C0112FB9EF3D2FBB86951" box="[1032,1102,496,516]" captionStart="Figures 54–61" captionStartId="16.[159,226,1517,1537]" captionTargetBox="[332,1268,199,1489]" captionTargetId="figure-169@16.[327,1268,197,1496]" captionTargetPageId="16" captionText="Figures 54–61. Lineostethus acuminatus sp. nov.: (54–56) male pygophore, dorsal, ventral and posterior views respectively; (57–58) left paramere, dorsal and lateral views; (59) phallus, lateral view; (60) female genital plates; (61) female receptaculum seminis. Abbreviations: ap, ventral apical projection of phallotheca; av, apical margin of vesica; b, bristles; bi, basal impression of foliaceous expansions; cs, capsula seminalis; ddr, distal ductus receptaculi; dr, dorsal rim; ds, ductus seminis distalis; fdr, foliaceous expansions of dorsal rim; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpVIII, gonapophyses VIII; gpIX, gonapophyses IX; id, internal duct of vesicular area; la8, laterotergites VIII; la9, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pa, posterolateral angle; par, paramere; ph, phallotheca; pi, pars intermedialis; psl, projection of superior layer of ventral rim; rs, ring sclerites; s, spiracle; tgIX, thickening of gonapophyses IX; v, vesica; vi, thickening of vaginal intima; vr, ventral rim; VII, urosternite VII; X, segment X. Scale bars: 54–58, 60 = 1 mm; 59, 61 = 0.5 mm." pageId="17" pageNumber="18">Fig. 61</figureCitation>
: cs).
</paragraph>
<paragraph id="230B36CFFF9C0112FCF3F02DFC276819" blockId="17.[807,1430,198,1740]" pageId="17" pageNumber="18">Measurements. Male (n = 5): head length, 1.54 ± 0.07 (1.43–1.63); width, 2.96 ± 0.16 (2.70–3.10); length of antennal segments: I, 0.40 ± 0.01 (0.40–0.42); II, 0.68 ± 0.04 (0.62–0.70); III, 0.67 ± 0.06 (0.60–0.74); IV, 0.89 ± 0.01 (0.88–0.90); V, 1.05 ± 0.07 (1.00–1.10); length of labial segments: I, 0.66 ± 0.05 (0.60–0.70); II, 1.34 ± 0.05 (1.30–1.40); III, 0.63 ± 0.05 (0.58–0.70); IV, 0.46 ± 0.05 (0.38–0.50); pronotum length, 1.66 ± 0.11 (1.52–1.80); width, 3.95 ± 0.18 (3.65–4.10); scutellum length, 3.30 ± 0.15 (3.08–3.48); width, 2.61 ± 0.12 (2.42–2.73); total length, 6.26 ± 0.30 (6.32–6.53); abdominal width, 4.28 ± 0.19 (4.00–4.45).</paragraph>
<paragraph id="230B36CFFF9C0112FCF3F175FC216FC0" blockId="17.[807,1430,198,1740]" pageId="17" pageNumber="18">Female (n = 5): head length, 1.55 ± 0.06 (1.45–1.63); width, 3.09 ± 0.13 (2.95–3.24); length of antennal segments: I, 0.42 ± 0.02 (0.40–0.46); II, 0.71 ± 0.06 (0.64–0.78); III, 0.65 ± 0.04 (0.60–0.70); IV, 0.81 ± 0.07 (0.74–0.90); V, 0.96 ± 0.03 (0.94–1.00); length of labial segments: I, 0.67 ± 0.04 (0.60– 0.70); II, 1.39 ± 0.08 (1.26–1.48); III, 0.65 ± 0.09 (0.60–0.80); IV, 0.43 ± 0.04 (0.40–0.48); pronotum length, 1.76 ± 0.08 (1.62–1.82); width, 4.07 ± 0.13 (3.88–4.20); scutellum length, 3.36 ± 0.08 (3.28–3.44); width, 2.68 ± 0.10 (2.64–2.79); total length, 6.67 ± 0.19 (6.60–6.96); abdominal width, 4.52 ± 0.14 (4.30–4.65).</paragraph>
<paragraph id="230B36CFFF9C0112FCF3F682FC1F6D4C" blockId="17.[807,1430,198,1740]" pageId="17" pageNumber="18">
Type material.
