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<mods:title id="3F4A595FF211117FBAA11DF67BF053F5">On two new species of arboreal crabs from phytotelms in Sarawak, Borneo (Crustacea: Brachyura: Gecarcinucidae: Arachnothelphusa)</mods:title>
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<paragraph id="207AA81DFE7FFFCCFF0F2129E6BAFD95" blockId="1.[151,493,582,609]" box="[151,493,582,609]" pageId="1" pageNumber="408">
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<taxonomicName id="E7C5D39EFE7FFFCCFF0F2129E6BAFD95" ID-CoL="7NQS2" authority="Ng, 1991" authorityName="Ng" authorityYear="1991" box="[151,493,582,609]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="12B1740FFE7FFFCCFF0F2129E6BAFD95" bold="true" box="[151,493,582,609]" pageId="1" pageNumber="408">
<emphasis id="12B1740FFE7FFFCCFF0F2129E62FFD94" bold="true" box="[151,376,582,609]" italics="true" pageId="1" pageNumber="408">Arachnothelphusa</emphasis>
<bibRefCitation id="4454D5ECFE7FFFCCFEE72128E6BAFD95" author="Ng, P. K. L." box="[383,493,582,609]" pageId="1" pageNumber="408" pagination="1 - 12" refId="ref6389" refString="Ng, P. K. L. (1991) Bornean freshwater crabs of the genus Arachnothelphusa gen. nov. (Crustacea: Decapoda: Brachyura: Gecarcinucidae). Zoologische Mededelingen, 65 (1), 1 - 12." type="journal article" year="1991">Ng, 1991</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="207AA81DFE7FFFCCFF0F21E0E3CCFD5D" blockId="1.[151,1179,654,681]" box="[151,1179,654,681]" pageId="1" pageNumber="408">
<emphasis id="12B1740FFE7FFFCCFF0F21E0E66EFD5D" bold="true" box="[151,313,654,681]" pageId="1" pageNumber="408">
<typeStatus id="FF7E16BFFE7FFFCCFF0F21E0E783FD5C" box="[151,212,655,681]" pageId="1" pageNumber="408">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="E7C5D39EFE7FFFCCFED921E0E430FD5D" authority="De Man, 1899" authorityName="De Man" authorityYear="1899" box="[321,871,654,681]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="melanippe" subGenus="Potamon">
<emphasis id="12B1740FFE7FFFCCFED921E0E6FCFD5D" box="[321,427,655,680]" italics="true" pageId="1" pageNumber="408">Potamon</emphasis>
<emphasis id="12B1740FFE7FFFCCFE2A21E1E6EBFD5D" bold="true" box="[434,444,654,680]" pageId="1" pageNumber="408">(</emphasis>
<emphasis id="12B1740FFE7FFFCCFE2421E0E571FD5D" box="[444,550,655,680]" italics="true" pageId="1" pageNumber="408">Potamon</emphasis>
<emphasis id="12B1740FFE7FFFCCFDBE21E1E567FD5D" bold="true" box="[550,560,654,680]" pageId="1" pageNumber="408">)</emphasis>
<emphasis id="12B1740FFE7FFFCCFDAF21E0E5E6FD5C" box="[567,689,655,681]" italics="true" pageId="1" pageNumber="408">melanippe</emphasis>
<emphasis id="12B1740FFE7FFFCCFD2021E0E430FD5D" bold="true" box="[696,871,654,681]" pageId="1" pageNumber="408">
De
<bibRefCitation id="4454D5ECFE7FFFCCFD7A21E0E430FD5D" author="Man, J. G. de" box="[738,871,654,681]" pageId="1" pageNumber="408" pagination="53 - 144" refId="ref5941" refString="Man, J. G. de (1899) Zoological Results of the Dutch Expedition to Central Borneo. The Crustacea. Part II, Brachyura. Notes Leyden Museum, 21, 53 - 144, pls. 5 - 12." type="journal article" year="1899">Man, 1899</bibRefCitation>
</emphasis>
</taxonomicName>
<emphasis id="12B1740FFE7FFFCCFCF021E1E3CCFD5D" bold="true" box="[872,1179,654,681]" pageId="1" pageNumber="408">, by original designation.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="68DFFB96FE7FFFCFFF0F2194E5E7FE7C" lastPageId="2" lastPageNumber="409" pageId="1" pageNumber="408" type="discussion">
<paragraph id="207AA81DFE7FFFCCFF0F2194E261FBE4" blockId="1.[151,1437,763,2013]" pageId="1" pageNumber="408">
<emphasis id="12B1740FFE7FFFCCFF0F2194E653FCE0" bold="true" box="[151,260,763,789]" pageId="1" pageNumber="408">Remarks</emphasis>
.
<bibRefCitation id="4454D5ECFE7FFFCCFE8B2194E6C1FCE0" author="Bott, R." box="[275,406,763,789]" pageId="1" pageNumber="408" pagination="1 - 338" refId="ref5190" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 526, 1 - 338." type="journal article" year="1970">Bott (1970)</bibRefCitation>
treated
<taxonomicName id="E7C5D39EFE7FFFCCFE6B2194E329FCE0" authority="Borradaile, 1900" authorityName="Borradaile" authorityYear="1900" box="[499,1150,763,789]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="kadamaianum" subGenus="Geothelphusa">
<emphasis id="12B1740FFE7FFFCCFE6B2194E50FFCE1" box="[499,600,763,788]" italics="true" pageId="1" pageNumber="408">Potamon</emphasis>
(
<emphasis id="12B1740FFE7FFFCCFDF22194E451FCE0" box="[618,774,763,789]" italics="true" pageId="1" pageNumber="408">Geothelphusa</emphasis>
)
<emphasis id="12B1740FFE7FFFCCFC8F2194E4E1FCE0" box="[791,950,763,789]" italics="true" pageId="1" pageNumber="408">kadamaianum</emphasis>
<bibRefCitation id="4454D5ECFE7FFFCCFC262194E329FCE0" author="Borradaile, L. A." box="[958,1150,763,789]" pageId="1" pageNumber="408" pagination="93 - 95" refId="ref5234" refString="Borradaile, L. A. (1900) On a small collection of decapod crustaceans from freshwater in North Borneo. Proceedings of the Zoological Society of London, 1900, 93 - 95. https: // doi. org / 10.1111 / j. 1096 - 3642.1890. tb 01706. x" type="journal article" year="1900">Borradaile, 1900</bibRefCitation>
</taxonomicName>
, as a subspecies of
<taxonomicName id="E7C5D39EFE7FFFCCFAC62194E539FCCC" authority="De Man, 1899" authorityName="De Man" authorityYear="1899" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="melanippe" subGenus="Potamon">
<emphasis id="12B1740FFE7FFFCCFAC62194E790FCCD" italics="true" pageId="1" pageNumber="408">Potamon</emphasis>
(
<emphasis id="12B1740FFE7FFFCCFF402070E66AFCCD" box="[216,317,799,824]" italics="true" pageId="1" pageNumber="408">Potamon</emphasis>
)
<emphasis id="12B1740FFE7FFFCCFED52070E695FCCC" box="[333,450,799,825]" italics="true" pageId="1" pageNumber="408">melanippe</emphasis>
De
<bibRefCitation id="4454D5ECFE7FFFCCFE692070E539FCCC" author="Man, J. G. de" box="[497,622,799,825]" pageId="1" pageNumber="408" pagination="53 - 144" refId="ref5941" refString="Man, J. G. de (1899) Zoological Results of the Dutch Expedition to Central Borneo. The Crustacea. Part II, Brachyura. Notes Leyden Museum, 21, 53 - 144, pls. 5 - 12." type="journal article" year="1899">Man, 1899</bibRefCitation>
</taxonomicName>
, placing both in
<taxonomicName id="E7C5D39EFE7FFFCCFCB02070E364FCCC" authority="Bott, 1969" authorityName="Bott" authorityYear="1969" box="[808,1075,799,825]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="12B1740FFE7FFFCCFCB02070E4E5FCCC" box="[808,946,799,825]" italics="true" pageId="1" pageNumber="408">Thelphusula</emphasis>
