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<document id="08E270270CD3969E70EB9C38D648B24E" ID-CLB-Dataset="28046" ID-DOI="10.11646/zootaxa.4508.2.8" ID-GBIF-Dataset="71421820-a02b-4fef-a8b6-e8b216c3e8f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607072" ID-ZooBank="3375CF8C-7952-4CF5-8E39-09DE2197119D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585884865" checkinUser="plazi" docAuthor="Wang, Yang, Dietrich, Christopher H. &amp; Zhang, Yalin" docDate="2018" docId="9A1987B1F20F2438D3BC5CDDB1F4FBFD" docLanguage="en" docName="zootaxa.4508.2.8.pdf" docOrigin="Zootaxa 4508 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Riseveinus hamatus Wang &amp; Dietrich &amp; Zhang 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="261" masterDocId="6620FFC9F20D243AD32B5D06B222FFAA" masterDocTitle="Four new species of the leafhopper genus Riseveinus Li (Hemiptera: Cicadellidae: Evacanthinae) from China and Thailand" masterLastPageNumber="266" masterPageNumber="259" pageNumber="261" updateTime="1698713016928" updateUser="ExternalLinkService">
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<mods:title id="FCCFD23CA1FC719EC529165285F01908">Four new species of the leafhopper genus Riseveinus Li (Hemiptera: Cicadellidae: Evacanthinae) from China and Thailand</mods:title>
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<mods:namePart id="B001A0C9FAF21420CC828C635591ACF9">Wang, Yang</mods:namePart>
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<mods:namePart id="2A44BAB5AE17F49389E50EA139F2E0B0">Dietrich, Christopher H.</mods:namePart>
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<mods:namePart id="029AB334A06EB687DB61B636A1D3031D">Zhang, Yalin</mods:namePart>
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<mods:title id="24DE12A0660D632ADE14EE75E4352EDF">Zootaxa</mods:title>
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<mods:date id="49465706D439DC88F894D09DEFF7A6CD">2018</mods:date>
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<mods:number id="A72C7097EEA1753A7032CF1DFA75E354">2018-10-31</mods:number>
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<treatment id="9A1987B1F20F2438D3BC5CDDB1F4FBFD" ID-DOI="http://doi.org/10.5281/zenodo.5957449" ID-GBIF-Taxon="154495798" ID-Zenodo-Dep="5957449" LSID="urn:lsid:plazi:treatment:9A1987B1F20F2438D3BC5CDDB1F4FBFD" httpUri="http://treatment.plazi.org/id/9A1987B1F20F2438D3BC5CDDB1F4FBFD" lastPageNumber="261" pageId="2" pageNumber="261">
<subSubSection id="5AAA652CF20F2438D3BC5CDDB3CFFE5C" box="[151,493,475,502]" pageId="2" pageNumber="261" type="nomenclature">
<paragraph id="120F36A7F20F2438D3BC5CDDB3CFFE5C" blockId="2.[151,493,475,535]" box="[151,493,475,502]" pageId="2" pageNumber="261">
<heading id="494781CBF20F2438D3BC5CDDB3CFFE5C" bold="true" box="[151,493,475,502]" fontSize="11" level="1" pageId="2" pageNumber="261" reason="1">
<emphasis id="20C4EAB5F20F2438D3BC5CDDB3CFFE5C" bold="true" box="[151,493,475,502]" pageId="2" pageNumber="261">