<materialsCitation id="93DC3C92FF9C0112FC62F6BDFAD16FBB" collectingDate="2008-12-07" collectionCode="DZUP" collectorName="Weirauch &amp; Forero" country="Costa Rica" latitude="10.81" location="Research Station" longLatPrecision="778" longitude="-85.62" municipality="Santa Rosa" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="FC0F886DFF9C0112FC62F6BDFBB96FE6" box="[1012,1103,1183,1203]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="5BA3765FFF9C0112FB0DF682FAEE6FE6" box="[1179,1304,1184,1203]" name="Costa Rica" pageId="17" pageNumber="18">COSTA RICA</collectingCountry>
,
<emphasis id="11C0EADDFF9C0112FAB2F682FA796FE6" box="[1316,1423,1184,1203]" italics="true" pageId="17" pageNumber="18">
<collectingRegion id="E170F82DFF9C0112FAB2F682FA796FE6" box="[1316,1423,1184,1203]" country="Costa Rica" name="Guanacaste" pageId="17" pageNumber="18">Guanacaste</collectingRegion>
</emphasis>
:
<collectingMunicipality id="C36FACB5FF9C0112FCBFF69CFC646F84" box="[809,914,1214,1233]" pageId="17" pageNumber="18">Santa Rosa</collectingMunicipality>
(NP,
<location id="266B6014FF9C0112FC52F69FFB916F85" LSID="urn:lsid:plazi:treatment:AB1D87D9FF820112FCF0F401FC076D99:266B6014FF9C0112FC52F69FFB916F85" box="[964,1127,1213,1232]" country="Costa Rica" latitude="10.81" longLatPrecision="778" longitude="-85.62" municipality="Santa Rosa" name="Research Station" pageId="17" pageNumber="18" stateProvince="Guanacaste">Research Station</location>
308.1,
<geoCoordinate id="46805008FF9C0112FB38F69CFB036F84" box="[1198,1269,1214,1233]" degrees="10.81" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="555" value="10.81">10.81N</geoCoordinate>
<geoCoordinate id="46805008FF9C0112FB6DF69FFABA6F85" box="[1275,1356,1213,1232]" degrees="85.62" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="555" value="-85.62">85.62W</geoCoordinate>
),
<date id="570A100FFF9C0112FACDF69CFCAA6FBB" pageId="17" pageNumber="18" value="2008-12-07">
<collectingDate id="474EE9E7FF9C0112FACDF69CFCAA6FBB" pageId="17" pageNumber="18" value="2008-12-07">7-XII-2008</collectingDate>
</date>
,
<collectorName id="8E415319FF9C0112FCF0F6F9FC336FBB" box="[870,965,1243,1262]" pageId="17" pageNumber="18">Weirauch</collectorName>
&amp;
<collectorName id="8E415319FF9C0112FC72F6FEFBD56FBA" box="[996,1059,1244,1263]" pageId="17" pageNumber="18">Forero</collectorName>
leg. (deposited in:
<collectionCode id="45A5AE0AFF9C0112FB74F6F9FAD76FBB" box="[1250,1313,1243,1262]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="17" pageNumber="18" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="93DC3C92FF9C0112FAA6F6FEFC026E7F" collectingDate="1977-07-28" collectionCode="DARC" collectorName="O'Brien" country="Honduras" location="Zamorano" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="El Paraiso" typeStatus="paratype">
<typeStatus id="FC0F886DFF9C0112FAA6F6FEFA676FBA" box="[1328,1425,1244,1263]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="5BA3765FFF9C0112FCBFF6DBFC596E59" box="[809,943,1273,1292]" name="Honduras" pageId="17" pageNumber="18">HONDURAS</collectingCountry>
:
<emphasis id="11C0EADDFF9C0112FC2AF6DBFBD56E59" box="[956,1059,1273,1292]" italics="true" pageId="17" pageNumber="18">
<collectingRegion id="E170F82DFF9C0112FC2AF6DBFBD56E59" box="[956,1059,1273,1292]" country="Honduras" name="El Paraiso" pageId="17" pageNumber="18">El Paraiso</collectingRegion>
</emphasis>
:
<location id="266B6014FF9C0112FBA0F6D8FB536E58" LSID="urn:lsid:plazi:treatment:AB1D87D9FF820112FCF0F401FC076D99:266B6014FF9C0112FBA0F6D8FB536E58" box="[1078,1189,1274,1293]" country="Honduras" name="Zamorano" pageId="17" pageNumber="18" stateProvince="El Paraiso">Zamorano</location>
,
<specimenCount id="35B2FD46FF9C0112FB25F6DBFAF76E59" box="[1203,1281,1273,1292]" count="1" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
