421 lines
66 KiB
XML
421 lines
66 KiB
XML
<document id="2AD5A7896A9C4ED9AF3843ABEB7CF915" ID-DOI="10.11646/zootaxa.5016.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5222295" ID-ZooBank="369BA2BD-7673-44EC-9D54-D846F112E4A4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629380692508" checkinUser="plazi" docAuthor="Ng, Peter K. L." docDate="2021" docId="A86C190BFE7CFFCBFF0F22B5E266FC3C" docLanguage="en" docName="zootaxa.5016.3.6.pdf" docOrigin="Zootaxa 5016 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arachnothelphusa rimba Ng 2021, n. sp." docType="treatment" docVersion="4" lastPageNumber="413" masterDocId="54556173FE7EFFCDFF98236FE757FFF5" masterDocTitle="On two new species of arboreal crabs from phytotelms in Sarawak, Borneo (Crustacea: Brachyura: Gecarcinucidae: Arachnothelphusa)" masterLastPageNumber="418" masterPageNumber="407" pageNumber="409" updateTime="1699070539357" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F9BBF98778260F8B9E627F8C4299E251">
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<mods:title id="26FEBA2FABED2123923470F55EB57816">On two new species of arboreal crabs from phytotelms in Sarawak, Borneo (Crustacea: Brachyura: Gecarcinucidae: Arachnothelphusa)</mods:title>
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<mods:name id="29596EF968AF19BC678D100ED31ECA01" type="personal">
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<mods:role id="72F0189719CC31B9BB75B78653777F3E">
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<mods:roleTerm id="6250B1EB85565FE045C51AD330E35795">Author</mods:roleTerm>
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<mods:namePart id="082E065894AFB98966528749096E66F5">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="A523ECB13563FA1C12D28273A423B5C1">Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore.</mods:affiliation>
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<mods:typeOfResource id="721BC4B702C7ADCC227F71395F4FD2AD">text</mods:typeOfResource>
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<mods:title id="BEF521C8C0DF6E94619A54727AA82823">Zootaxa</mods:title>
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<mods:part id="900E6CC6B230446B6574B10CB6601DC8">
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<mods:date id="62F43E01499B57191A1CE5F362D440FE">2021</mods:date>
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<mods:number id="BF8FB98D90F9E1345EFB04674FB6C674">2021-08-06</mods:number>
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<mods:number id="23DC33EA86CFC4844BCBCB26102F8B47">5016</mods:number>
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<mods:number id="A050E7F3D01B4EC942EF33ABE4908E9B">3</mods:number>
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<mods:start id="E396CBD5CEDB46109DABC118392DC6A2">407</mods:start>
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<mods:classification id="3E1BE3B091B8F82CBA2364AA720928C5">journal article</mods:classification>
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<mods:identifier id="B96823601247205D2BFE3EA7BF50EDA1" type="DOI">10.11646/zootaxa.5016.3.6</mods:identifier>
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<mods:identifier id="E151B7D94888506CCC509C0C3495585C" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A86C190BFE7CFFCBFF0F22B5E266FC3C" ID-DOI="http://doi.org/10.5281/zenodo.5222350" ID-Zenodo-Dep="5222350" LSID="urn:lsid:plazi:treatment:A86C190BFE7CFFCBFF0F22B5E266FC3C" httpUri="http://treatment.plazi.org/id/A86C190BFE7CFFCBFF0F22B5E266FC3C" lastPageId="6" lastPageNumber="413" pageId="2" pageNumber="409">
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<subSubSection id="68DFFB96FE7CFFCFFF0F22B5E546FE00" box="[151,529,474,501]" pageId="2" pageNumber="409" type="nomenclature">
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<paragraph id="207AA81DFE7CFFCFFF0F22B5E546FE00" blockId="2.[151,529,474,538]" box="[151,529,474,501]" pageId="2" pageNumber="409">
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<heading id="7B321F71FE7CFFCFFF0F22B5E546FE00" bold="true" box="[151,529,474,501]" fontSize="11" level="1" pageId="2" pageNumber="409" reason="1">
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<emphasis id="12B1740FFE7CFFCFFF0F22B5E546FE00" bold="true" box="[151,529,474,501]" pageId="2" pageNumber="409">
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<taxonomicName id="E7C5D39EFE7CFFCFFF0F22B5E690FE00" ID-CoL="8NFLZ" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[151,455,474,501]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="409" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
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<emphasis id="12B1740FFE7CFFCFFF0F22B5E690FE00" bold="true" box="[151,455,474,501]" italics="true" pageId="2" pageNumber="409">Arachnothelphusa rimba</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="0982C974FE7CFFCFFE5722B4E546FE00" box="[463,529,475,501]" pageId="2" pageNumber="409" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="68DFFB96FE7CFFCFFF0F2290E645FDEF" box="[151,274,511,538]" pageId="2" pageNumber="409" type="description">
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<paragraph id="207AA81DFE7CFFCFFF0F2290E645FDEF" blockId="2.[151,529,474,538]" box="[151,274,511,538]" pageId="2" pageNumber="409">
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(
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<figureCitation id="B8FEB498FE7CFFCFFF072290E65EFDEF" box="[159,265,511,538]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStartId-0="2.[151,196,1945,1969]" captionStartId-1="3.[151,196,1179,1203]" captionStartId-2="3.[151,196,1944,1968]" captionStartId-3="4.[151,196,1073,1097]" captionStartId-4="4.[151,196,1571,1595]" captionStartId-5="5.[151,196,1689,1713]" captionTargetBox-0="[275,1312,882,1919]" captionTargetBox-1="[151,1436,181,1155]" captionTargetBox-2="[151,1436,1300,1921]" captionTargetBox-3="[161,1426,188,1041]" captionTargetBox-4="[151,1436,1158,1547]" captionTargetBox-5="[151,1436,381,1666]" captionTargetId-0="figure-290@2.[275,1312,882,1919]" captionTargetId-1="figure-15@3.[151,1436,181,1155]" captionTargetId-2="figure-66@3.[151,1436,1300,1921]" captionTargetId-3="figure-24@4.[151,1436,181,1049]" captionTargetId-4="figure-74@4.[151,1436,1158,1547]" captionTargetId-5="figure-104@5.[151,1436,381,1666]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionTargetPageId-5="5" captionText-0="FIG. 1. Arachnothelphusa rimba n. sp., holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela." captionText-1="FIG. 2. Arachnothelphusa rimba n. sp. A, B, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088); C, male M1 (19.1 × 13.9 mm) (ZRC 2020.0086); D, male (20.2 × 15.4 mm) (ZRC 2020.0087).A, overall dorsal view; B–D, dorsal view of carapace. All specimens from Lanjak-Entimau." captionText-2="FIG. 3. Arachnothelphusa rimba n. sp., ambulatory legs. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau; C, D, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088), Lanjak-Entimau. A, C, right P4; B, D, right P5." captionText-3="FIG. 4. Arachnothelphusa rimba n. sp. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsal view); D, left G2. Scales = 0.5 mm." captionText-4="FIG. 5. Arachnothelphusa rimba n. sp., paratype female (22.9 × 16.5 mm) (ZRC 2020.0088), Lanjak-Entimau. A, pleon; B, vulvae." captionText-5="FIG. 6. Arachnothelphusa rimba n. sp., colour in life. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366); C–E, paratype male (20.2 × 15.4 mm) (ZRC 2020.0087), F, paratype male (19.1 × 13.9 mm) (ZRC 2020.0086); G, H, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088).All specimens from Lanjak-Entimau. Photographs: Tan Heok Hui." figureDoi-0="http://doi.org/10.5281/zenodo.5222297" figureDoi-1="http://doi.org/10.5281/zenodo.5222299" figureDoi-2="http://doi.org/10.5281/zenodo.5222301" figureDoi-3="http://doi.org/10.5281/zenodo.5222303" figureDoi-4="http://doi.org/10.5281/zenodo.5222305" figureDoi-5="http://doi.org/10.5281/zenodo.5222307" httpUri-0="https://zenodo.org/record/5222297/files/figure.png" httpUri-1="https://zenodo.org/record/5222299/files/figure.png" httpUri-2="https://zenodo.org/record/5222301/files/figure.png" httpUri-3="https://zenodo.org/record/5222303/files/figure.png" httpUri-4="https://zenodo.org/record/5222305/files/figure.png" httpUri-5="https://zenodo.org/record/5222307/files/figure.png" pageId="2" pageNumber="409">Figs. 1–6</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="68DFFB96FE7CFFCFFF0F2129E5DFFCA8" pageId="2" pageNumber="409" type="materials_examined">
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<paragraph id="207AA81DFE7CFFCFFF0F2129E5DFFCA8" blockId="2.[151,1436,582,861]" pageId="2" pageNumber="409">
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||
<emphasis id="12B1740FFE7CFFCFFF0F2129E62CFD95" bold="true" box="[151,379,582,608]" pageId="2" pageNumber="409">Material examined</emphasis>
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.