<bibRefCitation id="4454D5ECFE7FFFCCFC222070E364FCCC" author="Bott, R." box="[954,1075,799,825]" pageId="1" pageNumber="408" pagination="359 - 366" refId="ref5157" refString="Bott, R. (1969) Flusskrabben aus Asien und ihre Klassifikation (Crustacea, Decapoda). Senckenbergiana biologica, 50 (5 - 6), 359 - 366." type="journal article" year="1969">Bott, 1969</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="4454D5ECFE7FFFCCFBA62070E3E5FCCC" author="Ng, P. K. L." box="[1086,1202,799,825]" pageId="1" pageNumber="408" pagination="1 - 12" refId="ref6389" refString="Ng, P. K. L. (1991) Bornean freshwater crabs of the genus Arachnothelphusa gen. nov. (Crustacea: Decapoda: Brachyura: Gecarcinucidae). Zoologische Mededelingen, 65 (1), 1 - 12." type="journal article" year="1991">Ng (1991)</bibRefCitation>
redefined the genus, establishing
<taxonomicName id="E7C5D39EFE7FFFCCFEBC202CE6A6FCA8" authorityName="Ng" authorityYear="1991" box="[292,497,835,861]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="12B1740FFE7FFFCCFEBC202CE6A6FCA8" box="[292,497,835,861]" italics="true" pageId="1" pageNumber="408">Arachnothelphusa</emphasis>
</taxonomicName>
for these two species because of their long ambulatory legs as well as their distinctive G1 and G2 structures. Of the known species of
<taxonomicName id="E7C5D39EFE7FFFCCFD722008E4E0FC74" authorityName="Ng" authorityYear="1991" box="[746,951,871,897]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="12B1740FFE7FFFCCFD722008E4E0FC74" box="[746,951,871,897]" italics="true" pageId="1" pageNumber="408">Arachnothelphusa</emphasis>
</taxonomicName>
,
<taxonomicName id="E7C5D39EFE7FFFCCFC5E2008E313FC75" authorityName="Ng" authorityYear="1991" box="[966,1092,871,897]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="terrapes">
<emphasis id="12B1740FFE7FFFCCFC5E2008E313FC75" box="[966,1092,871,897]" italics="true" pageId="1" pageNumber="408">A. terrapes</emphasis>
</taxonomicName>
is probably the most atypical, with the external orbital tooth separated from the epibranchial tooth by a deep U-shaped sinus with the entire frontal region sunken in, the G1 terminal segment prominently curves upwards and the G2 distal segment is very short (
<bibRefCitation id="4454D5ECFE7FFFCCFF0720BCE653FC18" author="Ng, P. K. L." box="[159,260,979,1005]" pageId="1" pageNumber="408" pagination="1 - 12" refId="ref6389" refString="Ng, P. K. L. (1991) Bornean freshwater crabs of the genus Arachnothelphusa gen. nov. (Crustacea: Decapoda: Brachyura: Gecarcinucidae). Zoologische Mededelingen, 65 (1), 1 - 12." type="journal article" year="1991">Ng 1991</bibRefCitation>
;
<bibRefCitation id="4454D5ECFE7FFFCCFE8820BCE6ECFC18" author="Ng, P. K. L. &amp; Ng, P. Y. C." box="[272,443,979,1005]" pageId="1" pageNumber="408" pagination="89 - 112" refId="ref6976" refString="Ng, P. K. L. &amp; Ng, P. Y. C. (2018) The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). Zookeys, 760, 89 - 112. https: // doi. org / 10.3897 / zookeys. 760.24787" type="journal article" year="2018">Ng &amp; Ng 2018</bibRefCitation>
). All other congeners have a more normal carapace with the frontal margin level with the anterolateral margin, the G1 terminal segment is conical and straight, and the G2 distal segment is longer.
</paragraph>
<paragraph id="207AA81DFE7FFFCCFF5F2774E38DFA8C" blockId="1.[151,1437,763,2013]" pageId="1" pageNumber="408">
On the basis of the height of the carapace,
<taxonomicName id="E7C5D39EFE7FFFCCFD012774E47BFBC0" authorityName="De Man" authorityYear="1899" box="[665,812,1051,1077]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="melanippe">
<emphasis id="12B1740FFE7FFFCCFD012774E47BFBC0" box="[665,812,1051,1077]" italics="true" pageId="1" pageNumber="408">A. melanippe</emphasis>
</taxonomicName>
,
<taxonomicName id="E7C5D39EFE7FFFCCFCA02774E4B7FBC0" baseAuthorityName="Borradaile" baseAuthorityYear="1900" box="[824,992,1051,1077]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="kadamaiana">
<emphasis id="12B1740FFE7FFFCCFCA02774E4B7FBC0" box="[824,992,1051,1077]" italics="true" pageId="1" pageNumber="408">A. kadamaiana</emphasis>
</taxonomicName>
,
<taxonomicName id="E7C5D39EFE7FFFCCFC752774E3F4FBC1" authorityName="Grinang, Pui &amp; Ng" authorityYear="2015" box="[1005,1187,1051,1077]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="merarapensis">
<emphasis id="12B1740FFE7FFFCCFC752774E3F4FBC1" box="[1005,1187,1051,1077]" italics="true" pageId="1" pageNumber="408">A. merarapensis</emphasis>
</taxonomicName>
and
<taxonomicName id="E7C5D39EFE7FFFCCFB4F2774E2CBFBC0" baseAuthorityName="Ng &amp; Goh" baseAuthorityYear="1987" box="[1239,1436,1051,1077]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="rhadamanthysi">
<emphasis id="12B1740FFE7FFFCCFB4F2774E2CBFBC0" box="[1239,1436,1051,1077]" italics="true" pageId="1" pageNumber="408">A. rhadamanthysi</emphasis>
</taxonomicName>
as well as the new species here,
<taxonomicName id="E7C5D39EFE7FFFCCFD902750E509FBAC" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[520,606,1087,1113]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="bako" status="sp. nov.">
<emphasis id="12B1740FFE7FFFCCFD902750E509FBAC" box="[520,606,1087,1113]" italics="true" pageId="1" pageNumber="408">A. bako</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0982C974FE7FFFCCFDFF2750E5F1FBAC" box="[615,678,1087,1113]" pageId="1" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
, belong to one group where it is high and the structure looks more inflated (e.g.,
<figureCitation id="B8FEB498FE7FFFCCFEAD270CE6C1FB88" box="[309,406,1123,1149]" captionStart="FIG" captionStartId="7.[151,196,1733,1757]" captionTargetBox="[151,1436,345,1709]" captionTargetId="figure-96@7.[151,1436,345,1709]" captionTargetPageId="7" captionText="FIG. 7. Arachnothelphusa bako n. sp., holotype female (19.0 × 14.2 mm) (ZRC 2002.0098), Bako. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, third maxillipeds, anterior thoracic sternum and pleon; F, posterior margin of epistome; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5222309" httpUri="https://zenodo.org/record/5222309/files/figure.png" pageId="1" pageNumber="408">Figs. 7C</figureCitation>
,
<figureCitation id="B8FEB498FE7FFFCCFE3B270CE690FB88" box="[419,455,1123,1149]" captionStart="FIG" captionStartId="8.[151,196,1946,1970]" captionTargetBox="[207,1380,677,1922]" captionTargetId="figure-48@8.[207,1380,677,1922]" captionTargetPageId="8" captionText="FIG. 9. Arachnothelphusa kadamaiana (Borradaile, 1900), holotype female (18.9 × 13.5 mm) (SMF 4281), Kadamian River.A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, third maxillipeds, anterior thoracic sternum and pleon; F, posterior margin of epistome; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5222313" httpUri="https://zenodo.org/record/5222313/files/figure.png" pageId="1" pageNumber="408">9C</figureCitation>
).