,
<date id="570A100FFF9C0112FA99F6DBFA666E59" box="[1295,1424,1273,1292]" pageId="17" pageNumber="18" value="1977-07-28">
<collectingDate id="474EE9E7FF9C0112FA99F6DBFA666E59" box="[1295,1424,1273,1292]" pageId="17" pageNumber="18" value="1977-07-28">28-VII-1977</collectingDate>
</date>
,
<collectorName id="8E415319FF9C0112FCBFF735FC836E7F" box="[809,885,1303,1322]" pageId="17" pageNumber="18">O’Brien</collectorName>
leg. (
<collectionCode id="45A5AE0AFF9C0112FC3AF735FC196E7F" box="[940,1007,1303,1322]" pageId="17" pageNumber="18">DARC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="93DC3C92FF9C0112FC69F735FC926E1D" collectionCode="CAS" country="Costa Rica" location="COSTA RICA" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="El Paraiso" typeStatus="paratype">
<collectingCountry id="5BA3765FFF9C0112FC69F735FB726E7F" box="[1023,1156,1303,1322]" name="Costa Rica" pageId="17" pageNumber="18">COSTA RICA</collectingCountry>
: (C.a. E.P.),
<specimenCount id="35B2FD46FF9C0112FB6FF735FAC96E7F" box="[1273,1343,1303,1322]" count="1" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
, no date (
<collectionCode id="45A5AE0AFF9C0112FCA6F717FCA86E1D" box="[816,862,1333,1352]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="18" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="93DC3C92FF9C0112FCF9F717FAD66E33" collectingDate="1969-06-17" collectingDateMax="1969-06-27" collectingDateMin="1969-06-17" collectionCode="AMNH" collectorName="Toby Schuh &amp; Janet Crane" country="Costa Rica" location="Finca La Taboga" municipality="Las Canas" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<emphasis id="11C0EADDFF9C0112FCF9F717FC2D6E1D" box="[879,987,1333,1352]" italics="true" pageId="17" pageNumber="18">
<collectingRegion id="E170F82DFF9C0112FCF9F717FC2D6E1D" box="[879,987,1333,1352]" country="Costa Rica" name="Guanacaste" pageId="17" pageNumber="18">Guanacaste</collectingRegion>
</emphasis>
:
<collectingMunicipality id="C36FACB5FF9C0112FC7EF717FBBA6E1D" box="[1000,1100,1333,1352]" pageId="17" pageNumber="18">Las Cañas</collectingMunicipality>
(
<location id="266B6014FF9C0112FBCCF717FAF66E1C" LSID="urn:lsid:plazi:treatment:AB1D87D9FF820112FCF0F401FC076D99:266B6014FF9C0112FBCCF717FAF66E1C" box="[1114,1280,1333,1353]" country="Costa Rica" municipality="Las Canas" name="Finca La Taboga" pageId="17" pageNumber="18" stateProvince="Guanacaste">Finca La Taboga</location>
),
<specimenCount id="35B2FD46FF9C0112FA87F717FA9C6E1D" box="[1297,1386,1333,1352]" count="1" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
,
<date id="570A100FFF9C0112FAE3F717FC6F6E33" pageId="17" pageNumber="18" value="1969-06-17" valueMax="1969-06-27" valueMin="1969-06-17">
<collectingDate id="474EE9E7FF9C0112FAE3F717FC6F6E33" pageId="17" pageNumber="18" value="1969-06-17" valueMax="1969-06-27" valueMin="1969-06-17">17- 27-VI-1969</collectingDate>
</date>
,
<collectorName id="8E415319FF9C0112FC32F770FBEA6E33" box="[932,1052,1362,1382]" pageId="17" pageNumber="18">Toby Schuh</collectorName>
,
<collectorName id="8E415319FF9C0112FBB3F771FB6C6E33" box="[1061,1178,1363,1382]" pageId="17" pageNumber="18">Janet Crane</collectorName>
leg. (
<collectionCode id="45A5AE0AFF9C0112FB46F771FAEC6E33" box="[1232,1306,1363,1382]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="17" pageNumber="18">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="93DC3C92FF9C0112FABCF771FADE6EF7" collectingDate="2008-12-06" collectionCode="UCRC" collectorName="Weirauch &amp; Forero" country="Costa Rica" elevation="308" latitude="10.84" location="Research Station" longLatPrecision="778" longitude="-85.62" municipality="Santa Rosa" pageId="17" pageNumber="18" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Guanacaste" typeStatus="paratype">