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<materialsCitation id="90ADA240FE7CFFCFFE122128E391FD70" collectingDate="2019-10-12" collectionCode="ZRC" collectorName="K. O. Chan" pageId="2" pageNumber="409" specimenCode="ZRC 2020.0366" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="FF7E16BFFE7CFFCFFE122128E6A0FD94" box="[394,503,583,609]" pageId="2" pageNumber="409" type="holotype">Holotype</typeStatus>
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||
: male (22.8 ×
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||
<quantity id="E73D05F8FE7CFFCFFD032128E456FD94" box="[667,769,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="2" pageNumber="409" unit="mm" value="16.6">16.6 mm</quantity>
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||
) (
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||
<specimenCode id="70630066FE7CFFCFFC812128E482FD97" box="[793,981,583,610]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="409">ZRC 2020.0366</specimenCode>
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||
), station DZ-02; in tree holes, along upwards sloping forest trail behind base camp, at night, coll.
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||
<collectorName id="8D30CDCBFE7CFFCFFCAF2104E4E6FD70" box="[823,945,619,645]" pageId="2" pageNumber="409">K.O. Chan</collectorName>
|
||
<emphasis id="12B1740FFE7CFFCFFC222104E4A3FD70" box="[954,1012,619,645]" italics="true" pageId="2" pageNumber="409">et al.</emphasis>
|
||
,
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||
<date id="547B8EDDFE7CFFCFFB9B2104E391FD70" box="[1027,1222,619,645]" pageId="2" pageNumber="409" value="2019-10-12">
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||
<collectingDate id="443F7735FE7CFFCFFB9B2104E391FD70" box="[1027,1222,619,645]" pageId="2" pageNumber="409" value="2019-10-12">12 October 2019</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="90ADA240FE7CFFCFFB4A2104E5AAFD38" collectingDate="2019-10-12" collectionCode="ZRC" collectorName="K. O. Chan" pageId="2" pageNumber="409" specimenCode="ZRC 2020.0086" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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||
<typeStatus id="FF7E16BFFE7CFFCFFB4A2104E214FD70" box="[1234,1347,619,645]" pageId="2" pageNumber="409" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="36C36394FE7CFFCFFAD72104E2CBFD70" box="[1359,1436,619,645]" pageId="2" pageNumber="409" type="male">1 male</specimenCount>
|
||
(19.1 ×
|
||
<quantity id="E73D05F8FE7CFFCFFF6B21E0E60DFD5C" box="[243,346,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="2" pageNumber="409" unit="mm" value="13.9">13.9 mm</quantity>
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||
) (
|
||
<specimenCode id="70630066FE7CFFCFFEEA21E0E578FD5F" box="[370,559,655,682]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="409">ZRC 2020.0086</specimenCode>
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||
), station DZ-02; in tree holes, along upwards sloping forest trail behind base camp, at night, coll.
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||
<collectorName id="8D30CDCBFE7CFFCFFEE321DCE6A4FD38" box="[379,499,691,717]" pageId="2" pageNumber="409">K.O. Chan</collectorName>
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||
<emphasis id="12B1740FFE7CFFCFFE6221DCE565FD38" box="[506,562,691,717]" italics="true" pageId="2" pageNumber="409">et al.</emphasis>
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||
,
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||
<date id="547B8EDDFE7CFFCFFDA721DCE5AAFD38" box="[575,765,691,717]" pageId="2" pageNumber="409" value="2019-10-12">
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||
<collectingDate id="443F7735FE7CFFCFFDA721DCE5AAFD38" box="[575,765,691,717]" pageId="2" pageNumber="409" value="2019-10-12">12 October 2019</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="90ADA240FE7CFFCFFC9121DCE6ABFCE0" collectingDate="2019-10-17" collectionCode="ZRC" collectorName="Sungei Mujok & K. O. Chan & I. Isa" pageId="2" pageNumber="409" specimenCode="ZRC 2020.0087" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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||
<specimenCount id="36C36394FE7CFFCFFC9121DCE404FD38" box="[777,851,691,717]" pageId="2" pageNumber="409" type="male">1 male</specimenCount>
|
||
(20.2 ×
|
||
<quantity id="E73D05F8FE7CFFCFFC2A21DDE341FD38" box="[946,1046,690,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="2" pageNumber="409" unit="mm" value="15.4">15.4 mm</quantity>
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||
) (
|
||
<specimenCode id="70630066FE7CFFCFFBB421DCE3B0FD38" box="[1068,1255,691,717]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="409">ZRC 2020.0087</specimenCode>
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||
), station DZ-06, trail and stream behind base camp near boundary, in tree holes and on leaves,
|
||
<collectorName id="8D30CDCBFE7CFFCFFB8E21B8E3EDFD04" box="[1046,1210,727,753]" pageId="2" pageNumber="409">Sungei Mujok</collectorName>
|
||
, at night, coll.
|
||
<collectorName id="8D30CDCBFE7CFFCFFAFE21B8E785FCE0" pageId="2" pageNumber="409">K.O. Chan</collectorName>
|
||
&
|
||
<collectorName id="8D30CDCBFE7CFFCFFF6E2194E666FCE0" box="[246,305,763,789]" pageId="2" pageNumber="409">I. Isa</collectorName>
|
||
,
|
||
<date id="547B8EDDFE7CFFCFFEA52194E6ABFCE0" box="[317,508,763,789]" pageId="2" pageNumber="409" value="2019-10-17">
|
||
<collectingDate id="443F7735FE7CFFCFFEA52194E6ABFCE0" box="[317,508,763,789]" pageId="2" pageNumber="409" value="2019-10-17">17 October 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="90ADA240FE7CFFCFFD902194E4CBFCCC" collectingDate="2019-10-15" collectionCode="ZRC" collectorName="K. O. Chan" pageId="2" pageNumber="409" specimenCode="ZRC 2020.0088" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="36C36394FE7CFFCFFD902194E53EFCE0" box="[520,617,763,789]" pageId="2" pageNumber="409" type="female">1 female</specimenCount>
|
||
(22.9 ×
|
||
<quantity id="E73D05F8FE7CFFCFFD512195E479FCE0" box="[713,814,762,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="2" pageNumber="409" unit="mm" value="16.5">16.5 mm</quantity>
|
||
) (
|
||
<specimenCode id="70630066FE7CFFCFFCDD2194E357FCE0" box="[837,1024,763,789]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="409">ZRC 2020.0088</specimenCode>
|
||
), station DZ-04, along sloping forest trail behind base camp, at night, coll.
|
||
<collectorName id="8D30CDCBFE7CFFCFFDD42070E59DFCCC" box="[588,714,799,825]" pageId="2" pageNumber="409">K.O. Chan</collectorName>
|
||
,
|
||
<date id="547B8EDDFE7CFFCFFD4F2071E4CBFCCC" box="[727,924,798,825]" pageId="2" pageNumber="409" value="2019-10-15">
|
||
<collectingDate id="443F7735FE7CFFCFFD4F2071E4CBFCCC" box="[727,924,798,825]" pageId="2" pageNumber="409" value="2019-10-15">15 October 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
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||
.