<taxonomicName id="E7C5D39EFE7FFFCCFE44270CE459FB89" authorityName="Ng" authorityYear="1991" box="[476,782,1123,1149]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="terrapes">
<emphasis id="12B1740FFE7FFFCCFE44270CE459FB89" box="[476,782,1123,1149]" italics="true" pageId="1" pageNumber="408">Arachnothelphusa terrapes</emphasis>
</taxonomicName>
and
<taxonomicName id="E7C5D39EFE7FFFCCFCDE270CE4FEFB88" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[838,937,1123,1149]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
<emphasis id="12B1740FFE7FFFCCFCDE270CE4FEFB88" box="[838,937,1123,1149]" italics="true" pageId="1" pageNumber="408">A. rimba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0982C974FE7FFFCCFC28270CE4BAFB88" box="[944,1005,1123,1149]" pageId="1" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
belong to the second group where the carapace is not as high and the structure appears less inflated (e.g.,
<figureCitation id="B8FEB498FE7FFFCCFC1127E8E488FB54" box="[905,991,1159,1185]" captionStart="FIG" captionStartId="2.[151,196,1945,1969]" captionTargetBox="[275,1312,882,1919]" captionTargetId="figure-290@2.[275,1312,882,1919]" captionTargetPageId="2" captionText="FIG. 1. Arachnothelphusa rimba n. sp., holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5222297" httpUri="https://zenodo.org/record/5222297/files/figure.png" pageId="1" pageNumber="408">Fig. 1C</figureCitation>
). The difference, however, does not appear to be a phylogenetic one as the character cannot be correlated with the male pleonal and gonopodal features, and is probably associated with the habits and habitat of individual species. The proportions of the ambulatory legs is a useful character but should only be used between specimens of the same sex. As noted by Grinang
<emphasis id="12B1740FFE7FFFCCFA95279CE212FAF8" box="[1293,1349,1267,1293]" italics="true" pageId="1" pageNumber="408">et al.</emphasis>
(2015), female specimens of
<taxonomicName id="E7C5D39EFE7FFFCCFE192678E560FAC5" authorityName="Grinang, Pui &amp; Ng" authorityYear="2015" box="[385,567,1303,1329]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="merarapensis">
<emphasis id="12B1740FFE7FFFCCFE192678E560FAC5" box="[385,567,1303,1329]" italics="true" pageId="1" pageNumber="408">A. merarapensis</emphasis>
</taxonomicName>
generally have relatively longer ambulatory legs than males. This same condition is observed in
<taxonomicName id="E7C5D39EFE7FFFCCFEE82654E54DFAA0" baseAuthorityName="Borradaile" baseAuthorityYear="1900" box="[368,538,1339,1365]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="kadamaiana">
<emphasis id="12B1740FFE7FFFCCFEE82654E54DFAA0" box="[368,538,1339,1365]" italics="true" pageId="1" pageNumber="408">A. kadamaiana</emphasis>
</taxonomicName>
; with the male P2P5 relatively shorter (e.g., length to width ratio of P4 merus 4.344.36;
<figureCitation id="B8FEB498FE7FFFCCFE8E2630E6C1FA8C" box="[278,406,1375,1401]" captionStart="FIG" captionStartId="9.[151,196,1447,1471]" captionTargetBox="[151,1436,181,1423]" captionTargetId="figure-15@9.[151,1436,181,1423]" captionTargetPageId="9" captionText="FIG. 10. Ambulatory legs. AD, Arachnothelphusa bako n. sp., holotype female (19.0 × 14.2 mm) (ZRC 2002.0098), Bako; E, A. kadamaiana (Borradaile, 1900), holotype female (18.9 × 13.5 mm) (SMF 4281), Kadamian River; F, A. kadamaiana (Borradaile, 1900), male (21.1 × 15.8 mm) (ZRC 2002.0097), Crocker Range; G, A. kadamaiana (Borradaile, 1900), male (22.8 × 16.5 mm) (ZRC 2002.0097), Crocker Range. A, left P4; B, left P5 merus; C, EG, right P4; D, right P5." figureDoi="http://doi.org/10.5281/zenodo.5222331" httpUri="https://zenodo.org/record/5222331/files/figure.png" pageId="1" pageNumber="408">Fig. 10F, G</figureCitation>
) whereas in females, the P4 merus length to width ratio is 5.45 (
<figureCitation id="B8FEB498FE7FFFCCFBF02630E399FA8C" box="[1128,1230,1375,1401]" captionStart="FIG" captionStartId="9.[151,196,1447,1471]" captionTargetBox="[151,1436,181,1423]" captionTargetId="figure-15@9.[151,1436,181,1423]" captionTargetPageId="9" captionText="FIG. 10. Ambulatory legs. AD, Arachnothelphusa bako n. sp., holotype female (19.0 × 14.2 mm) (ZRC 2002.0098), Bako; E, A. kadamaiana (Borradaile, 1900), holotype female (18.9 × 13.5 mm) (SMF 4281), Kadamian River; F, A. kadamaiana (Borradaile, 1900), male (21.1 × 15.8 mm) (ZRC 2002.0097), Crocker Range; G, A. kadamaiana (Borradaile, 1900), male (22.8 × 16.5 mm) (ZRC 2002.0097), Crocker Range. A, left P4; B, left P5 merus; C, EG, right P4; D, right P5." figureDoi="http://doi.org/10.5281/zenodo.5222331" httpUri="https://zenodo.org/record/5222331/files/figure.png" pageId="1" pageNumber="408">Fig. 10E</figureCitation>
).
</paragraph>
<paragraph id="207AA81DFE7FFFCFFF5F26EDE5E7FE7C" blockId="1.[151,1437,763,2013]" lastBlockId="2.[151,1437,151,393]" lastPageId="2" lastPageNumber="409" pageId="1" pageNumber="408">
<emphasis id="12B1740FFE7FFFCCFF5F26EDE683FA69" bold="true" box="[199,468,1410,1436]" pageId="1" pageNumber="408">Comparative material</emphasis>
.
<taxonomicName id="E7C5D39EFE7FFFCCFE7F26ECE4A8FA68" authority="(De Man, 1899)" baseAuthorityName="De Man" baseAuthorityYear="1899" box="[487,1023,1411,1437]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="melanippe">
<emphasis id="12B1740FFE7FFFCCFE7F26ECE463FA68" box="[487,820,1411,1437]" italics="true" pageId="1" pageNumber="408">Arachnothelphusa melanippe</emphasis>
(De
<bibRefCitation id="4454D5ECFE7FFFCCFCEC26ECE4A0FA68" author="Man, J. G. de" box="[884,1015,1411,1437]" pageId="1" pageNumber="408" pagination="53 - 144" refId="ref5941" refString="Man, J. G. de (1899) Zoological Results of the Dutch Expedition to Central Borneo. The Crustacea. Part II, Brachyura. Notes Leyden Museum, 21, 53 - 144, pls. 5 - 12." type="journal article" year="1899">Man, 1899</bibRefCitation>
)
</taxonomicName>
:
<materialsCitation id="90ADA240FE7FFFCCFB9626ECE67CFA10" collectionCode="NNM" collectorName="C Buttikofer" country="Indonesia" location="Liang Koeboeng Mountains" municipality="Liang Kubung" pageId="1" pageNumber="408" specimenCode="D1303" specimenCount="1" stateProvince="Kalimantan Utara" typeStatus="lectotype">
<typeStatus id="FF7E16BFFE7FFFCCFB9626ECE329FA68" box="[1038,1150,1411,1437]" pageId="1" pageNumber="408" type="lectotype">Lectotype</typeStatus>
male (18.9 ×
<quantity id="E73D05F8FE7FFFCCFAB326ECE2C3FA68" box="[1323,1428,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="1" pageNumber="408" unit="mm" value="14.4">14.4 mm</quantity>
) (
<collectionCode id="46D430D8FE7FFFCCFF0726C8E7B5FA34" box="[159,226,1447,1473]" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="408" type="Museum">NNM</collectionCode>
<specimenCode id="70630066FE7FFFCCFF7226C8E66DFA34" box="[234,314,1447,1473]" collectionCode="NNM" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="408" type="Museum">D1303</specimenCode>
a),
<location id="251AFEC6FE7FFFCCFEC326C8E5CFFA34" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7FFFCCFEC326C8E5CFFA34" box="[347,664,1447,1473]" country="Indonesia" municipality="Liang Kubung" name="Liang Koeboeng Mountains" pageId="1" pageNumber="408" stateProvince="Kalimantan Utara">Liang Koeboeng Mountains</location>
(= present day
<collectingMunicipality id="C01E3267FE7FFFCCFCD426C8E4A3FA37" box="[844,1012,1447,1474]" pageId="1" pageNumber="408">Liang Kubung</collectingMunicipality>
),
<collectingRegion id="E20166FFFE7FFFCCFB9026C8E3D9FA34" box="[1032,1166,1447,1473]" country="Indonesia" name="Kalimantan Utara" pageId="1" pageNumber="408">Kalimantan</collectingRegion>
,
<collectingCountry id="58D2E88DFE7FFFCCFB0226C8E25DFA34" box="[1178,1290,1447,1473]" name="Indonesia" pageId="1" pageNumber="408">Indonesia</collectingCountry>
, coll.