<collectingMunicipality id="C36FACB5FF9C0112FABCF771FA636E33" box="[1322,1429,1363,1382]" pageId="17" pageNumber="18">Santa Rosa</collectingMunicipality>
(NP,
<location id="266B6014FF9C0112FCC1F753FC0B6ED1" LSID="urn:lsid:plazi:treatment:AB1D87D9FF820112FCF0F401FC076D99:266B6014FF9C0112FCC1F753FC0B6ED1" box="[855,1021,1393,1412]" country="Costa Rica" latitude="10.84" longLatPrecision="778" longitude="-85.62" municipality="Santa Rosa" name="Research Station" pageId="17" pageNumber="18" stateProvince="Guanacaste">Research Station</location>
,
<quantity id="E44C9B2AFF9C0112FB91F753FBB46ED1" box="[1031,1090,1393,1412]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="17" pageNumber="18" unit="m" value="308.0">
<elevation id="A899D1FCFF9C0112FB91F753FBB46ED1" box="[1031,1090,1393,1412]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="17" pageNumber="18" unit="m" value="308.0">308m</elevation>
</quantity>
,
<geoCoordinate id="46805008FF9C0112FBDAF753FB6A6ED1" box="[1100,1180,1393,1412]" degrees="10.84" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="555" value="10.84">10.84°N</geoCoordinate>
<geoCoordinate id="46805008FF9C0112FB34F753FB0C6ED1" box="[1186,1274,1393,1412]" degrees="85.62" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="555" value="-85.62">85.62°W</geoCoordinate>
),
<specimenCount id="35B2FD46FF9C0112FA9DF753FAA16ED1" box="[1291,1367,1393,1412]" count="2" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
and
<specimenCount id="35B2FD46FF9C0112FA1FF753FC836EF4" count="4" pageId="17" pageNumber="18" type="female">4 females</specimenCount>
,
<date id="570A100FFF9C0112FCEAF7ACFC056EF4" box="[892,1011,1422,1441]" pageId="17" pageNumber="18" value="2008-12-06">
<collectingDate id="474EE9E7FF9C0112FCEAF7ACFC056EF4" box="[892,1011,1422,1441]" pageId="17" pageNumber="18" value="2008-12-06">06-XII-2008</collectingDate>
</date>
,
<collectorName id="8E415319FF9C0112FC6DF7ACFBAE6EF4" box="[1019,1112,1422,1441]" pageId="17" pageNumber="18">Weirauch</collectorName>
&amp;
<collectorName id="8E415319FF9C0112FBE7F7ADFB596EF7" box="[1137,1199,1423,1442]" pageId="17" pageNumber="18">Forero</collectorName>
leg. (
<collectionCode id="45A5AE0AFF9C0112FB76F7ADFAD46EF7" box="[1248,1314,1423,1442]" country="USA" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="17" pageNumber="18" type="University or college">UCRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="93DC3C92FF9C0112FAB9F7ACFA666E88" collectingDate="2008-12-04" collectionCode="UFRG" collectorName="Weirauch &amp; Forero" country="Costa Rica" elevation="26" latitude="10.35" location="Nr" longLatPrecision="779" longitude="-85.35" municipality="Palo Verde National Park" pageId="17" pageNumber="18" specimenCount="11" specimenCount-female="8" specimenCount-male="3" stateProvince="Guanacaste" typeStatus="paratype">
<collectingMunicipality id="C36FACB5FF9C0112FAB9F7ACFC5C6EEA" pageId="17" pageNumber="18">Palo Verde National Park</collectingMunicipality>
(
<location id="266B6014FF9C0112FC22F78FFC386E95" LSID="urn:lsid:plazi:treatment:AB1D87D9FF820112FCF0F401FC076D99:266B6014FF9C0112FC22F78FFC386E95" box="[948,974,1453,1472]" country="Costa Rica" latitude="10.35" longLatPrecision="779" longitude="-85.35" municipality="Palo Verde National Park" name="Nr" pageId="17" pageNumber="18" stateProvince="Guanacaste">Nr</location>
OTS research station
<quantity id="E44C9B2AFF9C0112FB0FF78EFB316EEA" box="[1177,1223,1452,1471]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="17" pageNumber="18" unit="m" value="26.0">
<elevation id="A899D1FCFF9C0112FB0FF78EFB316EEA" box="[1177,1223,1452,1471]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="17" pageNumber="18" unit="m" value="26.0">26m</elevation>
</quantity>
,
<geoCoordinate id="46805008FF9C0112FB59F78EFAEB6EEA" box="[1231,1309,1452,1471]" degrees="10.35" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="555" value="10.35">10.35°N</geoCoordinate>