<taxonomicName id="D5B04D24F20F2438D3BC5CDDB3AAFE5F" ID-CoL="8JYS2" authority="Wang &amp; Dietrich &amp; Zhang, 2018" authorityName="Wang &amp; Dietrich &amp; Zhang" authorityYear="2018" box="[151,392,475,501]" class="Insecta" family="Cicadellidae" genus="Riseveinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="20C4EAB5F20F2438D3BC5CDDB3AAFE5F" bold="true" box="[151,392,475,501]" italics="true" pageId="2" pageNumber="261">Riseveinus hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3BF757CEF20F2438D2BB5CDAB3CFFE5C" box="[400,493,476,502]" pageId="2" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5AAA652CF20F2438D3BC5CF8B2CFFCB9" pageId="2" pageNumber="261" type="description">
<paragraph id="120F36A7F20F2438D3BC5CF8B307FDBD" blockId="2.[151,493,475,535]" box="[151,293,510,535]" pageId="2" pageNumber="261">
(
<figureCitation id="8A8B2A22F20F2438D3B45CF8B33CFDBD" box="[159,286,510,535]" captionStart="FIGURES 1117" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[191,1392,198,1915]" captionTargetId="figure@4.[185,1405,193,1920]" captionTargetPageId="4" captionText="FIGURES 1117. Riseveinus hamatus sp. nov.: 11. male habitus, dorsal view; 12. male habitus, lateral view; 13. male face; 14. male pygofer side, lateral view; 15. aedeagal, connective and style, ventral view; 16. aedeagal, lateral view; 17. subgenital plate, ventral view." httpUri="https://zenodo.org/record/2607076/files/figure.png" pageId="2" pageNumber="261">Figs 1117</figureCitation>
)
</paragraph>
<paragraph id="120F36A7F20F2438D3BC5F40B628FD29" blockId="2.[151,1437,582,1111]" pageId="2" pageNumber="261">
<emphasis id="20C4EAB5F20F2438D3BC5F40B30BFDF5" bold="true" box="[151,297,582,607]" pageId="2" pageNumber="261">Description.</emphasis>
Body length (incl. forewing):
<specimenCount id="04B6FD2EF20F2438D1555F41B0B7FDF4" box="[638,661,583,606]" pageId="2" pageNumber="261" type="male"></specimenCount>
,
<quantity id="D5489B42F20F2438D18A5F40B104FDF4" box="[673,806,582,606]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" metricValueMax="8.5" metricValueMin="8.3" pageId="2" pageNumber="261" unit="mm" value="8.4" valueMax="8.5" valueMin="8.3">8.3-8.5 mm</quantity>
. Head width (incl. eyes):
<quantity id="D5489B42F20F2438D76E5F40B6EDFDF4" box="[1093,1231,582,606]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="2" pageNumber="261" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.51.6 mm</quantity>
; head length:
<quantity id="D5489B42F20F2438D6405F40B2D2FD28" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="2" pageNumber="261" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5 1.6 mm</quantity>
. Crown (
<figureCitation id="8A8B2A22F20F2438D27D5F6CB387FD29" box="[342,421,618,643]" captionStart="FIGURES 1117" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[191,1392,198,1915]" captionTargetId="figure@4.[185,1405,193,1920]" captionTargetPageId="4" captionText="FIGURES 1117. Riseveinus hamatus sp. nov.: 11. male habitus, dorsal view; 12. male habitus, lateral view; 13. male face; 14. male pygofer side, lateral view; 15. aedeagal, connective and style, ventral view; 16. aedeagal, lateral view; 17. subgenital plate, ventral view." httpUri="https://zenodo.org/record/2607076/files/figure.png" pageId="2" pageNumber="261">Fig. 11</figureCitation>
) about 3/4 length of pronotum and scutellum together.