<collectorName id="8D30CDCBFE7FFFCCFAC826C8E7B3FA10" pageId="1" pageNumber="408">C Büttikofer</collectorName>
, 1897
</materialsCitation>
;
<materialsCitation id="90ADA240FE7FFFCCFEA026A4E556F9FC" collectionCode="NNM" collectorName="C Buttikofer" country="Indonesia" location="Liang Koeboeng Mountains" municipality="Liang Kubung" pageId="1" pageNumber="408" specimenCode="D1303" specimenCount="1" stateProvince="Kalimantan Utara" typeStatus="paralectotype">
<typeStatus id="FF7E16BFFE7FFFCCFEA026A4E698FA10" box="[312,463,1483,1509]" pageId="1" pageNumber="408" type="paralectotype">paralectotype</typeStatus>
female (21.4 ×
<quantity id="E73D05F8FE7FFFCCFD1D26A4E5BBFA10" box="[645,748,1483,1509]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="1" pageNumber="408" unit="mm" value="16.7">16.7 mm</quantity>
) (
<collectionCode id="46D430D8FE7FFFCCFC9B26A4E411FA10" box="[771,838,1483,1509]" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="408" type="Museum">NNM</collectionCode>
<specimenCode id="70630066FE7FFFCCFCD626A4E4C9FA13" box="[846,926,1483,1510]" collectionCode="NNM" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="408" type="Museum">D1303</specimenCode>
b), same data as lectotype.
<taxonomicName id="E7C5D39EFE7FFFCCFB5726A4E556F9FC" authority="(Borradaile, 1900)" baseAuthorityName="Borradaile" baseAuthorityYear="1900" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="kadamaiana">
<emphasis id="12B1740FFE7FFFCCFB5726A4E676F9FC" italics="true" pageId="1" pageNumber="408">Arachnothelphusa kadamaiana</emphasis>
(
<bibRefCitation id="4454D5ECFE7FFFCCFEAB2680E6AEF9FC" author="Borradaile, L. A." box="[307,505,1519,1545]" pageId="1" pageNumber="408" pagination="93 - 95" refId="ref5234" refString="Borradaile, L. A. (1900) On a small collection of decapod crustaceans from freshwater in North Borneo. Proceedings of the Zoological Society of London, 1900, 93 - 95. https: // doi. org / 10.1111 / j. 1096 - 3642.1890. tb 01706. x" type="journal article" year="1900">Borradaile, 1900</bibRefCitation>
)
</taxonomicName>
</materialsCitation>
:
<materialsCitation id="90ADA240FE7FFFCCFD972680E633F9D8" collectionCode="SMF" collectorName="East &amp; Borneo" country="Malaysia" location="Kadamian River" pageId="1" pageNumber="408" specimenCode="SMF 4281" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="FF7E16BFFE7FFFCCFD972680E520F9FC" box="[527,631,1519,1545]" pageId="1" pageNumber="408" type="holotype">Holotype</typeStatus>
female (18.9 ×
<quantity id="E73D05F8FE7FFFCCFCAD2681E4CAF9FC" box="[821,925,1518,1545]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="1" pageNumber="408" unit="mm" value="13.5">13.5 mm</quantity>
) (
<specimenCode id="70630066FE7FFFCCFC2E2680E361F9FC" box="[950,1078,1519,1545]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="408">SMF 4281</specimenCode>
),
<location id="251AFEC6FE7FFFCCFBD42680E25EF9FC" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7FFFCCFBD42680E25EF9FC" box="[1100,1289,1519,1545]" country="Malaysia" name="Kadamian River" pageId="1" pageNumber="408" stateProvince="Sabah">Kadamian River</location>
,
<collectingRegion id="E20166FFFE7FFFCCFA8F2680E209F9FC" box="[1303,1374,1519,1545]" country="Malaysia" name="Sabah" pageId="1" pageNumber="408">Sabah</collectingRegion>
,
<collectorName id="8D30CDCBFE7FFFCCFAF42680E2CBF9FC" box="[1388,1436,1519,1545]" pageId="1" pageNumber="408">East</collectorName>
<collectingCountry id="58D2E88DFE7FFFCCFF0F257CE657F9D8" box="[151,256,1555,1581]" name="Malaysia" pageId="1" pageNumber="408">Malaysia</collectingCountry>
,
<collectorName id="8D30CDCBFE7FFFCCFE96257CE633F9D8" box="[270,356,1555,1581]" pageId="1" pageNumber="408">Borneo</collectorName>
</materialsCitation>
;
<materialsCitation id="90ADA240FE7FFFCCFEE9257CE30DF9D8" box="[369,1114,1555,1581]" collectionCode="SMF" collectorName="East &amp; Borneo" country="Malaysia" location="Kadamian River" pageId="1" pageNumber="408" specimenCode="SMF 4282" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<specimenCount id="36C36394FE7FFFCCFEE9257CE6EAF9D8" box="[369,445,1555,1581]" pageId="1" pageNumber="408" type="male">1 male</specimenCount>
(20.1 ×
<quantity id="E73D05F8FE7FFFCCFDBA257CE5DEF9D8" box="[546,649,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="1" pageNumber="408" unit="mm" value="14.9">14.9 mm</quantity>
) (
<specimenCode id="70630066FE7FFFCCFD39257CE477F9D8" box="[673,800,1555,1581]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="408">SMF 4282</specimenCode>
), same locality as holotype
</materialsCitation>
;
<materialsCitation id="90ADA240FE7FFFCCFBFE257CE262F9A4" collectingDate="1992-08-12" collectionCode="ZRC" collectorName="R. F. Inger" country="Malaysia" county="Borneo" location="Poring" municipality="Basin" pageId="1" pageNumber="408" specimenCode="ZRC 2009.0094" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
<specimenCount id="36C36394FE7FFFCCFBFE257CE39EF9D8" box="[1126,1225,1555,1581]" pageId="1" pageNumber="408" type="female">1 female</specimenCount>
(23.2 ×
<quantity id="E73D05F8FE7FFFCCFAB6257CE2C3F9D8" box="[1326,1428,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" pageId="1" pageNumber="408" unit="mm" value="17.1">17.1 mm</quantity>
) (
<specimenCode id="70630066FE7FFFCCFF072558E60AF9A4" box="[159,349,1591,1617]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="408">ZRC 2009.0094</specimenCode>
),
<location id="251AFEC6FE7FFFCCFEEA2558E697F9A4" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7FFFCCFEEA2558E697F9A4" box="[370,448,1591,1617]" country="Malaysia" county="Borneo" municipality="Basin" name="Poring" pageId="1" pageNumber="408" stateProvince="Sabah">Poring</location>
,
<collectingMunicipality id="C01E3267FE7FFFCCFE552558E55AF9A4" box="[461,525,1591,1617]" pageId="1" pageNumber="408">Basin</collectingMunicipality>
1A,
<collectingRegion id="E20166FFFE7FFFCCFDD12558E5C7F9A4" box="[585,656,1591,1617]" country="Malaysia" name="Sabah" pageId="1" pageNumber="408">Sabah</collectingRegion>
,
<collectingCountry id="58D2E88DFE7FFFCCFD062558E450F9A4" box="[670,775,1591,1617]" name="Malaysia" pageId="1" pageNumber="408">Malaysia</collectingCountry>
,
<collectingCounty id="C91BD091FE7FFFCCFC8E2558E43CF9A4" box="[790,875,1591,1617]" pageId="1" pageNumber="408">Borneo</collectingCounty>
, coll.