|
||
<materialsCitation id="90ADA240FE7CFFCFFC3E2070E5D3FCA8" collectorName="East & Borneo" country="Malaysia" location="All" municipality="Lanjak-Entimau Wildlife Sanctuary" pageId="2" pageNumber="409" specimenCount="1" typeStatus="paratype">
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||
<location id="251AFEC6FE7CFFCFFC3E2070E49DFCCC" LSID="urn:lsid:plazi:treatment:A86C190BFE7CFFCBFF0F22B5E266FC3C:251AFEC6FE7CFFCFFC3E2070E49DFCCC" box="[934,970,799,825]" country="Malaysia" municipality="Lanjak-Entimau Wildlife Sanctuary" name="All" pageId="2" pageNumber="409">All</location>
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||
locations from
|
||
<collectingMunicipality id="C01E3267FE7CFFCFFB192070E65FFCA8" pageId="2" pageNumber="409">Lanjak-Entimau Wildlife Sanctuary</collectingMunicipality>
|
||
, Sarawak,
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||
<collectorName id="8D30CDCBFE7CFFCFFE1A202CE6E5FCA8" box="[386,434,835,861]" pageId="2" pageNumber="409">East</collectorName>
|
||
<collectingCountry id="58D2E88DFE7CFFCFFE21202CE575FCA8" box="[441,546,835,861]" name="Malaysia" pageId="2" pageNumber="409">Malaysia</collectingCountry>
|
||
,
|
||
<collectorName id="8D30CDCBFE7CFFCFFDB6202CE5D3FCA8" box="[558,644,835,861]" pageId="2" pageNumber="409">Borneo</collectorName>
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||
</materialsCitation>
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||
.
|
||
</paragraph>
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||
</subSubSection>
|
||
<caption id="74BAF895FE7CFFCFFF0F24F6E425F80C" ID-DOI="http://doi.org/10.5281/zenodo.5222297" ID-Zenodo-Dep="5222297" httpUri="https://zenodo.org/record/5222297/files/figure.png" pageId="2" pageNumber="409" startId="2.[151,196,1945,1969]" targetBox="[275,1312,882,1919]" targetPageId="2">
|
||
<paragraph id="207AA81DFE7CFFCFFF0F24F6E425F80C" blockId="2.[151,1437,1945,2041]" pageId="2" pageNumber="409">
|
||
<emphasis id="12B1740FFE7CFFCFFF0F24F6E7B3F844" bold="true" box="[151,228,1945,1969]" pageId="2" pageNumber="409">FIG. 1.</emphasis>
|
||
<taxonomicName id="E7C5D39EFE7CFFCFFF7324F6E6BFF844" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[235,488,1945,1969]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="409" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
|
||
<emphasis id="12B1740FFE7CFFCFFF7324F6E6BFF844" box="[235,488,1945,1969]" italics="true" pageId="2" pageNumber="409">Arachnothelphusa rimba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0982C974FE7CFFCFFE7624F5E571F844" box="[494,550,1946,1969]" pageId="2" pageNumber="409" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="74BAF895FE7DFFCEFF0F27F4E6BCFB0E" ID-DOI="http://doi.org/10.5281/zenodo.5222299" ID-Zenodo-Dep="5222299" httpUri="https://zenodo.org/record/5222299/files/figure.png" pageId="3" pageNumber="410" startId="3.[151,196,1179,1203]" targetBox="[151,1436,181,1155]" targetPageId="3">
|
||
<paragraph id="207AA81DFE7DFFCEFF0F27F4E6BCFB0E" blockId="3.[151,1436,1179,1275]" pageId="3" pageNumber="410">
|
||
<emphasis id="12B1740FFE7DFFCEFF0F27F4E7B2FB46" bold="true" box="[151,229,1179,1204]" pageId="3" pageNumber="410">FIG. 2.</emphasis>
|
||
<taxonomicName id="E7C5D39EFE7DFFCEFF7527F4E6BCFB46" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[237,491,1179,1203]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
|
||
<emphasis id="12B1740FFE7DFFCEFF7527F4E6BCFB46" box="[237,491,1179,1203]" italics="true" pageId="3" pageNumber="410">Arachnothelphusa rimba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0982C974FE7DFFCEFE6A27F3E57DFB46" box="[498,554,1180,1203]" pageId="3" pageNumber="410" rank="species">n. sp.</taxonomicNameLabel>
|
||
A, B, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088); C, male M1 (19.1 × 13.9 mm) (ZRC 2020.0086); D, male (20.2 × 15.4 mm) (ZRC 2020.0087). A, overall dorsal view; B–D, dorsal view of carapace. All specimens from Lanjak-Entimau.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="74BAF895FE7DFFCEFF0F24F7E24BF821" ID-DOI="http://doi.org/10.5281/zenodo.5222301" ID-Zenodo-Dep="5222301" httpUri="https://zenodo.org/record/5222301/files/figure.png" pageId="3" pageNumber="410" startId="3.[151,196,1944,1968]" targetBox="[151,1436,1300,1921]" targetPageId="3">
|
||
<paragraph id="207AA81DFE7DFFCEFF0F24F7E24BF821" blockId="3.[151,1436,1944,2005]" pageId="3" pageNumber="410">
|
||
<emphasis id="12B1740FFE7DFFCEFF0F24F7E7B3F845" bold="true" box="[151,228,1944,1969]" pageId="3" pageNumber="410">FIG. 3.</emphasis>
|
||
<taxonomicName id="E7C5D39EFE7DFFCEFF7224F7E6B0F845" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[234,487,1944,1968]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
|
||
<emphasis id="12B1740FFE7DFFCEFF7224F7E6B0F845" box="[234,487,1944,1968]" italics="true" pageId="3" pageNumber="410">Arachnothelphusa rimba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0982C974FE7DFFCEFE7524F6E572F845" box="[493,549,1945,1968]" pageId="3" pageNumber="410" rank="species">n. sp.</taxonomicNameLabel>
|
||
, ambulatory legs. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau; C, D, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088), Lanjak-Entimau. A, C, right P4; B, D, right P5.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="74BAF895FE7AFFC9FF0F275EE23EFB98" ID-DOI="http://doi.org/10.5281/zenodo.5222303" ID-Zenodo-Dep="5222303" httpUri="https://zenodo.org/record/5222303/files/figure.png" pageId="4" pageNumber="411" startId="4.[151,196,1073,1097]" targetBox="[161,1426,188,1041]" targetPageId="4">
|
||
<paragraph id="207AA81DFE7AFFC9FF0F275EE23EFB98" blockId="4.[151,1436,1073,1134]" pageId="4" pageNumber="411">
|
||
<emphasis id="12B1740FFE7AFFC9FF0F275EE7B2FBBC" bold="true" box="[151,229,1073,1098]" pageId="4" pageNumber="411">FIG. 4.</emphasis>
|
||
<taxonomicName id="E7C5D39EFE7AFFC9FF74275DE6BDFBBF" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[236,490,1074,1098]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="411" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
|
||
<emphasis id="12B1740FFE7AFFC9FF74275DE6BDFBBF" box="[236,490,1074,1098]" italics="true" pageId="4" pageNumber="411">Arachnothelphusa rimba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0982C974FE7AFFC9FE69275DE57FFBBC" box="[497,552,1074,1097]" pageId="4" pageNumber="411" rank="species">n. sp.</taxonomicNameLabel>
|
||
A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsal view); D, left G2. Scales = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="74BAF895FE7AFFC9FF0F254CE7B5F9AA" ID-DOI="http://doi.org/10.5281/zenodo.5222305" ID-Zenodo-Dep="5222305" httpUri="https://zenodo.org/record/5222305/files/figure.png" pageId="4" pageNumber="411" startId="4.[151,196,1571,1595]" targetBox="[151,1436,1158,1547]" targetPageId="4">
|
||
<paragraph id="207AA81DFE7AFFC9FF0F254CE7B5F9AA" blockId="4.[151,1437,1571,1631]" pageId="4" pageNumber="411">
|
||
<emphasis id="12B1740FFE7AFFC9FF0F254CE7B2F9CE" bold="true" box="[151,229,1571,1595]" pageId="4" pageNumber="411">FIG. 5.</emphasis>
|
||
<taxonomicName id="E7C5D39EFE7AFFC9FF76254CE6BAF9CE" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[238,493,1571,1595]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="411" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
|
||
<emphasis id="12B1740FFE7AFFC9FF76254CE6BAF9CE" box="[238,493,1571,1595]" italics="true" pageId="4" pageNumber="411">Arachnothelphusa rimba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0982C974FE7AFFC9FE6C254CE579F9CF" box="[500,558,1571,1594]" pageId="4" pageNumber="411" rank="species">n. sp.</taxonomicNameLabel>