</paragraph>
<paragraph id="120F36A7F20F2438D3EC5F88B2CFFCB9" blockId="2.[151,1437,582,1111]" pageId="2" pageNumber="261">
Anal tube with barblike ventral process (
<figureCitation id="8A8B2A22F20F2438D18B5F88B0D5FD0D" box="[672,759,654,679]" captionStart="FIGURES 1117" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[191,1392,198,1915]" captionTargetId="figure@4.[185,1405,193,1920]" captionTargetPageId="4" captionText="FIGURES 1117. Riseveinus hamatus sp. nov.: 11. male habitus, dorsal view; 12. male habitus, lateral view; 13. male face; 14. male pygofer side, lateral view; 15. aedeagal, connective and style, ventral view; 16. aedeagal, lateral view; 17. subgenital plate, ventral view." httpUri="https://zenodo.org/record/2607076/files/figure.png" pageId="2" pageNumber="261">Fig. 14</figureCitation>
). Aedeagal shaft short, swollen at base and then gradually narrowing towards apex in lateral view, without apical processes; apodeme bilobed, dorsolateral extensions short, rounded; paired ventrolateral processes longer than aedeagal shaft, and close to aedeagal shaft in lateral view (
<figureCitation id="8A8B2A22F20F2438D6415FD0B2C2FCB9" captionStart="FIGURES 1117" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[191,1392,198,1915]" captionTargetId="figure@4.[185,1405,193,1920]" captionTargetPageId="4" captionText="FIGURES 1117. Riseveinus hamatus sp. nov.: 11. male habitus, dorsal view; 12. male habitus, lateral view; 13. male face; 14. male pygofer side, lateral view; 15. aedeagal, connective and style, ventral view; 16. aedeagal, lateral view; 17. subgenital plate, ventral view." httpUri="https://zenodo.org/record/2607076/files/figure.png" pageId="2" pageNumber="261">Figs 15, 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5AAA652CF20F2438D3EC5E18B719FCD5" pageId="2" pageNumber="261" type="materials_examined">
<paragraph id="120F36A7F20F2438D3EC5E18B719FCD5" blockId="2.[151,1437,582,1111]" pageId="2" pageNumber="261">
<emphasis id="20C4EAB5F20F2438D3EC5E18B357FC9D" bold="true" box="[199,373,798,823]" pageId="2" pageNumber="261">Type material.</emphasis>
<materialsCitation id="A2D83CFAF20F2438D2565E18B131FCF1" ID-GBIF-Occurrence="2028138920" collectingDate="2004-06-18" collectionCode="NMNS" collectorName="C. H. Dietrich" country="Taiwan" latitude="24.251667" location="Dashuishan Forest Rd." longLatPrecision="21" longitude="120.916115" pageId="2" pageNumber="261" specimenCount="1" specimenCount-male="1" stateProvince="Taichung" typeStatus="holotype">
<typeStatus id="CD0B8805F20F2438D2565E18B3C7FC9D" box="[381,485,798,823]" pageId="2" pageNumber="261" type="holotype">Holotype</typeStatus>
<specimenCount id="04B6FD2EF20F2438D2C65E19B020FC9C" box="[493,514,799,822]" pageId="2" pageNumber="261" type="male"></specimenCount>
(
<collectionCode id="74A1AE62F20F2438D13A5E19B045FC9D" box="[529,615,799,823]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="2" pageNumber="261">NMNS</collectionCode>
),
<collectingCountry id="6AA77637F20F2438D1505E19B0C8FC9D" box="[635,746,799,823]" name="Taiwan" pageId="2" pageNumber="261">TAIWAN</collectingCountry>
,
<collectingRegion id="D074F845F20F2438D1DD5E18B143FC9D" box="[758,865,798,823]" country="Taiwan" name="Taichung" pageId="2" pageNumber="261">Taichung</collectingRegion>
,
<location id="176F607CF20F2438D0465E18B64FFC9D" LSID="urn:lsid:plazi:treatment:9A1987B1F20F2438D3BC5CDDB1F4FBFD:176F607CF20F2438D0465E18B64FFC9D" box="[877,1133,798,823]" country="Taiwan" latitude="24.251667" longLatPrecision="21" longitude="120.916115" name="Dashuishan Forest Rd." pageId="2" pageNumber="261" stateProvince="Taichung">Dashuishan Forest Rd.</location>
km 20, 1250m,
<geoCoordinate id="77845060F20F2438D60D5E18B7BEFC9D" box="[1318,1436,798,823]" direction="north" orientation="latitude" pageId="2" pageNumber="261" precision="15" value="24.251667">24°15'6&quot;N</geoCoordinate>
<geoCoordinate id="77845060F20F2438D3BC5E44B30AFCF1" box="[151,296,834,859]" direction="east" orientation="longitude" pageId="2" pageNumber="261" precision="15" value="120.916115">120°54'58&quot;E</geoCoordinate>
,
<date id="660E1067F20F2438D21E5E44B38DFCF1" box="[309,431,834,859]" pageId="2" pageNumber="261" value="2004-06-18">
<collectingDate id="764AE98FF20F2438D21E5E44B38DFCF1" box="[309,431,834,859]" pageId="2" pageNumber="261" value="2004-06-18">18.vi.2004</collectingDate>
</date>
,
<collectorName id="BF455371F20F2438D2975E45B07AFCF1" box="[444,600,834,859]" pageId="2" pageNumber="261">C.H. Dietrich</collectorName>
, 16-1, sweeping
</materialsCitation>
.