<collectorName id="8D30CDCBFE7FFFCCFC2B2558E371F9A4" box="[947,1062,1591,1617]" pageId="1" pageNumber="408">R.F. Inger</collectorName>
<emphasis id="12B1740FFE7FFFCCFBA82558E33CF9A4" box="[1072,1131,1591,1617]" italics="true" pageId="1" pageNumber="408">et al.</emphasis>
,
<date id="547B8EDDFE7FFFCCFBE32558E262F9A4" box="[1147,1333,1591,1617]" pageId="1" pageNumber="408" value="1992-08-12">
<collectingDate id="443F7735FE7FFFCCFBE32558E262F9A4" box="[1147,1333,1591,1617]" pageId="1" pageNumber="408" value="1992-08-12">12 August 1992</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="90ADA240FE7FFFCCFADB2558E6EDF96C" collectingDate="2001-04-24" collectionCode="ZRC" collectorName="I. Das" country="Malaysia" latitude="5.45" location="Crocker Range" longLatPrecision="1305" longitude="116.05" pageId="1" pageNumber="408" specimenCode="ZRC 2002.0097" specimenCount="3" specimenCount-male="3" stateProvince="Sabah" typeStatus="holotype">
<specimenCount id="36C36394FE7FFFCCFADB2558E2CBF9A4" box="[1347,1436,1591,1617]" pageId="1" pageNumber="408" type="male">3 males</specimenCount>
(21.1 ×
<quantity id="E73D05F8FE7FFFCCFF692535E60EF980" box="[241,345,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="1" pageNumber="408" unit="mm" value="15.8">15.8 mm</quantity>
, 22.8 ×
<quantity id="E73D05F8FE7FFFCCFE2E2535E54AF980" box="[438,541,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="408" unit="mm" value="16.5">16.5 mm</quantity>
, 25.3 ×
<quantity id="E73D05F8FE7FFFCCFDE32535E5B6F980" box="[635,737,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="408" unit="mm" value="18.5">18.5 mm</quantity>
) (
<specimenCode id="70630066FE7FFFCCFD612534E4E2F980" box="[761,949,1627,1653]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="408">ZRC 2002.0097</specimenCode>
),
<location id="251AFEC6FE7FFFCCFC502534E322F980" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7FFFCCFC502534E322F980" box="[968,1141,1627,1653]" country="Malaysia" latitude="5.45" longLatPrecision="1305" longitude="116.05" name="Crocker Range" pageId="1" pageNumber="408" stateProvince="Sabah">Crocker Range</location>
,
<collectingRegion id="E20166FFFE7FFFCCFB192534E39FF980" box="[1153,1224,1627,1653]" country="Malaysia" name="Sabah" pageId="1" pageNumber="408">Sabah</collectingRegion>
,
<geoCoordinate id="45F1CEDAFE7FFFCCFB4D2535E272F980" box="[1237,1317,1626,1653]" degrees="5" direction="north" minutes="27" orientation="latitude" pageId="1" pageNumber="408" precision="925" value="5.45">5º27N</geoCoordinate>
<geoCoordinate id="45F1CEDAFE7FFFCCFAB52534E2CFF980" box="[1325,1432,1627,1653]" degrees="116" direction="east" minutes="03" orientation="longitude" pageId="1" pageNumber="408" precision="925" value="116.05">116º03E</geoCoordinate>
, coll.
<collectorName id="8D30CDCBFE7FFFCCFF562510E644F96C" box="[206,275,1663,1689]" pageId="1" pageNumber="408">I. Das</collectorName>
,
<date id="547B8EDDFE7FFFCCFE852510E6EDF96C" box="[285,442,1663,1689]" pageId="1" pageNumber="408" value="2001-04-24">
<collectingDate id="443F7735FE7FFFCCFE852510E6EDF96C" box="[285,442,1663,1689]" pageId="1" pageNumber="408" value="2001-04-24">24 April 2001</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="90ADA240FE7FFFCCFE5C2510E6DFF914" collectingDate="1986-03-25" collectionCode="ZRC" collectorName="P. Chapman" country="Malaysia" county="Borneo" latitude="5.55" location="Simud Puteh Cave" longLatPrecision="1305" longitude="118.1" municipality="East" pageId="1" pageNumber="408" specimenCode="ZRC 1990.0443" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<taxonomicName id="E7C5D39EFE7FFFCCFE5C2510E342F96C" authority="(Ng &amp; Goh, 1987)" baseAuthorityName="Ng &amp; Goh" baseAuthorityYear="1987" box="[452,1045,1663,1689]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="rhadamanthysi">
<emphasis id="12B1740FFE7FFFCCFE5C2510E468F96C" box="[452,831,1663,1689]" italics="true" pageId="1" pageNumber="408">Arachnothelphusa rhadamanthysi</emphasis>
(
<bibRefCitation id="4454D5ECFE7FFFCCFCD52510E35AF96C" author="Ng, P. K. L. &amp; Goh, R." box="[845,1037,1663,1689]" pageId="1" pageNumber="408" pagination="313 - 330" refId="ref6717" refString="Ng, P. K. L. &amp; Goh, R. (1987) Cavernicolous freshwater crabs (Crustacea Decapoda, Brachyura) from Sabah, Borneo. Stygologia, 3 (4), 313 - 330, pls. 1 - 3." type="journal article" year="1987">Ng &amp; Goh, 1987</bibRefCitation>
)
</taxonomicName>
:
<typeStatus id="FF7E16BFFE7FFFCCFB872510E3D6F96C" box="[1055,1153,1663,1689]" pageId="1" pageNumber="408" type="holotype">holotype</typeStatus>
female (16.9 × 12.0 mm) (
<specimenCode id="70630066FE7FFFCCFF0725CCE60FF948" box="[159,344,1699,1725]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="408">ZRC 1990.0443</specimenCode>
),
<location id="251AFEC6FE7FFFCCFEF325CCE56AF948" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7FFFCCFEF325CCE56AF948" box="[363,573,1699,1725]" country="Malaysia" county="Borneo" latitude="5.55" longLatPrecision="1305" longitude="118.1" municipality="East" name="Simud Puteh Cave" pageId="1" pageNumber="408" stateProvince="Sabah">Simud Puteh Cave</location>
,
<location id="251AFEC6FE7FFFCCFDD025CCE598F948" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7FFFCCFDD025CCE598F948" box="[584,719,1699,1725]" country="Malaysia" county="Borneo" latitude="5.55" longLatPrecision="1305" longitude="118.1" municipality="East" name="Gomantong" pageId="1" pageNumber="408" stateProvince="Sabah">Gomantong</location>
,
<collectingRegion id="E20166FFFE7FFFCCFD4125CCE477F948" box="[729,800,1699,1725]" country="Malaysia" name="Sabah" pageId="1" pageNumber="408">Sabah</collectingRegion>
,
<collectingMunicipality id="C01E3267FE7FFFCCFCB325CCE40CF948" box="[811,859,1699,1725]" pageId="1" pageNumber="408">East</collectingMunicipality>
<collectingCountry id="58D2E88DFE7FFFCCFCFA25CCE49CF948" box="[866,971,1699,1725]" name="Malaysia" pageId="1" pageNumber="408">Malaysia</collectingCountry>
,
<collectingCounty id="C91BD091FE7FFFCCFC4E25CCE37CF948" box="[982,1067,1699,1725]" pageId="1" pageNumber="408">Borneo</collectingCounty>
,
<geoCoordinate id="45F1CEDAFE7FFFCCFBAE25CDE3DFF948" box="[1078,1160,1698,1725]" degrees="5" direction="north" minutes="33" orientation="latitude" pageId="1" pageNumber="408" precision="925" value="5.55">5°33N</geoCoordinate>
<geoCoordinate id="45F1CEDAFE7FFFCCFB1725CCE3ACF948" box="[1167,1275,1699,1725]" degrees="118" direction="east" minutes="06" orientation="longitude" pageId="1" pageNumber="408" precision="925" value="118.1">118°06E</geoCoordinate>
, coll.
<collectorName id="8D30CDCBFE7FFFCCFAA625CCE79DF914" pageId="1" pageNumber="408">P. Chapman</collectorName>
,
<date id="547B8EDDFE7FFFCCFF4F25A9E6DFF914" box="[215,392,1734,1761]" pageId="1" pageNumber="408" value="1986-03-25">
<collectingDate id="443F7735FE7FFFCCFF4F25A9E6DFF914" box="[215,392,1734,1761]" pageId="1" pageNumber="408" value="1986-03-25">25 March 1986</collectingDate>
</date>
</materialsCitation>
.