|
||
, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088), Lanjak-Entimau. A, pleon; B, vulvae.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="68DFFB96FE7AFFC8FF5F25E2E61DFE90" lastPageId="5" lastPageNumber="412" pageId="4" pageNumber="411" type="diagnosis">
|
||
<paragraph id="207AA81DFE7AFFC8FF5F25E2E61DFE90" blockId="4.[151,1437,1677,2028]" lastBlockId="5.[151,1437,150,357]" lastPageId="5" lastPageNumber="412" pageId="4" pageNumber="411">
|
||
<emphasis id="12B1740FFE7AFFC9FF5F25E2E66DF952" bold="true" box="[199,314,1677,1703]" pageId="4" pageNumber="411">Diagnosis</emphasis>
|
||
. Carapace with dorsal surface gently surface punctate, rugose, finely granular; branchial regions gently inflated; epigastric and postorbital cristae distinct; cervical and H-shaped gastric grooves deep, not confluent; anterolateral margins distinctly convex, finely granulate but not serrate; antero- and posterolateral regions rugose; epibranchial tooth very low to not visible, barely separated from external orbital tooth; external orbital tooth acutely triangular, outer margin almost straight, subequal in length to inner margin (
|
||
<figureCitation id="B8FEB498FE7AFFC9FC782472E316F8C2" box="[992,1089,1821,1847]" captionStart="FIG" captionStartId="2.[151,196,1945,1969]" captionTargetBox="[275,1312,882,1919]" captionTargetId="figure-290@2.[275,1312,882,1919]" captionTargetPageId="2" captionText="FIG. 1. Arachnothelphusa rimba n. sp., holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5222297" httpUri="https://zenodo.org/record/5222297/files/figure.png" pageId="4" pageNumber="411">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="B8FEB498FE7AFFC9FBD42472E3C4F8C2" box="[1100,1171,1821,1847]" captionStart="FIG" captionStartId="3.[151,196,1179,1203]" captionTargetBox="[151,1436,181,1155]" captionTargetId="figure-15@3.[151,1436,181,1155]" captionTargetPageId="3" captionText="FIG. 2. Arachnothelphusa rimba n. sp. A, B, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088); C, male M1 (19.1 × 13.9 mm) (ZRC 2020.0086); D, male (20.2 × 15.4 mm) (ZRC 2020.0087).A, overall dorsal view; B–D, dorsal view of carapace. All specimens from Lanjak-Entimau." figureDoi="http://doi.org/10.5281/zenodo.5222299" httpUri="https://zenodo.org/record/5222299/files/figure.png" pageId="4" pageNumber="411">2B–D</figureCitation>
|
||
). Basis-ischium of third maxilliped rectangular, long, length to width ratio 1.78; oblique median sulcus deep (
|
||
<figureCitation id="B8FEB498FE7AFFC9FBFE242EE396F8AE" box="[1126,1217,1857,1883]" captionStart="FIG" captionStartId="2.[151,196,1945,1969]" captionTargetBox="[275,1312,882,1919]" captionTargetId="figure-290@2.[275,1312,882,1919]" captionTargetPageId="2" captionText="FIG. 1. Arachnothelphusa rimba n. sp., holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5222297" httpUri="https://zenodo.org/record/5222297/files/figure.png" pageId="4" pageNumber="411">Fig. 1D</figureCitation>
|
||
). Outer surfaces of chelipeds rugose; carpus rugose, with granules, inner angle with triangular tooth (
|
||
<figureCitation id="B8FEB498FE7AFFC9FBB5240AE39AF875" box="[1069,1229,1893,1920]" captionStart="FIG" captionStartId="2.[151,196,1945,1969]" captionTargetBox="[275,1312,882,1919]" captionTargetId="figure-290@2.[275,1312,882,1919]" captionTargetPageId="2" captionText="FIG. 1. Arachnothelphusa rimba n. sp., holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5222297" httpUri="https://zenodo.org/record/5222297/files/figure.png" pageId="4" pageNumber="411">Fig. 1A, G, H</figureCitation>
|
||
). Posterior margin of epistome wide, median love distinctly triangular (
|
||
<figureCitation id="B8FEB498FE7AFFC9FD4624E6E463F856" box="[734,820,1929,1955]" captionStart="FIG" captionStartId="2.[151,196,1945,1969]" captionTargetBox="[275,1312,882,1919]" captionTargetId="figure-290@2.[275,1312,882,1919]" captionTargetPageId="2" captionText="FIG. 1. Arachnothelphusa rimba n. sp., holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5222297" httpUri="https://zenodo.org/record/5222297/files/figure.png" pageId="4" pageNumber="411">Fig. 1C</figureCitation>
|
||
). P2–P5 very long; merus unarmed, dorsal margin gently gently serrated; outer surface gently rugose; P5 merus slightly longer than length of carapace; length to width ratio of P4 and P5 merus ca. 5.27 and 5.20, respectively, for males, ca. 5.79 and 5.92, respectively, for female (
|
||
<figureCitation id="B8FEB498FE7AFFC8FAFC24BEE7EBFF44" captionStart="FIG" captionStartId="2.[151,196,1945,1969]" captionTargetBox="[275,1312,882,1919]" captionTargetId="figure-290@2.[275,1312,882,1919]" captionTargetPageId="2" captionText="FIG. 1. Arachnothelphusa rimba n. sp., holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5222297" httpUri="https://zenodo.org/record/5222297/files/figure.png" lastPageId="5" lastPageNumber="412" pageId="4" pageNumber="411">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="B8FEB498FE7BFFC8FF5023F8E7BAFF44" box="[200,237,151,177]" captionStart="FIG" captionStartId="3.[151,196,1179,1203]" captionTargetBox="[151,1436,181,1155]" captionTargetId="figure-15@3.[151,1436,181,1155]" captionTargetPageId="3" captionText="FIG. 2. Arachnothelphusa rimba n. sp. A, B, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088); C, male M1 (19.1 × 13.9 mm) (ZRC 2020.0086); D, male (20.2 × 15.4 mm) (ZRC 2020.0087).A, overall dorsal view; B–D, dorsal view of carapace. All specimens from Lanjak-Entimau." figureDoi="http://doi.org/10.5281/zenodo.5222299" httpUri="https://zenodo.org/record/5222299/files/figure.png" pageId="5" pageNumber="412">2A</figureCitation>
|
||
,
|
||
<figureCitation id="B8FEB498FE7BFFC8FF6123F8E65EFF44" box="[249,265,151,177]" captionStart="FIG" captionStartId="3.[151,196,1944,1968]" captionTargetBox="[151,1436,1300,1921]" captionTargetId="figure-66@3.[151,1436,1300,1921]" captionTargetPageId="3" captionText="FIG. 3. Arachnothelphusa rimba n. sp., ambulatory legs. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau; C, D, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088), Lanjak-Entimau. A, C, right P4; B, D, right P5." figureDoi="http://doi.org/10.5281/zenodo.5222301" httpUri="https://zenodo.org/record/5222301/files/figure.png" pageId="5" pageNumber="412">3</figureCitation>
|
||
). Male pleon distinctly T-shaped, lateral margins of somites 5 and 6 distinctly concave; somite 6 longer than telson; lateral margins of telson gently concave (
|
||
<figureCitation id="B8FEB498FE7BFFC8FC9823D4E42EFF20" box="[768,889,187,213]" captionStart="FIG" captionStartId="2.[151,196,1945,1969]" captionTargetBox="[275,1312,882,1919]" captionTargetId="figure-290@2.[275,1312,882,1919]" captionTargetPageId="2" captionText="FIG. 1. Arachnothelphusa rimba n. sp., holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5222297" httpUri="https://zenodo.org/record/5222297/files/figure.png" pageId="5" pageNumber="412">Fig. 1E, F</figureCitation>
|
||
). G1 slender, sinuous, gently curving outwards; terminal segment relatively short, proximal part cylindrical, distal part suddenly tapering to long slender point, about 0.33 times length of subterminal segment (
|
||
<figureCitation id="B8FEB498FE7BFFC8FD21226CE462FEE8" box="[697,821,259,285]" captionStart="FIG" captionStartId="4.[151,196,1073,1097]" captionTargetBox="[161,1426,188,1041]" captionTargetId="figure-24@4.[151,1436,181,1049]" captionTargetPageId="4" captionText="FIG. 4. Arachnothelphusa rimba n. sp. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsal view); D, left G2. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5222303" httpUri="https://zenodo.org/record/5222303/files/figure.png" pageId="5" pageNumber="412">Fig. 4A–C</figureCitation>