<geoCoordinate id="46805008FF9C0112FAB7F78EFA8E6EEA" box="[1313,1400,1452,1471]" degrees="85.35" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="555" value="-85.35">85.35°W</geoCoordinate>
),
<specimenCount id="35B2FD46FF9C0112FA10F78FFCA96E88" count="3" pageId="17" pageNumber="18" type="male">3 males</specimenCount>
and
<specimenCount id="35B2FD46FF9C0112FC1BF7E8FC1F6E88" box="[909,1001,1482,1501]" count="8" pageId="17" pageNumber="18" type="female">8 females</specimenCount>
,
<date id="570A100FFF9C0112FC67F7E8FBAA6E88" box="[1009,1116,1482,1501]" pageId="17" pageNumber="18" value="2008-12-04">
<collectingDate id="474EE9E7FF9C0112FC67F7E8FBAA6E88" box="[1009,1116,1482,1501]" pageId="17" pageNumber="18" value="2008-12-04">4-XII-2008</collectingDate>
</date>
,
<collectorName id="8E415319FF9C0112FBF2F7E8FB376E88" box="[1124,1217,1482,1501]" pageId="17" pageNumber="18">Weirauch</collectorName>
&amp;
<collectorName id="8E415319FF9C0112FB4DF7E9FAEF6E8B" box="[1243,1305,1483,1502]" pageId="17" pageNumber="18">Forero</collectorName>
leg. (
<collectionCode id="45A5AE0AFF9C0112FADCF7E8FA7F6E88" box="[1354,1417,1482,1501]" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="17" pageNumber="18">UFRG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="93DC3C92FF9C0112FCB1F7CAFC126D4C" collectingDate="1953-12" collectionCode="AMNH" collectorName="Collector &amp; N. L. H. Kraus" country="Panama" location="Taboga" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Los Santos" typeStatus="paratype">
<collectingCountry id="5BA3765FFF9C0112FCB1F7CAFC746EAE" box="[807,898,1512,1531]" name="Panama" pageId="17" pageNumber="18">PANAMA</collectingCountry>
,
<emphasis id="11C0EADDFF9C0112FC18F7CBFC1B6EAE" box="[910,1005,1512,1531]" italics="true" pageId="17" pageNumber="18">
<collectingRegion id="E170F82DFF9C0112FC18F7CBFC1B6EAE" box="[910,1005,1512,1531]" country="Panama" name="Los Santos" pageId="17" pageNumber="18">Los Santos</collectingRegion>
</emphasis>
:
<location id="266B6014FF9C0112FC6EF7CAFBC96EA9" LSID="urn:lsid:plazi:treatment:AB1D87D9FF820112FCF0F401FC076D99:266B6014FF9C0112FC6EF7CAFBC96EA9" box="[1016,1087,1512,1532]" country="Panama" name="Taboga" pageId="17" pageNumber="18" stateProvince="Los Santos">Taboga</location>
,
<specimenCount id="35B2FD46FF9C0112FBD1F7CAFB7F6EAE" box="[1095,1161,1512,1531]" count="1" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
,
<date id="570A100FFF9C0112FB07F7CAFB1F6EAE" box="[1169,1257,1512,1531]" pageId="17" pageNumber="18" value="1953-12">
<collectingDate id="474EE9E7FF9C0112FB07F7CAFB1F6EAE" box="[1169,1257,1512,1531]" pageId="17" pageNumber="18" value="1953-12">XII-1953</collectingDate>
</date>
,
<collectorName id="8E415319FF9C0112FB67F7CAFAB86EAE" box="[1265,1358,1512,1531]" pageId="17" pageNumber="18">Collector</collectorName>
:
<collectorName id="8E415319FF9C0112FAC3F7CBFCA86D4C" pageId="17" pageNumber="18">N.L.H. Kraus</collectorName>
leg. (
<collectionCode id="45A5AE0AFF9C0112FC02F424FC296D4C" box="[916,991,1542,1561]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="17" pageNumber="18">AMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="230B36CFFF9C0112FCF3F406FB936D00" blockId="17.[807,1430,198,1740]" pageId="17" pageNumber="18">
Distribution (
<figureCitation id="BB8F2A4AFF9C0112FC78F401FBC36D62" box="[1006,1077,1571,1591]" captionStart="Figure 78" captionStartId="22.[159,217,936,956]" captionTargetBox="[164,1434,198,913]" captionTargetId="figure-21@22.[159,1436,197,916]" captionTargetPageId="22" captionText="Figure 78. Distributional map for studied specimens of the species of Lineostethus. Countries with species records are in green." pageId="17" pageNumber="18">Fig. 78</figureCitation>
).