<taxonomicName id="E7C5D39EFE7FFFCCFE0D25A8E469F914" authority="Ng, 1991" authorityName="Ng" authorityYear="1991" box="[405,830,1735,1761]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="terrapes">
<emphasis id="12B1740FFE7FFFCCFE0D25A8E59FF915" box="[405,712,1735,1761]" italics="true" pageId="1" pageNumber="408">Arachnothelphusa terrapes</emphasis>
<bibRefCitation id="4454D5ECFE7FFFCCFD4825A8E469F914" author="Ng, P. K. L." box="[720,830,1735,1761]" pageId="1" pageNumber="408" pagination="1 - 12" refId="ref6389" refString="Ng, P. K. L. (1991) Bornean freshwater crabs of the genus Arachnothelphusa gen. nov. (Crustacea: Decapoda: Brachyura: Gecarcinucidae). Zoologische Mededelingen, 65 (1), 1 - 12." type="journal article" year="1991">Ng, 1991</bibRefCitation>
</taxonomicName>
:
<materialsCitation id="90ADA240FE7FFFCCFCD325A8E662F8DC" collectingDate="1990-10-23" collectionCode="ZRC" collectorName="H. K. Voris" country="Malaysia" county="Borneo" location="Danum Valley Field Centre" municipality="Lahad Datu" pageId="1" pageNumber="408" specimenCode="ZRC 1992.7918" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="FF7E16BFFE7FFFCCFCD325A8E4E4F914" box="[843,947,1735,1761]" pageId="1" pageNumber="408" type="holotype">Holotype</typeStatus>
male (17.6 ×
<quantity id="E73D05F8FE7FFFCCFBCE25A8E3EBF914" box="[1110,1212,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="1" pageNumber="408" unit="mm" value="13.3">13.3 mm</quantity>
) (
<specimenCode id="70630066FE7FFFCCFB4C25A8E2C7F914" box="[1236,1424,1735,1761]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="408">ZRC 1992.7918</specimenCode>
),
<location id="251AFEC6FE7FFFCCFF0F2584E685F8F0" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7FFFCCFF0F2584E685F8F0" box="[151,466,1771,1797]" country="Malaysia" county="Borneo" municipality="Lahad Datu" name="Danum Valley Field Centre" pageId="1" pageNumber="408" stateProvince="Sabah">Danum Valley Field Centre</location>
, station 507, in dry stump on ridge,
<collectingMunicipality id="C01E3267FE7FFFCCFCEB2584E4ADF8F0" box="[883,1018,1771,1797]" pageId="1" pageNumber="408">Lahad Datu</collectingMunicipality>
,
<collectingRegion id="E20166FFFE7FFFCCFB902584E318F8F0" box="[1032,1103,1771,1797]" country="Malaysia" name="Sabah" pageId="1" pageNumber="408">Sabah</collectingRegion>
,
<collectingCounty id="C91BD091FE7FFFCCFBC42584E3E6F8F0" box="[1116,1201,1771,1797]" pageId="1" pageNumber="408">Borneo</collectingCounty>
, coll.
<collectorName id="8D30CDCBFE7FFFCCFB6F2584E223F8F0" box="[1271,1396,1771,1797]" pageId="1" pageNumber="408">H.K. Voris</collectorName>
,
<date id="547B8EDDFE7FFFCCFA182584E662F8DC" pageId="1" pageNumber="408" value="1990-10-23">
<collectingDate id="443F7735FE7FFFCCFA182584E662F8DC" pageId="1" pageNumber="408" value="1990-10-23">23 October 1990</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="90ADA240FE7FFFCCFEDA2460E6B1F8B8" collectingDate="1989-07-21" collectionCode="ZRC" collectorName="S. C. Choy" country="Malaysia" county="Borneo" location="Danum Valley" municipality="Lahad Datu" pageId="1" pageNumber="408" specimenCode="ZRC 1992.7919" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="FF7E16BFFE7FFFCCFEDA2460E6F5F8DC" box="[322,418,1807,1833]" pageId="1" pageNumber="408" type="paratype">paratype</typeStatus>
female (25.7 ×
<quantity id="E73D05F8FE7FFFCCFDC32460E595F8DC" box="[603,706,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="1" pageNumber="408" unit="mm" value="18.6">18.6 mm</quantity>
) (
<specimenCode id="70630066FE7FFFCCFD422460E4C1F8DC" box="[730,918,1807,1833]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="408">ZRC 1992.7919</specimenCode>
),
<location id="251AFEC6FE7FFFCCFC322460E319F8DC" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7FFFCCFC322460E319F8DC" box="[938,1102,1807,1833]" country="Malaysia" county="Borneo" municipality="Lahad Datu" name="Danum Valley" pageId="1" pageNumber="408" stateProvince="Sabah">Danum Valley</location>
,
<collectingMunicipality id="C01E3267FE7FFFCCFBC22460E3B5F8DC" box="[1114,1250,1807,1833]" pageId="1" pageNumber="408">Lahad Datu</collectingMunicipality>
,
<collectingRegion id="E20166FFFE7FFFCCFB772460E261F8DC" box="[1263,1334,1807,1833]" country="Malaysia" name="Sabah" pageId="1" pageNumber="408">Sabah</collectingRegion>
,
<collectingCounty id="C91BD091FE7FFFCCFADB2460E2CFF8DC" box="[1347,1432,1807,1833]" pageId="1" pageNumber="408">Borneo</collectingCounty>
, coll.
<collectorName id="8D30CDCBFE7FFFCCFF48245CE611F8B8" box="[208,326,1843,1869]" pageId="1" pageNumber="408">S.C. Choy</collectorName>
,
<date id="547B8EDDFE7FFFCCFEC9245CE6B1F8B8" box="[337,486,1843,1869]" pageId="1" pageNumber="408" value="1989-07-21">
<collectingDate id="443F7735FE7FFFCCFEC9245CE6B1F8B8" box="[337,486,1843,1869]" pageId="1" pageNumber="408" value="1989-07-21">21 July 1989</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="90ADA240FE7FFFCCFE6B245CE6B6F860" collectingDate="2017-07-20" collectionCode="ZRC" collectorName="Lahad Datu &amp; Borneo" country="Malaysia" location="Danum Valley" pageId="1" pageNumber="408" specimenCode="ZRC 2017.1205" specimenCount="28" specimenCount-female="1" specimenCount-juvenile="26" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<specimenCount id="36C36394FE7FFFCCFE6B245CE569F8B8" box="[499,574,1843,1869]" pageId="1" pageNumber="408" type="male">1 male</specimenCount>
(30.8 ×
<quantity id="E73D05F8FE7FFFCCFD38245DE45FF8B8" box="[672,776,1842,1869]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="1" pageNumber="408" unit="mm" value="20.5">20.5 mm</quantity>
),
<specimenCount id="36C36394FE7FFFCCFC84245CE42AF8B8" box="[796,893,1843,1869]" pageId="1" pageNumber="408" type="female">1 female</specimenCount>
(30.1 ×
<quantity id="E73D05F8FE7FFFCCFC47245DE312F8B8" box="[991,1093,1842,1869]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="1" pageNumber="408" unit="mm" value="20.5">20.5 mm</quantity>
, with
<specimenCount id="36C36394FE7FFFCCFB13245CE25EF8B8" box="[1163,1289,1843,1869]" pageId="1" pageNumber="408" type="juvenile">26 juvenile</specimenCount>
crabs) (
<specimenCode id="70630066FE7FFFCCFAFC245CE644F884" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="408">ZRC 2017.1205</specimenCode>
), from water-filled tree buttress, ca.
<quantity id="E73D05F8FE7FFFCCFD2D2439E5ACF884" box="[693,763,1878,1905]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="408" unit="cm" value="35.0">35 cm</quantity>
above ground
<location id="251AFEC6FE7FFFCCFC312438E31AF884" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7FFFCCFC312438E31AF884" box="[937,1101,1879,1905]" country="Malaysia" name="Danum Valley" pageId="1" pageNumber="408" stateProvince="Sabah">Danum Valley</location>
,
<collectorName id="8D30CDCBFE7FFFCCFBC12438E3B6F884" box="[1113,1249,1879,1905]" pageId="1" pageNumber="408">Lahad Datu</collectorName>
,
<collectingRegion id="E20166FFFE7FFFCCFB762438E262F884" box="[1262,1333,1879,1905]" country="Malaysia" name="Sabah" pageId="1" pageNumber="408">Sabah</collectingRegion>
,
<collectorName id="8D30CDCBFE7FFFCCFADB2438E2CFF884" box="[1347,1432,1879,1905]" pageId="1" pageNumber="408">Borneo</collectorName>
,
<collectingCountry id="58D2E88DFE7FFFCCFF0F2414E657F860" box="[151,256,1915,1941]" name="Malaysia" pageId="1" pageNumber="408">Malaysia</collectingCountry>
, coll.
<date id="547B8EDDFE7FFFCCFED02414E6B6F860" box="[328,481,1915,1941]" pageId="1" pageNumber="408" value="2017-07-20">
<collectingDate id="443F7735FE7FFFCCFED02414E6B6F860" box="[328,481,1915,1941]" pageId="1" pageNumber="408" value="2017-07-20">20 July 2017</collectingDate>
</date>
</materialsCitation>
.