|
||
). G2 with short distal segment, about 0.27 times length of basal segment (
|
||
<figureCitation id="B8FEB498FE7BFFC8FEFF2248E693FEB4" box="[359,452,295,321]" captionStart="FIG" captionStartId="4.[151,196,1073,1097]" captionTargetBox="[161,1426,188,1041]" captionTargetId="figure-24@4.[151,1436,181,1049]" captionTargetPageId="4" captionText="FIG. 4. Arachnothelphusa rimba n. sp. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsal view); D, left G2. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5222303" httpUri="https://zenodo.org/record/5222303/files/figure.png" pageId="5" pageNumber="412">Fig. 4D</figureCitation>
|
||
). Vulvae large, occupying submedian part of sternite 6, without obvious sternal vulvar cover (
|
||
<figureCitation id="B8FEB498FE7BFFC8FF7D2224E66AFE90" box="[229,317,330,357]" captionStart="FIG" captionStartId="4.[151,196,1571,1595]" captionTargetBox="[151,1436,1158,1547]" captionTargetId="figure-74@4.[151,1436,1158,1547]" captionTargetPageId="4" captionText="FIG. 5. Arachnothelphusa rimba n. sp., paratype female (22.9 × 16.5 mm) (ZRC 2020.0088), Lanjak-Entimau. A, pleon; B, vulvae." figureDoi="http://doi.org/10.5281/zenodo.5222305" httpUri="https://zenodo.org/record/5222305/files/figure.png" pageId="5" pageNumber="412">Fig. 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="74BAF895FE7BFFC8FF0F25F6E336F90C" ID-DOI="http://doi.org/10.5281/zenodo.5222307" ID-Zenodo-Dep="5222307" httpUri="https://zenodo.org/record/5222307/files/figure.png" pageId="5" pageNumber="412" startId="5.[151,196,1689,1713]" targetBox="[151,1436,381,1666]" targetPageId="5">
|
||
<paragraph id="207AA81DFE7BFFC8FF0F25F6E336F90C" blockId="5.[151,1436,1689,1786]" pageId="5" pageNumber="412">
|
||
<emphasis id="12B1740FFE7BFFC8FF0F25F6E7B3F944" bold="true" box="[151,228,1689,1714]" pageId="5" pageNumber="412">FIG. 6.</emphasis>
|
||
<taxonomicName id="E7C5D39EFE7BFFC8FF7225F6E6B0F944" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[234,487,1689,1713]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="412" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
|
||
<emphasis id="12B1740FFE7BFFC8FF7225F6E6B0F944" box="[234,487,1689,1713]" italics="true" pageId="5" pageNumber="412">Arachnothelphusa rimba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0982C974FE7BFFC8FE7525F5E572F944" box="[493,549,1690,1713]" pageId="5" pageNumber="412" rank="species">n. sp.</taxonomicNameLabel>
|
||
, colour in life. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366); C–E, paratype male (20.2 × 15.4 mm) (ZRC 2020.0087), F, paratype male (19.1 × 13.9 mm) (ZRC 2020.0086); G, H, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088). All specimens from Lanjak-Entimau. Photographs: Tan Heok Hui.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="68DFFB96FE7BFFC8FF5F2448E422F827" pageId="5" pageNumber="412" type="description">
|
||
<paragraph id="207AA81DFE7BFFC8FF5F2448E422F827" blockId="5.[151,1437,1831,2038]" pageId="5" pageNumber="412">
|
||
<emphasis id="12B1740FFE7BFFC8FF5F2448E66FF8B4" bold="true" box="[199,312,1831,1857]" pageId="5" pageNumber="412">Variation</emphasis>
|
||
.
|
||
<materialsCitation id="90ADA240FE7BFFC8FEDE2447E425F827" collectionCode="ZRC" pageId="5" pageNumber="412" specimenCode="ZRC 2020.0087, ZRC 2020.0088" specimenCount="4" typeStatus="paratype">
|
||
One of the
|
||
<typeStatus id="FF7E16BFFE7BFFC8FE5C2447E573F8B7" box="[452,548,1832,1858]" pageId="5" pageNumber="412" type="paratype">paratype</typeStatus>
|
||
male specimens (20.2 ×
|
||
<quantity id="E73D05F8FE7BFFC8FCA72448E4F2F8B7" box="[831,933,1831,1858]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="5" pageNumber="412" unit="mm" value="15.4">15.4 mm</quantity>
|
||
,
|
||
<specimenCode id="70630066FE7BFFC8FC292447E33FF8B7" box="[945,1128,1832,1858]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="412">ZRC 2020.0087</specimenCode>
|
||
) has a relatively more rectangular carapace than the other specimens, with the branchial regions less prominently swollen laterally (
|
||
<figureCitation id="B8FEB498FE7BFFC8FAAD2423E2C7F893" box="[1333,1424,1868,1894]" captionStart="FIG" captionStartId="3.[151,196,1179,1203]" captionTargetBox="[151,1436,181,1155]" captionTargetId="figure-15@3.[151,1436,181,1155]" captionTargetPageId="3" captionText="FIG. 2. Arachnothelphusa rimba n. sp. A, B, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088); C, male M1 (19.1 × 13.9 mm) (ZRC 2020.0086); D, male (20.2 × 15.4 mm) (ZRC 2020.0087).A, overall dorsal view; B–D, dorsal view of carapace. All specimens from Lanjak-Entimau." figureDoi="http://doi.org/10.5281/zenodo.5222299" httpUri="https://zenodo.org/record/5222299/files/figure.png" pageId="5" pageNumber="412">Fig. 2D</figureCitation>
|
||
). In all other aspects, however, it agrees with the other types; its G1 structure being identical to the
|
||
<typeStatus id="FF7E16BFFE7BFFC8FB4E241FE26BF87F" box="[1238,1340,1904,1930]" pageId="5" pageNumber="412" type="holotype">holotype</typeStatus>
|
||
. The left side of the anterolateral margin of the
|
||
<typeStatus id="FF7E16BFFE7BFFC8FDDC24FBE5F3F85B" box="[580,676,1940,1966]" pageId="5" pageNumber="412" type="paratype">paratype</typeStatus>
|
||
female (22.9 ×
|
||
<quantity id="E73D05F8FE7BFFC8FCCF24FCE4EAF85B" box="[855,957,1939,1966]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="5" pageNumber="412" unit="mm" value="16.5">16.5 mm</quantity>
|
||
,
|
||
<specimenCode id="70630066FE7BFFC8FC5124FBE328F85B" box="[969,1151,1940,1966]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="412">ZRC 2020.0088</specimenCode>
|
||
) is damaged, with a deep cleft, suggesting there had been an injury in the past (
|
||
<figureCitation id="B8FEB498FE7BFFC8FD7624D7E430F827" box="[750,871,1976,2002]" captionStart="FIG" captionStartId="3.[151,196,1179,1203]" captionTargetBox="[151,1436,181,1155]" captionTargetId="figure-15@3.[151,1436,181,1155]" captionTargetPageId="3" captionText="FIG. 2. Arachnothelphusa rimba n. sp. A, B, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088); C, male M1 (19.1 × 13.9 mm) (ZRC 2020.0086); D, male (20.2 × 15.4 mm) (ZRC 2020.0087).A, overall dorsal view; B–D, dorsal view of carapace. All specimens from Lanjak-Entimau." figureDoi="http://doi.org/10.5281/zenodo.5222299" httpUri="https://zenodo.org/record/5222299/files/figure.png" pageId="5" pageNumber="412">Fig. 2A, B</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68DFFB96FE7BFFCBFF5F24B3E78AFE91" lastPageId="6" lastPageNumber="413" pageId="5" pageNumber="412" type="etymology">
|
||
<paragraph id="207AA81DFE7BFFCBFF5F24B3E599FF21" blockId="5.[151,1437,1831,2038]" lastBlockId="6.[151,1437,150,969]" lastPageId="6" lastPageNumber="413" pageId="5" pageNumber="412">
|
||
<emphasis id="12B1740FFE7BFFC8FF5F24B3E61FF803" bold="true" box="[199,328,2012,2038]" pageId="5" pageNumber="412">Etymology</emphasis>
|
||
. The species name is derived from the initiative that resulted in its discovery – Research for Intensified Management of Bio-Rich Areas of
|
||
<collectingCountry id="58D2E88DFE78FFCBFDCB23F9E5E4FF45" box="[595,691,150,176]" name="Malaysia" pageId="6" pageNumber="413">Sarawak</collectingCountry>
|
||
(RIMBA) which was started by the
|
||
<collectingCountry id="58D2E88DFE78FFCBFBD223F9E3FDFF45" box="[1098,1194,150,176]" name="Malaysia" pageId="6" pageNumber="413">Sarawak</collectingCountry>
|
||