<collectingCountry id="5BA3765FFF9C0112FBDEF406FB5B6D62" box="[1096,1197,1572,1591]" name="Honduras" pageId="17" pageNumber="18">Honduras</collectingCountry>
:
<emphasis id="11C0EADDFF9C0112FB2EF406FAE16D63" box="[1208,1303,1572,1591]" italics="true" pageId="17" pageNumber="18">
<collectingRegion id="E170F82DFF9C0112FB2EF406FAE16D63" box="[1208,1303,1572,1591]" country="Honduras" name="El Paraiso" pageId="17" pageNumber="18">El Paraiso</collectingRegion>
</emphasis>
;
<collectingCountry id="5BA3765FFF9C0112FAB3F406FA646D62" box="[1317,1426,1572,1591]" name="Costa Rica" pageId="17" pageNumber="18">Costa Rica</collectingCountry>
:
<emphasis id="11C0EADDFF9C0112FCBFF460FC636D00" box="[809,917,1602,1621]" italics="true" pageId="17" pageNumber="18">
<collectingRegion id="E170F82DFF9C0112FCBFF460FC636D00" box="[809,917,1602,1621]" country="Costa Rica" name="Guanacaste" pageId="17" pageNumber="18">Guanacaste</collectingRegion>
</emphasis>
;
<collectingCountry id="5BA3765FFF9C0112FC37F460FC056D00" box="[929,1011,1602,1621]" name="Panama" pageId="17" pageNumber="18">Panama</collectingCountry>
:
<emphasis id="11C0EADDFF9C0112FC6BF460FBA86D00" box="[1021,1118,1602,1621]" italics="true" pageId="17" pageNumber="18">
<collectingRegion id="E170F82DFF9C0112FC6BF460FBA86D00" box="[1021,1118,1602,1621]" country="Panama" name="Los Santos" pageId="17" pageNumber="18">Los Santos</collectingRegion>
</emphasis>
.
</paragraph>
<paragraph id="230B36CFFF9C0112FCF3F47DFC076D99" blockId="17.[807,1430,198,1740]" pageId="17" pageNumber="18">
Etymology. The epithet refers to the shape of the apex of each projection of the superior layer of ventral rim of pygophore; Latin adjective masculine,
<taxonomicName id="E4B44D4CFF9C0112FB23F4BEFAD76DFB" authorityName="Garbelotto &amp; Kochenborger &amp; Campos" authorityYear="2018" box="[1205,1313,1692,1710]" class="Insecta" family="Pentatomidae" genus="Lineostethus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acuminatus">
<emphasis id="11C0EADDFF9C0112FB23F4BEFAD76DFB" box="[1205,1313,1692,1710]" italics="true" pageId="17" pageNumber="18">acuminatus</emphasis>
</taxonomicName>
(
<emphasis id="11C0EADDFF9C0112FAA7F4BEFAB26DFB" box="[1329,1348,1692,1710]" italics="true" pageId="17" pageNumber="18">-a</emphasis>
,
<emphasis id="11C0EADDFF9C0112FAC5F4BEFA8E6DFB" box="[1363,1400,1692,1710]" italics="true" pageId="17" pageNumber="18">-um</emphasis>
) = pointed, sharpened.
</paragraph>
</subSubSection>
</treatment>
</document>
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