<taxonomicName id="E7C5D39EFE7FFFCCFE752414E3DBF860" authority="Grinang, Pui &amp; Ng, 2015" authorityName="Grinang, Pui &amp; Ng" authorityYear="2015" box="[493,1164,1914,1941]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="merarapensis">
<emphasis id="12B1740FFE7FFFCCFE752414E40CF861" box="[493,859,1915,1941]" italics="true" pageId="1" pageNumber="408">Arachnothelphusa merarapensis</emphasis>
<bibRefCitation id="4454D5ECFE7FFFCCFCFC2414E3DBF860" author="Grinang, J. &amp; Pui, Y. M. &amp; Ng, P. K. L." box="[868,1164,1914,1941]" pageId="1" pageNumber="408" pagination="454 - 460" refId="ref5602" refString="Grinang, J., Pui, Y. M. &amp; Ng, P. K. L. (2015) A new species of tree-hole dwelling crab of the genus Arachnothelphusa Ng, 1991 (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from northern Sarawak, Malaysia, Borneo. Raffles Bulletin of Zoology, 63, 454 - 460." type="journal article" year="2015">Grinang, Pui &amp; Ng, 2015</bibRefCitation>
</taxonomicName>
:
<materialsCitation id="90ADA240FE7FFFCFFB002414E513FF44" collectingDate="2014-10-31" collectionCode="ZRC" collectorName="J. Grinang &amp; Y. M. Pui" country="Malaysia" county="Borneo" elevation="485" lastPageId="2" lastPageNumber="409" latitude="4.3737226" location="Merarap Hot Spring Resort" longLatPrecision="1" longitude="115.43614" municipality="Lawas" pageId="1" pageNumber="408" specimenCode="ZRC 2016.0297" specimenCount="1" typeStatus="holotype">
<typeStatus id="FF7E16BFFE7FFFCCFB002414E257F860" box="[1176,1280,1915,1941]" pageId="1" pageNumber="408" type="holotype">Holotype</typeStatus>
male (22.5 ×
<quantity id="E73D05F8FE7FFFCCFF0F24F0E7A8F84C" box="[151,255,1951,1977]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="1" pageNumber="408" unit="mm" value="16.8">16.8 mm</quantity>
) (
<specimenCode id="70630066FE7FFFCCFE8024F0E681F84F" box="[280,470,1951,1978]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="408">ZRC 2016.0297</specimenCode>
), water-filled tree-hole, ca.
<quantity id="E73D05F8FE7FFFCCFC8C24F0E43DF84C" box="[788,874,1951,1977]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="408" unit="cm" value="100.0">100 cm</quantity>
above ground, steep dipterocarp forest,
<location id="251AFEC6FE7FFFCCFAA524F0E63DF828" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7FFFCCFAA524F0E63DF828" country="Malaysia" county="Borneo" latitude="4.3737226" longLatPrecision="1" longitude="115.43614" municipality="Lawas" name="Merarap Hot Spring Resort" pageId="1" pageNumber="408">Merarap Hot Spring Resort</location>
,
<collectingMunicipality id="C01E3267FE7FFFCCFEEF24ACE694F828" box="[375,451,1987,2013]" pageId="1" pageNumber="408">Lawas</collectingMunicipality>
, northern
<collectingCountry id="58D2E88DFE7FFFCCFDA124ACE5CBF828" box="[569,668,1987,2013]" name="Malaysia" pageId="1" pageNumber="408">Sarawak</collectingCountry>
,
<collectingCountry id="58D2E88DFE7FFFCCFD3124ACE445F828" box="[681,786,1987,2013]" name="Malaysia" pageId="1" pageNumber="408">Malaysia</collectingCountry>
,
<collectingCounty id="C91BD091FE7FFFCCFCB824ACE422F828" box="[800,885,1987,2013]" pageId="1" pageNumber="408">Borneo</collectingCounty>
,
<geoCoordinate id="45F1CEDAFE7FFFCCFC1A24ADE347F828" box="[898,1040,1986,2013]" degrees="4" direction="north" minutes="22" orientation="latitude" pageId="1" pageNumber="408" precision="1" seconds="25.4" value="4.3737226">4º2225.4”N</geoCoordinate>
<geoCoordinate id="45F1CEDAFE7FFFCCFB8124ADE396F828" box="[1049,1217,1986,2013]" degrees="115" direction="east" minutes="26" orientation="longitude" pageId="1" pageNumber="408" precision="1" seconds="10.1" value="115.43614">115º2610.1”E</geoCoordinate>
,
<elevation id="ABE84F2EFE7FFFCCFB5724ADE214F828" box="[1231,1347,1986,2013]" metricMagnitude="2" metricUnit="m" metricValue="4.85" pageId="1" pageNumber="408" unit="m" value="485.0">
<quantity id="E73D05F8FE7FFFCCFB5724ADE24FF828" box="[1231,1304,1986,2013]" metricMagnitude="2" metricUnit="m" metricValue="4.85" pageId="1" pageNumber="408" unit="m" value="485.0">485 m</quantity>
asl
</elevation>
, coll.
<collectorName id="8D30CDCBFE7FFFCFFA1224ACE7A4FF44" lastPageId="2" lastPageNumber="409" pageId="1" pageNumber="408">J. Grinang</collectorName>
&amp;
<collectorName id="8D30CDCBFE7CFFCFFE8C23F8E62DFF44" box="[276,378,151,177]" pageId="2" pageNumber="409">Y.M. Pui</collectorName>
,
<date id="547B8EDDFE7CFFCFFE1D23F8E513FF44" box="[389,580,151,177]" pageId="2" pageNumber="409" value="2014-10-31">
<collectingDate id="443F7735FE7CFFCFFE1D23F8E513FF44" box="[389,580,151,177]" pageId="2" pageNumber="409" value="2014-10-31">31 October 2014</collectingDate>
</date>
</materialsCitation>
.
<taxonomicName id="E7C5D39EFE7CFFCFFDD523F8E308FF44" authority="Grinang &amp; Ng, 2021" authorityName="Grinang &amp; Ng" authorityYear="2021" box="[589,1119,151,177]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="409" phylum="Arthropoda" rank="species" species="sarang">
<emphasis id="12B1740FFE7CFFCFFDD523F8E438FF45" box="[589,879,151,177]" italics="true" pageId="2" pageNumber="409">Arachnothelphusa sarang</emphasis>
<bibRefCitation id="4454D5ECFE7CFFCFFCED23F8E308FF44" author="Grinang, J. &amp; Ng, P. K. L." box="[885,1119,151,177]" pageId="2" pageNumber="409" pagination="1 - 7" refId="ref5542" refString="Grinang, J. &amp; Ng, P. K. L. (2021) A new species of the genus Arachnothelphusa Ng, 1991 (Crustacea: Decapoda: Gecarcinucidae) from a limestone cave in Sarawak (Malaysian Borneo). Raffles Bulletin of Zoology. Raffles Bulletin of Zoology, 69, 1 - 7." type="journal article" year="2021">Grinang &amp; Ng, 2021</bibRefCitation>
</taxonomicName>
:
<materialsCitation id="90ADA240FE7CFFCFFBF123F8E785FF0C" collectingDate="2005-08-20" collectionCode="ZRC" collectorName="H. H. Tan" country="Malaysia" location="Bukit Sarang" municipality="Bintulu" pageId="2" pageNumber="409" specimenCode="ZRC 2020.0098" specimenCount="1" typeStatus="holotype">
<typeStatus id="FF7E16BFFE7CFFCFFBF123F8E386FF44" box="[1129,1233,151,177]" pageId="2" pageNumber="409" type="holotype">Holotype</typeStatus>
male (20.4 ×
<quantity id="E73D05F8FE7CFFCFFAF323F8E793FF20" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="2" pageNumber="409" unit="mm" value="14.7">14.7 mm</quantity>
) (
<specimenCode id="70630066FE7CFFCFFF4223D4E6C2FF20" box="[218,405,187,213]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="409">ZRC 2020.0098</specimenCode>
), limestone cave,
<location id="251AFEC6FE7CFFCFFDC423D4E5A5FF20" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7CFFCFFDC423D4E5A5FF20" box="[604,754,187,213]" country="Malaysia" municipality="Bintulu" name="Bukit Sarang" pageId="2" pageNumber="409">Bukit Sarang</location>
,
<collectingMunicipality id="C01E3267FE7CFFCFFD6423D4E404FF20" box="[764,851,187,213]" pageId="2" pageNumber="409">Bintulu</collectingMunicipality>
,
<collectingCountry id="58D2E88DFE7CFFCFFCC623D4E496FF20" box="[862,961,187,213]" name="Malaysia" pageId="2" pageNumber="409">Sarawak</collectingCountry>
,
<collectingCountry id="58D2E88DFE7CFFCFFC5423D4E362FF20" box="[972,1077,187,213]" name="Malaysia" pageId="2" pageNumber="409">Malaysia</collectingCountry>
, coll.