<materialsCitation id="A2D83CFAF20F2438D0345E45B715FCD5" ID-GBIF-Occurrence="2028138923" collectingDate="2004-06-18" collectionCode="NWAFU, INHS" collectorName="J. H. Zahniser" country="Taiwan" elevation="1600" latitude="24.246668" location="Dashuishan Forest Rd." longLatPrecision="21" longitude="120.935555" pageId="2" pageNumber="261" specimenCount="2" specimenCount-male="2" stateProvince="Taichung" typeStatus="paratype">
<typeStatus id="CD0B8805F20F2438D0345E45B1B2FCF1" box="[799,912,835,859]" pageId="2" pageNumber="261" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="04B6FD2EF20F2438D0B75E45B1F6FCF1" box="[924,980,835,859]" pageId="2" pageNumber="261" type="male">2♂♂</specimenCount>
(
<collectionCode id="74A1AE62F20F2438D0C85E45B66EFCF1" box="[995,1100,835,859]" pageId="2" pageNumber="261">NWAFU</collectionCode>
,
<collectionCode id="74A1AE62F20F2438D7735E45B6BFFCF1" box="[1112,1181,835,859]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="2" pageNumber="261" type="Other type of research institution/biorepository">INHS</collectionCode>
),
<collectingCountry id="6AA77637F20F2438D7995E45B703FCF1" box="[1202,1313,835,859]" name="Taiwan" pageId="2" pageNumber="261">TAIWAN</collectingCountry>
,
<collectingRegion id="D074F845F20F2438D6065E44B7BAFCF1" box="[1325,1432,834,859]" country="Taiwan" name="Taichung" pageId="2" pageNumber="261">Taichung</collectingRegion>
,
<location id="176F607CF20F2438D3BC5E60B3B4FCD5" LSID="urn:lsid:plazi:treatment:9A1987B1F20F2438D3BC5CDDB1F4FBFD:176F607CF20F2438D3BC5E60B3B4FCD5" box="[151,406,870,895]" country="Taiwan" latitude="24.246668" longLatPrecision="21" longitude="120.935555" name="Dashuishan Forest Rd." pageId="2" pageNumber="261" stateProvince="Taichung">Dashuishan Forest Rd.</location>
km 23,
<quantity id="D5489B42F20F2438D2DA5E60B063FCD5" box="[497,577,870,895]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="261" unit="m" value="1600.0">
<elevation id="999DD194F20F2438D2DA5E60B063FCD5" box="[497,577,870,895]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="261" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<geoCoordinate id="77845060F20F2438D1665E61B0F2FCD5" box="[589,720,871,895]" direction="north" orientation="latitude" pageId="2" pageNumber="261" precision="15" value="24.246668">24°14'48&quot;N</geoCoordinate>
<geoCoordinate id="77845060F20F2438D1F35E60B17AFCD5" box="[728,856,870,895]" direction="east" orientation="longitude" pageId="2" pageNumber="261" precision="15" value="120.935555">120°56'8&quot;E</geoCoordinate>
<date id="660E1067F20F2438D04B5E60B1F8FCD5" box="[864,986,870,895]" pageId="2" pageNumber="261" value="2004-06-18">
<collectingDate id="764AE98FF20F2438D04B5E60B1F8FCD5" box="[864,986,870,895]" pageId="2" pageNumber="261" value="2004-06-18">18.vi.2004</collectingDate>
</date>
,
<collectorName id="BF455371F20F2438D0CE5E61B65FFCD5" box="[997,1149,870,895]" pageId="2" pageNumber="261">J.H. Zahniser</collectorName>
, 17-1, sweeping
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5AAA652CF20F2438D3EC5E8CB315FC6D" pageId="2" pageNumber="261" type="etymology">
<paragraph id="120F36A7F20F2438D3EC5E8CB315FC6D" blockId="2.[151,1437,582,1111]" pageId="2" pageNumber="261">
<emphasis id="20C4EAB5F20F2438D3EC5E8CB36CFC09" bold="true" box="[199,334,906,931]" pageId="2" pageNumber="261">Etymology.</emphasis>
The specific epithet “
<emphasis id="20C4EAB5F20F2438D1635E8DB0BEFC09" box="[584,668,907,931]" italics="true" pageId="2" pageNumber="261">hamata</emphasis>