Forestry Corporation in 2015. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="207AA81DFE78FFCBFF5F23B1E78AFE91" blockId="6.[151,1437,150,969]" pageId="6" pageNumber="413">
|
||
<emphasis id="12B1740FFE78FFCBFF5F23B1E64CFF0D" bold="true" box="[199,283,222,248]" pageId="6" pageNumber="413">Colour</emphasis>
|
||
. In life, the carapace is brownish to brownish purple (
|
||
<figureCitation id="B8FEB498FE78FFCBFCF323B1E4F9FF0C" box="[875,942,222,249]" captionStart="FIG" captionStartId="5.[151,196,1689,1713]" captionTargetBox="[151,1436,381,1666]" captionTargetId="figure-104@5.[151,1436,381,1666]" captionTargetPageId="5" captionText="FIG. 6. Arachnothelphusa rimba n. sp., colour in life. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366); C–E, paratype male (20.2 × 15.4 mm) (ZRC 2020.0087), F, paratype male (19.1 × 13.9 mm) (ZRC 2020.0086); G, H, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088).All specimens from Lanjak-Entimau. Photographs: Tan Heok Hui." figureDoi="http://doi.org/10.5281/zenodo.5222307" httpUri="https://zenodo.org/record/5222307/files/figure.png" pageId="6" pageNumber="413">Fig. 6</figureCitation>
|
||
). The chelae and proximal part of the fingers are purplish-brown, with the distal third of the fingers cream to pale-yellow (
|
||
<figureCitation id="B8FEB498FE78FFCBFC76226DE3F2FEE8" box="[1006,1189,258,285]" captionStart="FIG" captionStartId="5.[151,196,1689,1713]" captionTargetBox="[151,1436,381,1666]" captionTargetId="figure-104@5.[151,1436,381,1666]" captionTargetPageId="5" captionText="FIG. 6. Arachnothelphusa rimba n. sp., colour in life. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366); C–E, paratype male (20.2 × 15.4 mm) (ZRC 2020.0087), F, paratype male (19.1 × 13.9 mm) (ZRC 2020.0086); G, H, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088).All specimens from Lanjak-Entimau. Photographs: Tan Heok Hui." figureDoi="http://doi.org/10.5281/zenodo.5222307" httpUri="https://zenodo.org/record/5222307/files/figure.png" pageId="6" pageNumber="413">Fig. 6B, D–F, H</figureCitation>
|
||
). The ambulatory legs are purplish-brown, with the distal third of the dactylus pale yellow (
|
||
<figureCitation id="B8FEB498FE78FFCBFC362249E4A2FEB4" box="[942,1013,294,321]" captionStart="FIG" captionStartId="5.[151,196,1689,1713]" captionTargetBox="[151,1436,381,1666]" captionTargetId="figure-104@5.[151,1436,381,1666]" captionTargetPageId="5" captionText="FIG. 6. Arachnothelphusa rimba n. sp., colour in life. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366); C–E, paratype male (20.2 × 15.4 mm) (ZRC 2020.0087), F, paratype male (19.1 × 13.9 mm) (ZRC 2020.0086); G, H, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088).All specimens from Lanjak-Entimau. Photographs: Tan Heok Hui." figureDoi="http://doi.org/10.5281/zenodo.5222307" httpUri="https://zenodo.org/record/5222307/files/figure.png" pageId="6" pageNumber="413">Fig. 6</figureCitation>
|
||
). The ventral surfaces are yellowishwhite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68DFFB96FE78FFCBFF5F2201E266FC3C" pageId="6" pageNumber="413" type="discussion">
|
||
<paragraph id="207AA81DFE78FFCBFF5F2201E36DFD05" blockId="6.[151,1437,150,969]" pageId="6" pageNumber="413">
|
||
<emphasis id="12B1740FFE78FFCBFF5F2201E663FE7D" bold="true" box="[199,308,366,392]" pageId="6" pageNumber="413">Remarks</emphasis>
|
||
. The relatively flatter carapace allies
|
||
<taxonomicName id="E7C5D39EFE78FFCBFD4C2200E462FE7D" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[724,821,366,393]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
|
||
<emphasis id="12B1740FFE78FFCBFD4C2200E462FE7D" box="[724,821,366,393]" italics="true" pageId="6" pageNumber="413">A. rimba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0982C974FE78FFCBFCA32200E421FE7C" box="[827,886,367,393]" pageId="6" pageNumber="413" rank="species">n. sp.</taxonomicNameLabel>
|
||
with
|
||
<taxonomicName id="E7C5D39EFE78FFCBFC2C2200E378FE7D" authorityName="Ng" authorityYear="1991" box="[948,1071,367,393]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="terrapes">
|
||
<emphasis id="12B1740FFE78FFCBFC2C2200E378FE7D" box="[948,1071,367,393]" italics="true" pageId="6" pageNumber="413">A. terrapes</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="E20166FFFE78FFCBFBEA2201E3EDFE7D" box="[1138,1210,366,392]" country="Malaysia" name="Sabah" pageId="6" pageNumber="413">Sabah</collectingRegion>
|
||
. They can, however, easily be separated;
|
||
<taxonomicName id="E7C5D39EFE78FFCBFEEF22FCE68FFE59" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[375,472,402,429]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
|
||
<emphasis id="12B1740FFE78FFCBFEEF22FCE68FFE59" box="[375,472,402,429]" italics="true" pageId="6" pageNumber="413">A. rimba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0982C974FE78FFCBFE4622FCE54EFE58" box="[478,537,403,429]" pageId="6" pageNumber="413" rank="species">n. sp.</taxonomicNameLabel>
|
||
has the anterolateral margin entire (
|
||
<figureCitation id="B8FEB498FE78FFCBFC3022FDE35EFE59" box="[936,1033,402,428]" captionStart="FIG" captionStartId="2.[151,196,1945,1969]" captionTargetBox="[275,1312,882,1919]" captionTargetId="figure-290@2.[275,1312,882,1919]" captionTargetPageId="2" captionText="FIG. 1. Arachnothelphusa rimba n. sp., holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5222297" httpUri="https://zenodo.org/record/5222297/files/figure.png" pageId="6" pageNumber="413">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="B8FEB498FE78FFCBFB8D22FDE30EFE59" box="[1045,1113,402,428]" captionStart="FIG" captionStartId="3.[151,196,1179,1203]" captionTargetBox="[151,1436,181,1155]" captionTargetId="figure-15@3.[151,1436,181,1155]" captionTargetPageId="3" captionText="FIG. 2. Arachnothelphusa rimba n. sp. A, B, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088); C, male M1 (19.1 × 13.9 mm) (ZRC 2020.0086); D, male (20.2 × 15.4 mm) (ZRC 2020.0087).A, overall dorsal view; B–D, dorsal view of carapace. All specimens from Lanjak-Entimau." figureDoi="http://doi.org/10.5281/zenodo.5222299" httpUri="https://zenodo.org/record/5222299/files/figure.png" pageId="6" pageNumber="413">2B–D</figureCitation>
|
||
) (versus with deep U-shaped notch in
|
||
<taxonomicName id="E7C5D39EFE78FFCBFF6022D8E624FE25" authorityName="Ng" authorityYear="1991" box="[248,371,439,465]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="terrapes">
|
||
<emphasis id="12B1740FFE78FFCBFF6022D8E624FE25" box="[248,371,439,465]" italics="true" pageId="6" pageNumber="413">A. terrapes</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="4454D5ECFE78FFCBFE3C22D8E55EFE25" author="Ng, P. K. L." box="[420,521,438,465]" pageId="6" pageNumber="413" pagination="1 - 12" refId="ref6389" refString="Ng, P. K. L. (1991) Bornean freshwater crabs of the genus Arachnothelphusa gen. nov. (Crustacea: Decapoda: Brachyura: Gecarcinucidae). Zoologische Mededelingen, 65 (1), 1 - 12." type="journal article" year="1991">Ng 1991</bibRefCitation>
|
||
: fig. 5); the palm and fingers of the adult male cheliped are proportionately shorter (
|
||
<figureCitation id="B8FEB498FE78FFCBFF3822B5E7A2FE01" box="[160,245,474,500]" captionStart="FIG" captionStartId="2.[151,196,1945,1969]" captionTargetBox="[275,1312,882,1919]" captionTargetId="figure-290@2.[275,1312,882,1919]" captionTargetPageId="2" captionText="FIG. 1. Arachnothelphusa rimba n. sp., holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5222297" httpUri="https://zenodo.org/record/5222297/files/figure.png" pageId="6" pageNumber="413">Fig. 1G</figureCitation>
|
||
) (versus palm and fingers longer and more slender; cf.