<collectorName id="8D30CDCBFE7CFFCFFBE023D4E388FF20" box="[1144,1247,187,213]" pageId="2" pageNumber="409">H.H. Tan</collectorName>
<emphasis id="12B1740FFE7CFFCFFB7D23D4E24AFF20" box="[1253,1309,187,213]" italics="true" pageId="2" pageNumber="409">et al.</emphasis>
,
<date id="547B8EDDFE7CFFCFFAB223D4E785FF0C" pageId="2" pageNumber="409" value="2005-08-20">
<collectingDate id="443F7735FE7CFFCFFAB223D4E785FF0C" pageId="2" pageNumber="409" value="2005-08-20">20 August 2005</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="90ADA240FE7CFFCFFF7823B0E64BFEE8" collectingDate="2005-08-20" collectionCode="ZRC" collectorName="H. H. Tan" country="Malaysia" location="Bukit Sarang" municipality="Bintulu" pageId="2" pageNumber="409" specimenCode="ZRC 2020.0099" specimenCount="5" specimenCount-female="4" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="FF7E16BFFE7CFFCFFF7823B0E607FF0C" box="[224,336,223,249]" pageId="2" pageNumber="409" type="paratype">paratypes</typeStatus>
:
<specimenCount id="36C36394FE7CFFCFFEC323B0E6FFFF0C" box="[347,424,223,249]" pageId="2" pageNumber="409" type="male">1 male</specimenCount>
(18.7 ×
<quantity id="E73D05F8FE7CFFCFFD9523B0E520FF0C" box="[525,631,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="409" unit="mm" value="14.8">14.8 mm</quantity>
),
<specimenCount id="36C36394FE7CFFCFFD1323B0E5AEFF0C" box="[651,761,223,249]" pageId="2" pageNumber="409" type="female">4 females</specimenCount>
(15.819.8 × 12.0
<quantity id="E73D05F8FE7CFFCFFC4423B1E315FF0C" box="[988,1090,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="2" pageNumber="409" unit="mm" value="15.8">15.8 mm</quantity>
) (
<specimenCode id="70630066FE7CFFCFFBC223B0E240FF0C" box="[1114,1303,223,249]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="409">ZRC 2020.0099</specimenCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="90ADA240FE7CFFCFFEB0226CE3E8FEB4" collectingDate="2005-08-20" collectionCode="ZRC" collectorName="H. H. Tan" country="Malaysia" county="Bintulu" location="Batu Gelam" municipality="Bukit Sarang" pageId="2" pageNumber="409" specimenCode="ZRC 2020.0100" specimenCount="17" specimenCount-female="7" specimenCount-male="10" typeStatus="holotype">
<specimenCount id="36C36394FE7CFFCFFEB0226CE6DAFEE8" box="[296,397,259,285]" pageId="2" pageNumber="409" type="male">10 males</specimenCount>
(7.411.2 ×
<quantity id="E73D05F8FE7CFFCFFDB8226DE5FCFEE8" box="[544,683,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.699999999999999" metricValueMax="9.6" metricValueMin="5.8" pageId="2" pageNumber="409" unit="mm" value="7.699999999999999" valueMax="9.6" valueMin="5.8">5.89.6 mm</quantity>
),
<specimenCount id="36C36394FE7CFFCFFD27226CE47BFEE8" box="[703,812,259,285]" pageId="2" pageNumber="409" type="female">7 females</specimenCount>
(7.512.7 ×
<quantity id="E73D05F8FE7CFFCFFC27226DE31FFEE8" box="[959,1096,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.85" metricValueMax="9.9" metricValueMin="5.8" pageId="2" pageNumber="409" unit="mm" value="7.85" valueMax="9.9" valueMin="5.8">5.89.9 mm</quantity>
) (
<specimenCode id="70630066FE7CFFCFFBF8226CE24BFEE8" box="[1120,1308,259,285]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="409">ZRC 2020.0100</specimenCode>
), limestone cave,
<location id="251AFEC6FE7CFFCFFF412248E634FEB4" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7CFFCFFF412248E634FEB4" box="[217,355,295,321]" country="Malaysia" county="Bintulu" municipality="Bukit Sarang" name="Batu Gelam" pageId="2" pageNumber="409">Batu Gelam</location>
,
<collectingMunicipality id="C01E3267FE7CFFCFFEF72248E550FEB4" box="[367,519,295,321]" pageId="2" pageNumber="409">Bukit Sarang</collectingMunicipality>
,
<collectingCounty id="C91BD091FE7CFFCFFD8A2248E53EFEB4" box="[530,617,295,321]" pageId="2" pageNumber="409">Bintulu</collectingCounty>
,
<collectingCountry id="58D2E88DFE7CFFCFFDED2248E58FFEB4" box="[629,728,295,321]" name="Malaysia" pageId="2" pageNumber="409">Sarawak</collectingCountry>
,
<collectingCountry id="58D2E88DFE7CFFCFFD7C2248E41AFEB4" box="[740,845,295,321]" name="Malaysia" pageId="2" pageNumber="409">Malaysia</collectingCountry>
, coll.
<collectorName id="8D30CDCBFE7CFFCFFC0A2248E4ABFEB4" box="[914,1020,295,321]" pageId="2" pageNumber="409">H.H. Tan</collectorName>
,
<date id="547B8EDDFE7CFFCFFB912248E3E8FEB4" box="[1033,1215,294,321]" pageId="2" pageNumber="409" value="2005-08-20">
<collectingDate id="443F7735FE7CFFCFFB912248E3E8FEB4" box="[1033,1215,294,321]" pageId="2" pageNumber="409" value="2005-08-20">20 August 2005</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="90ADA240FE7CFFCFFB532248E5FAFE7C" collectingDate="2005-08-18" collectionCode="ZRC" collectorName="H. H. Tan" country="Malaysia" county="Bintulu" location="Batu Kelelut" municipality="Bukit Sarang" pageId="2" pageNumber="409" specimenCode="ZRC 2020.0351" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="36C36394FE7CFFCFFB532248E240FEB4" box="[1227,1303,295,321]" pageId="2" pageNumber="409" type="male">1 male</specimenCount>
(12.1 ×
<quantity id="E73D05F8FE7CFFCFFAE12248E790FE90" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="2" pageNumber="409" unit="mm" value="9.9">9.9 mm</quantity>
),
<specimenCount id="36C36394FE7CFFCFFF422224E66CFE90" box="[218,315,331,357]" pageId="2" pageNumber="409" type="female">1 female</specimenCount>
(12.9 ×
<quantity id="E73D05F8FE7CFFCFFE012224E6A9FE90" box="[409,510,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="409" unit="mm" value="10.4">10.4 mm</quantity>
) (
<specimenCode id="70630066FE7CFFCFFD8C2224E599FE90" box="[532,718,330,357]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="409">ZRC 2020.0351</specimenCode>
), limestone cave,
<location id="251AFEC6FE7CFFCFFC0E2224E370FE90" LSID="urn:lsid:plazi:treatment:A86C190BFE7FFFCFFF0F2129E5E7FE7C:251AFEC6FE7CFFCFFC0E2224E370FE90" box="[918,1063,331,357]" country="Malaysia" county="Bintulu" municipality="Bukit Sarang" name="Batu Kelelut" pageId="2" pageNumber="409">Batu Kelelut</location>
,
<collectingMunicipality id="C01E3267FE7CFFCFFBAA2224E39FFE90" box="[1074,1224,331,357]" pageId="2" pageNumber="409">Bukit Sarang</collectingMunicipality>
,
<collectingCounty id="C91BD091FE7CFFCFFB4B2224E27DFE90" box="[1235,1322,331,357]" pageId="2" pageNumber="409">Bintulu</collectingCounty>
,
<collectingCountry id="58D2E88DFE7CFFCFFAAD2224E2CFFE90" box="[1333,1432,331,357]" name="Malaysia" pageId="2" pageNumber="409">Sarawak</collectingCountry>
,
<collectingCountry id="58D2E88DFE7CFFCFFF0F2200E657FE7C" box="[151,256,367,393]" name="Malaysia" pageId="2" pageNumber="409">Malaysia</collectingCountry>
, coll.
<collectorName id="8D30CDCBFE7CFFCFFEDC2200E6FCFE7C" box="[324,427,367,393]" pageId="2" pageNumber="409">H.H. Tan</collectorName>
<emphasis id="12B1740FFE7CFFCFFE292200E6BDFE7C" box="[433,490,367,393]" italics="true" pageId="2" pageNumber="409">et al.</emphasis>
,
<date id="547B8EDDFE7CFFCFFE6F2200E5FAFE7C" box="[503,685,366,393]" pageId="2" pageNumber="409" value="2005-08-18">
<collectingDate id="443F7735FE7CFFCFFE6F2200E5FAFE7C" box="[503,685,366,393]" pageId="2" pageNumber="409" value="2005-08-18">18 August 2005</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>