”, meaning hooked, refers to the barblike ventral process of the anal tube (
<figureCitation id="8A8B2A22F20F2438D3FD5EA8B308FC6D" box="[214,298,942,967]" captionStart="FIGURES 1117" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[191,1392,198,1915]" captionTargetId="figure@4.[185,1405,193,1920]" captionTargetPageId="4" captionText="FIGURES 1117. Riseveinus hamatus sp. nov.: 11. male habitus, dorsal view; 12. male habitus, lateral view; 13. male face; 14. male pygofer side, lateral view; 15. aedeagal, connective and style, ventral view; 16. aedeagal, lateral view; 17. subgenital plate, ventral view." httpUri="https://zenodo.org/record/2607076/files/figure.png" pageId="2" pageNumber="261">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5AAA652CF20F2438D3EC5ED4B1F4FBFD" pageId="2" pageNumber="261" type="discussion">
<paragraph id="120F36A7F20F2438D3EC5ED4B1F4FBFD" blockId="2.[151,1437,582,1111]" pageId="2" pageNumber="261">
<emphasis id="20C4EAB5F20F2438D3EC5ED4B319FC41" bold="true" box="[199,315,978,1003]" pageId="2" pageNumber="261">Remarks.</emphasis>
This species is very similar to
<taxonomicName id="D5B04D24F20F2438D18C5ED5B1A4FC41" authority="(Jacobi)" authorityName="Jacobi" baseAuthorityName="Jacobi" box="[679,902,978,1003]" class="Insecta" family="Cicadellidae" genus="Dussana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="20C4EAB5F20F2438D18C5ED5B100FC40" box="[679,802,979,1002]" italics="true" pageId="2" pageNumber="261">R. sinensis</emphasis>
(Jacobi)
</taxonomicName>
externally, but differs as follows: 1) anal tube with barblike ventral process (
<figureCitation id="8A8B2A22F20F2438D2C05EF0B01CFBA5" box="[491,574,1014,1039]" captionStart="FIGURES 1117" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[191,1392,198,1915]" captionTargetId="figure@4.[185,1405,193,1920]" captionTargetPageId="4" captionText="FIGURES 1117. Riseveinus hamatus sp. nov.: 11. male habitus, dorsal view; 12. male habitus, lateral view; 13. male face; 14. male pygofer side, lateral view; 15. aedeagal, connective and style, ventral view; 16. aedeagal, lateral view; 17. subgenital plate, ventral view." httpUri="https://zenodo.org/record/2607076/files/figure.png" pageId="2" pageNumber="261">Fig. 14</figureCitation>
); absent in
<taxonomicName id="D5B04D24F20F2438D1945EF1B115FBA4" box="[703,823,1015,1038]" class="Insecta" family="Cicadellidae" genus="Dussana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="20C4EAB5F20F2438D1945EF1B115FBA4" box="[703,823,1015,1038]" italics="true" pageId="2" pageNumber="261">R. sinensis</emphasis>
</taxonomicName>
); 2) in lateral view, aedeagal shaft swollen at base and then narrowing towards apex gradually (aedeagal shaft same width from base to apex in
<taxonomicName id="D5B04D24F20F2438D7A3591DB723FB98" box="[1160,1281,1051,1074]" class="Insecta" family="Cicadellidae" genus="Dussana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="20C4EAB5F20F2438D7A3591DB723FB98" box="[1160,1281,1051,1074]" italics="true" pageId="2" pageNumber="261">R. sinensis</emphasis>
</taxonomicName>
); 3) aedeagal ventral process evenly curved near apex (sinuate near apex in
<taxonomicName id="D5B04D24F20F2438D0655939B1E4FBFC" box="[846,966,1087,1110]" class="Insecta" family="Cicadellidae" genus="Dussana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="20C4EAB5F20F2438D0655939B1E4FBFC" box="[846,966,1087,1110]" italics="true" pageId="2" pageNumber="261">R. sinensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>