|
||
<bibRefCitation id="4454D5ECFE78FFCBFCC222B4E355FE01" author="Ng, P. K. L. & Ng, P. Y. C." box="[858,1026,474,501]" pageId="6" pageNumber="413" pagination="89 - 112" refId="ref6976" refString="Ng, P. K. L. & Ng, P. Y. C. (2018) The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). Zookeys, 760, 89 - 112. https: // doi. org / 10.3897 / zookeys. 760.24787" type="journal article" year="2018">Ng & Ng 2018</bibRefCitation>
|
||
: fig. 5D); the G1 terminal segment is straight but with the distal part sharply tapering to an acute tip (
|
||
<figureCitation id="B8FEB498FE78FFCBFCC42291E483FDED" box="[860,980,510,536]" captionStart="FIG" captionStartId="4.[151,196,1073,1097]" captionTargetBox="[161,1426,188,1041]" captionTargetId="figure-24@4.[151,1436,181,1049]" captionTargetPageId="4" captionText="FIG. 4. Arachnothelphusa rimba n. sp. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsal view); D, left G2. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5222303" httpUri="https://zenodo.org/record/5222303/files/figure.png" pageId="6" pageNumber="413">Fig. 4A–C</figureCitation>
|
||
) (versus terminal segment gently curved upwards in
|
||
<taxonomicName id="E7C5D39EFE78FFCBFE84214CE6CEFDC9" authorityName="Ng" authorityYear="1991" box="[284,409,547,573]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="terrapes">
|
||
<emphasis id="12B1740FFE78FFCBFE84214CE6CEFDC9" box="[284,409,547,573]" italics="true" pageId="6" pageNumber="413">A. terrapes</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="4454D5ECFE78FFCBFE55214CE564FDC9" author="Ng, P. K. L." box="[461,563,546,573]" pageId="6" pageNumber="413" pagination="1 - 12" refId="ref6389" refString="Ng, P. K. L. (1991) Bornean freshwater crabs of the genus Arachnothelphusa gen. nov. (Crustacea: Decapoda: Brachyura: Gecarcinucidae). Zoologische Mededelingen, 65 (1), 1 - 12." type="journal article" year="1991">Ng 1991</bibRefCitation>
|
||
: fig. 2D–G); and the G2 distal segment is relatively shorter (
|
||
<figureCitation id="B8FEB498FE78FFCBFB7B214DE26CFDC9" box="[1251,1339,546,572]" captionStart="FIG" captionStartId="4.[151,196,1073,1097]" captionTargetBox="[161,1426,188,1041]" captionTargetId="figure-24@4.[151,1436,181,1049]" captionTargetPageId="4" captionText="FIG. 4. Arachnothelphusa rimba n. sp. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsal view); D, left G2. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5222303" httpUri="https://zenodo.org/record/5222303/files/figure.png" pageId="6" pageNumber="413">Fig. 4D</figureCitation>
|
||
) (versus distal segment longer in
|
||
<taxonomicName id="E7C5D39EFE78FFCBFE362128E57CFD95" authorityName="Ng" authorityYear="1991" box="[430,555,583,609]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="terrapes">
|
||
<emphasis id="12B1740FFE78FFCBFE362128E57CFD95" box="[430,555,583,609]" italics="true" pageId="6" pageNumber="413">A. terrapes</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="4454D5ECFE78FFCBFDF82128E591FD95" author="Ng, P. K. L." box="[608,710,582,609]" pageId="6" pageNumber="413" pagination="1 - 12" refId="ref6389" refString="Ng, P. K. L. (1991) Bornean freshwater crabs of the genus Arachnothelphusa gen. nov. (Crustacea: Decapoda: Brachyura: Gecarcinucidae). Zoologische Mededelingen, 65 (1), 1 - 12." type="journal article" year="1991">Ng 1991</bibRefCitation>
|
||
: fig. 6H). The G1 of
|
||
<taxonomicName id="E7C5D39EFE78FFCBFC2F2128E34DFD95" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[951,1050,582,609]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
|
||
<emphasis id="12B1740FFE78FFCBFC2F2128E34DFD95" box="[951,1050,582,609]" italics="true" pageId="6" pageNumber="413">A. rimba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0982C974FE78FFCBFBBA2128E308FD94" box="[1058,1119,583,609]" pageId="6" pageNumber="413" rank="species">n. sp.</taxonomicNameLabel>
|
||
superficially resembles that of
|
||
<taxonomicName id="E7C5D39EFE78FFCBFF202104E619FD71" baseAuthorityName="De Man" baseAuthorityYear="1899" box="[184,334,618,645]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="melanippe">
|
||
<emphasis id="12B1740FFE78FFCBFF202104E619FD71" box="[184,334,618,645]" italics="true" pageId="6" pageNumber="413">A. melanippe</emphasis>
|
||
</taxonomicName>
|
||
but that of the latter is proportionately almost straight and is cone-shaped, gradually tapering to a sharp point (
|
||
<bibRefCitation id="4454D5ECFE78FFCBFEA221E1E6F6FD5D" author="Ng, P. K. L." box="[314,417,654,680]" pageId="6" pageNumber="413" pagination="1 - 12" refId="ref6389" refString="Ng, P. K. L. (1991) Bornean freshwater crabs of the genus Arachnothelphusa gen. nov. (Crustacea: Decapoda: Brachyura: Gecarcinucidae). Zoologische Mededelingen, 65 (1), 1 - 12." type="journal article" year="1991">Ng 1991</bibRefCitation>
|
||
: fig. 2C–F); in
|
||
<taxonomicName id="E7C5D39EFE78FFCBFDC821E0E5E4FD5D" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[592,691,654,681]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
|
||
<emphasis id="12B1740FFE78FFCBFDC821E0E5E4FD5D" box="[592,691,654,681]" italics="true" pageId="6" pageNumber="413">A. rimba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0982C974FE78FFCBFD2321E0E5ADFD5C" box="[699,762,655,681]" pageId="6" pageNumber="413" rank="species">n. sp.</taxonomicNameLabel>
|
||
, the G1 terminal segment is straight but with the distal part sharply tapering to an acute tip (
|
||
<figureCitation id="B8FEB498FE78FFCBFD9A21DDE529FD39" box="[514,638,690,716]" captionStart="FIG" captionStartId="4.[151,196,1073,1097]" captionTargetBox="[161,1426,188,1041]" captionTargetId="figure-24@4.[151,1436,181,1049]" captionTargetPageId="4" captionText="FIG. 4. Arachnothelphusa rimba n. sp. A, B, holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsal view); D, left G2. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5222303" httpUri="https://zenodo.org/record/5222303/files/figure.png" pageId="6" pageNumber="413">Fig. 4A–C</figureCitation>
|
||
). Unlike
|
||
<taxonomicName id="E7C5D39EFE78FFCBFD7D21DCE410FD39" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[741,839,690,717]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="rimba" status="sp. nov.">
|
||
<emphasis id="12B1740FFE78FFCBFD7D21DCE410FD39" box="[741,839,690,717]" italics="true" pageId="6" pageNumber="413">A. rimba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0982C974FE78FFCBFCD621DCE4DDFD38" box="[846,906,691,717]" pageId="6" pageNumber="413" rank="species">n. sp.</taxonomicNameLabel>
|
||
which has an entire anterolateral margin (
|
||
<figureCitation id="B8FEB498FE78FFCBFAFC21DDE7EEFD05" captionStart="FIG" captionStartId="2.[151,196,1945,1969]" captionTargetBox="[275,1312,882,1919]" captionTargetId="figure-290@2.[275,1312,882,1919]" captionTargetPageId="2" captionText="FIG. 1. Arachnothelphusa rimba n. sp., holotype male (22.8 × 16.6 mm) (ZRC 2020.0366), Lanjak-Entimau. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, anterior thoracic sternum and pleon; F, pleon; G, outer view of right chela; H, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5222297" httpUri="https://zenodo.org/record/5222297/files/figure.png" pageId="6" pageNumber="413">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="B8FEB498FE78FFCBFF5E21B9E65AFD05" box="[198,269,726,752]" captionStart="FIG" captionStartId="3.[151,196,1179,1203]" captionTargetBox="[151,1436,181,1155]" captionTargetId="figure-15@3.[151,1436,181,1155]" captionTargetPageId="3" captionText="FIG. 2. Arachnothelphusa rimba n. sp. A, B, paratype female (22.9 × 16.5 mm) (ZRC 2020.0088); C, male M1 (19.1 × 13.9 mm) (ZRC 2020.0086); D, male (20.2 × 15.4 mm) (ZRC 2020.0087).A, overall dorsal view; B–D, dorsal view of carapace. All specimens from Lanjak-Entimau." figureDoi="http://doi.org/10.5281/zenodo.5222299" httpUri="https://zenodo.org/record/5222299/files/figure.png" pageId="6" pageNumber="413">2B–D</figureCitation>
|
||
), that of
|
||
<taxonomicName id="E7C5D39EFE78FFCBFEF721B8E554FD05" baseAuthorityName="De Man" baseAuthorityYear="1899" box="[367,515,726,753]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="melanippe">
|
||
<emphasis id="12B1740FFE78FFCBFEF721B8E554FD05" box="[367,515,726,753]" italics="true" pageId="6" pageNumber="413">A. melanippe</emphasis>
|
||
</taxonomicName>
|
||
has a distinct epibranchial tooth (
|
||
<bibRefCitation id="4454D5ECFE78FFCBFCE521B9E4B3FD05" author="Ng, P. K. L." box="[893,996,726,752]" pageId="6" pageNumber="413" pagination="1 - 12" refId="ref6389" refString="Ng, P. K. L. (1991) Bornean freshwater crabs of the genus Arachnothelphusa gen. nov. (Crustacea: Decapoda: Brachyura: Gecarcinucidae). Zoologische Mededelingen, 65 (1), 1 - 12." type="journal article" year="1991">Ng 1991</bibRefCitation>
|
||
: fig. 1).
|
||
</paragraph>
|
||
<paragraph id="207AA81DFE78FFCBFF5F2195E266FC3C" blockId="6.[151,1437,150,969]" pageId="6" pageNumber="413">
|
||
The first report of crabs from the Lanjak-Entimau Wildlife Sanctuary was by
|
||
<bibRefCitation id="4454D5ECFE78FFCBFBA42194E395FCE0" author="Ng, P. K. L." box="[1084,1218,762,789]" pageId="6" pageNumber="413" pagination="57 - 72" refId="ref6489" refString="Ng, P. K. L. (1995 b) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng (1995b)</bibRefCitation>
|
||
, who recorded two potamids,
|
||
<taxonomicName id="E7C5D39EFE78FFCBFE952071E571FCCC" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[269,550,798,825]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="aethes">
|
||
<emphasis id="12B1740FFE78FFCBFE952071E6E5FCCD" box="[269,434,798,824]" italics="true" pageId="6" pageNumber="413">Ibanum aethes</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E7C5D39EFE78FFCBFDA92071E4F9FCCC" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[561,942,798,825]" class="Malacostraca" family="Potamidae" genus="Isolapotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="stuebingi">
|
||
<emphasis id="12B1740FFE78FFCBFDA92071E46DFCCD" box="[561,826,798,824]" italics="true" pageId="6" pageNumber="413">Isolapotamon stuebingi</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
; and one sesarmid
|
||
<taxonomicName id="E7C5D39EFE78FFCBFB1E2070E785FCA8" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="12B1740FFE78FFCBFB1E2070E23CFCCD" box="[1158,1387,798,825]" italics="true" pageId="6" pageNumber="413">Geosesarma katibas</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="E7C5D39EFE78FFCBFF47202DE6BEFCA9" authorityName="Ng" authorityYear="1995" box="[223,489,834,860]" class="Malacostraca" family="Potamidae" genus="Isolapotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="stuebingi">
|
||
<emphasis id="12B1740FFE78FFCBFF47202DE6BEFCA9" box="[223,489,834,860]" italics="true" pageId="6" pageNumber="413">Isolapotamon stuebingi</emphasis>
|
||
</taxonomicName>
|
||
has since been synonymised under
|
||
<taxonomicName id="E7C5D39EFE78FFCBFC1F202CE327FCA9" authority="Ng, 1987" authorityName="Ng" authorityYear="1987" box="[903,1136,834,861]" class="Malacostraca" family="Potamidae" genus="Isolapotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="nimboni">
|
||
<emphasis id="12B1740FFE78FFCBFC1F202CE4ABFCA9" box="[903,1020,834,860]" italics="true" pageId="6" pageNumber="413">I. nimboni</emphasis>
|
||
Ng, 1987
|
||
</taxonomicName>
|
||
(Ng 2004;
|
||
<bibRefCitation id="4454D5ECFE78FFCBFB6E202CE783FC75" author="Ng, P. K. L. & Grinang, J." pageId="6" pageNumber="413" pagination="299 - 325" refId="ref6764" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity, Sarawak Museum Journal, New Series, 59 (Special Issue No. 6), 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
).
|
||
<taxonomicName id="E7C5D39EFE78FFCBFF702009E6D9FC75" authorityName="Ng" authorityYear="1995" box="[232,398,870,896]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="aethes">
|
||
<emphasis id="12B1740FFE78FFCBFF702009E6D9FC75" box="[232,398,870,896]" italics="true" pageId="6" pageNumber="413">Ibanum aethes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E7C5D39EFE78FFCBFE052009E5CBFC75" authorityName="Ng" authorityYear="1987" box="[413,668,870,896]" class="Malacostraca" family="Potamidae" genus="Isolapotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="nimboni">
|
||
<emphasis id="12B1740FFE78FFCBFE052009E5CBFC75" box="[413,668,870,896]" italics="true" pageId="6" pageNumber="413">Isolapotamon nimboni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E7C5D39EFE78FFCBFD4D2008E4ECFC75" authorityName="Ng" authorityYear="1995" box="[725,955,870,897]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="12B1740FFE78FFCBFD4D2008E4ECFC75" box="[725,955,870,897]" italics="true" pageId="6" pageNumber="413">Geosesarma katibas</emphasis>
|
||
</taxonomicName>
|
||
were collected in the survey of 2019, and addition, two other gecarcinucids,
|
||
<taxonomicName id="E7C5D39EFE78FFCBFD8520E5E37EFC50" authority="(von Martens, 1868)" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[541,1065,906,933]" class="Malacostraca" family="Gecarcinucidae" genus="Perithelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis id="12B1740FFE78FFCBFD8520E5E46CFC51" box="[541,827,906,932]" italics="true" pageId="6" pageNumber="413">Perithelphusa borneensis</emphasis>
|
||
(von
|
||
<bibRefCitation id="4454D5ECFE78FFCBFCE620E4E376FC50" author="Martens, E. von" box="[894,1057,906,933]" pageId="6" pageNumber="413" pagination="1 - 64" refId="ref5985" refString="Martens, E. von (1868) Ueber einige ostasiatische Susswasserthiere. Archiv fur Naturgeschichte, 34 (1), 1 - 64, pl. 1. https: // doi. org / 10.5962 / bhl. part. 20475" type="journal article" year="1868">Martens, 1868</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and a new species of
|
||
<taxonomicName id="E7C5D39EFE78FFCBFABB20E5E60DFC3D" authority="Bott, 1969" authorityName="Bott" authorityYear="1969" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="413" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="12B1740FFE78FFCBFABB20E5E78DFC3D" italics="true" pageId="6" pageNumber="413">Sundathelphusa</emphasis>
|
||
<bibRefCitation id="4454D5ECFE78FFCBFF7920C0E60DFC3D" author="Bott, R." box="[225,346,942,969]" pageId="6" pageNumber="413" pagination="359 - 366" refId="ref5157" refString="Bott, R. (1969) Flusskrabben aus Asien und ihre Klassifikation (Crustacea, Decapoda). Senckenbergiana biologica, 50 (5 - 6), 359 - 366." type="journal article" year="1969">Bott, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
, were also found. The latter will be treated later in a revision of the genus from Borneo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |