treatments-xml/data/47/A1/37/47A137B854F75944B6ECCCB9896B43AB.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne &amp; Haring, Elisabeth" docDate="2021" docId="47A137B854F75944B6ECCCB9896B43AB" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Charpentieria itala subsp. zalloti Mattia, Reier &amp; Haring, 2021, nom. nov." docType="treatment" docVersion="4" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1668151235153" updateUser="ExternalLinkService">
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<mods:title>Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>De Mattia, Willy</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Reier, Susanne</mods:namePart>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Haring, Elisabeth</mods:namePart>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:date>2021</mods:date>
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<mods:number>2021-12-14</mods:number>
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<treatment ID-GBIF-Taxon="191645954" LSID="urn:lsid:plazi:treatment:47A137B854F75944B6ECCCB9896B43AB" httpUri="http://treatment.plazi.org/id/47A137B854F75944B6ECCCB9896B43AB" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="47A137B8-54F7-5944-B6EC-CCB9896B43AB" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. zalloti" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" status="nom. nov." subSpecies="zalloti">Charpentieria itala zalloti</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">nom. nov.</taxonomicNameLabel>
</paragraph>
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<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Charpentieria itala zalloti nom. nov., Udine 48.1 whole distal genital organs 48.2 internal distal part of genital organs. Charpentieria itala serravalensis (Nordsieck, 1963), Serravalle, Vittorio Veneto 48.3 whole distal genital organs 48.4 internal distal part of genital organs. Charpentieria itala trepida (Kaeufel, 1928), Cima Ora, Anfo 48.5 whole distal genital organs 48.6 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure48" httpUri="https://binary.pensoft.net/fig/622840" pageId="0" pageNumber="1">Figs 48.1-48.2, 50.1-50.2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
During field collecting (WDM), five populations belonging to 
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
</taxonomicName>
ssp. were found in the surrounding of Udine and Cividale (Friuli, Italy). These populations show a remarkably reduced size of the shell comparing to the nominate subspecies or 
<taxonomicName lsidName="C. itala subsp. serravalensis" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="serravalensis">
<emphasis italics="true" pageId="0" pageNumber="1">C. itala serravalensis</emphasis>
</taxonomicName>
, that are the closest 
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
</taxonomicName>
subspecies in the west (Figs 
<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Charpentieria itala itala (G. von Martens, 1824), Barbarano Vicentino 44.1, 44.2 whole distal genital organs 44.3 internal distal part of genital organs 44.4 detail of the pseudopapilla. Charpentieria itala albopustolata (De Cristofori &amp; Jan, 1832), Brescia 44.5 whole distal genital organs 44.6 internal distal part of genital organs. Charpentieria itala allatollae (Kaeufel, 1928), Val Ampola, Storo 44.7 whole distal genital organs 44.8 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure44" httpUri="https://binary.pensoft.net/fig/622836" pageId="0" pageNumber="1">44</figureCitation>
, 
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Charpentieria itala zalloti nom. nov., Udine 48.1 whole distal genital organs 48.2 internal distal part of genital organs. Charpentieria itala serravalensis (Nordsieck, 1963), Serravalle, Vittorio Veneto 48.3 whole distal genital organs 48.4 internal distal part of genital organs. Charpentieria itala trepida (Kaeufel, 1928), Cima Ora, Anfo 48.5 whole distal genital organs 48.6 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure48" httpUri="https://binary.pensoft.net/fig/622840" pageId="0" pageNumber="1">48</figureCitation>
). Moreover, the sutures are almost colourless, the shell is translucent and the peristome is continuous (Figs 
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Charpentieria itala zalloti nom. nov., Udine 50.1 - 50.2 shell. Charpentieria itala serravalensis (Nordsieck, 1963), Serravalle, Vittorio Veneto 50.3 shell. Charpentieria itala trepida (Kaeufel, 1928), Cima Ora, Anfo 50.4 shell. Charpentieria itala triumplinae Nardi, 2011, Lavenone, Brescia 50.5 shell. Charpentieria itala variscoi (Pini, 1883), Val Taleggio, Bergamo 50.6 shell." figureDoi="10.3897/zookeys.1077.67081.figure50" httpUri="https://binary.pensoft.net/fig/622842" pageId="0" pageNumber="1">50</figureCitation>
.1, 50.2). 
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="169 - 203" refId="B58" refString="Nordsieck, H, 1963b. Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen. Archiv fuer Molluskenkunde 92: 169 - 203" title="Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen." volume="92" year="1963 b">Nordsieck (1963b)</bibRefCitation>
stated that the eastern distributional limit for 
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
</taxonomicName>
is represented by the axis Pordenone-Ampezzo. Thus, these Friulian populations were neglected or simply not known by him. Nevertheless, these populations of 
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
</taxonomicName>
were not unknown to past authors. 
<bibRefCitation author="Pirona, A" journalOrPublisher="Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti" pageId="0" pageNumber="1" pagination="675 - 708" refId="B79" refString="Pirona, A, 1865. Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti 10: 675 - 708" title="Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli." volume="10" year="1865">Pirona (1865</bibRefCitation>
: 692) cited &quot; 
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia alboguttata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alboguttata">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia alboguttata</emphasis>
</taxonomicName>
Wagn. [= 
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
</taxonomicName>
]. Nei muri vecchi muscosi (On old mossy walls); Udine, Tricesimo, Dolegna, Sacile, Portogruaro&quot;. He listed two morphotypes: var. a, 
<emphasis italics="true" pageId="0" pageNumber="1">major</emphasis>
for the western part (Caneva and Portogruaro) and var. b, 
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia minor" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia minor</emphasis>
</taxonomicName>
for Udine and Dolegna. In his description, for var. b, 
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia minor" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia minor</emphasis>
</taxonomicName>
<bibRefCitation author="Pirona, A" journalOrPublisher="Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti" pageId="0" pageNumber="1" pagination="675 - 708" refId="B79" refString="Pirona, A, 1865. Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti 10: 675 - 708" title="Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli." volume="10" year="1865">Pirona (1865)</bibRefCitation>
pointed out the translucent shell (
<emphasis italics="true" pageId="0" pageNumber="1">cornea</emphasis>
) and the continuous peristome (
<emphasis italics="true" pageId="0" pageNumber="1">peristomate continuo</emphasis>
) and these details, as previously stated, are regularly found also in the easternmost populations recently collected. Concerning the anatomy of the genital organs, these populations present the absence of the (2) longitudinal epiphallar pleats (Figs 
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Charpentieria itala zalloti nom. nov., Udine 48.1 whole distal genital organs 48.2 internal distal part of genital organs. Charpentieria itala serravalensis (Nordsieck, 1963), Serravalle, Vittorio Veneto 48.3 whole distal genital organs 48.4 internal distal part of genital organs. Charpentieria itala trepida (Kaeufel, 1928), Cima Ora, Anfo 48.5 whole distal genital organs 48.6 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure48" httpUri="https://binary.pensoft.net/fig/622840" pageId="0" pageNumber="1">48</figureCitation>
.1-48.2). As regards the identity of the var. a, 
<emphasis italics="true" pageId="0" pageNumber="1">major</emphasis>
, following 
<normalizedToken originalValue="Pirona’s">Pirona's</normalizedToken>
description (
<emphasis italics="true" pageId="0" pageNumber="1">
saturate 
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria rubiginea" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rubiginea">Charpentieria rubiginea</taxonomicName>
</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="1">peristomate soluto</emphasis>
) it can be postulated that he dealt with the nominate subspecies.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The type locality of the nominate subspecies is less than 50 km west of Portogruaro. Thus, following 
<bibRefCitation author="Pirona, A" journalOrPublisher="Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti" pageId="0" pageNumber="1" pagination="675 - 708" refId="B79" refString="Pirona, A, 1865. Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti 10: 675 - 708" title="Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli." volume="10" year="1865">Pirona (1865)</bibRefCitation>
and recent research, the distributional area of 
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
</taxonomicName>
is further extended eastward as far as the Friuli plain and Cividale/Dolegna in the east.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Despite 
<bibRefCitation author="Pirona, A" journalOrPublisher="Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti" pageId="0" pageNumber="1" pagination="675 - 708" refId="B79" refString="Pirona, A, 1865. Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti 10: 675 - 708" title="Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli." volume="10" year="1865">
<normalizedToken originalValue="Pirona’s">Pirona's</normalizedToken>
(1865)
</bibRefCitation>
thorough description, 
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia minor" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia minor</emphasis>
</taxonomicName>
Pirona, 1865 is several times preoccupied by: 
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia minor" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia minor</emphasis>
</taxonomicName>
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
, 1850, currently a synonym of 
<taxonomicName authority="contracta" class="Gastropoda" family="Clausiliidae" genus="Medora" kingdom="Animalia" lsidName="Medora contracta subsp. contracta" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="contracta" subSpecies="contracta">
<emphasis italics="true" pageId="0" pageNumber="1">Medora contracta contracta</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
, 1842); 
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia minor" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia minor</emphasis>
</taxonomicName>
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
, 1850, currently a synonym of 
<taxonomicName authority="almissana" class="Gastropoda" family="Clausiliidae" genus="Medora" kingdom="Animalia" lsidName="Medora almissana subsp. almissana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="almissana" subSpecies="almissana">
<emphasis italics="true" pageId="0" pageNumber="1">Medora almissana almissana</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
, 1847); 
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia minor" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia minor</emphasis>
</taxonomicName>
Charpentier, 1852, currently a synonym of 
<taxonomicName baseAuthorityName="Charpentier" baseAuthorityYear="1852" class="Gastropoda" family="Clausiliidae" genus="Albinaria" kingdom="Animalia" lsidName="Albinaria profuga" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="profuga">
<emphasis italics="true" pageId="0" pageNumber="1">Albinaria profuga</emphasis>
</taxonomicName>
(Charpentier, 1852); 
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia minor" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia minor</emphasis>
</taxonomicName>
A. Schmidt, 1856, currently a synonym of 
<taxonomicName authorityName="Coutagne" authorityYear="1886" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia rugosa subsp. provincialis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="rugosa" subSpecies="provincialis">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia rugosa provincialis</emphasis>
</taxonomicName>
Coutagne, 1886; 
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia minor" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia minor</emphasis>
</taxonomicName>
Walderdorff, 1864, currently a synonym of 
<taxonomicName baseAuthorityName="Walderdorff" baseAuthorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Delima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Delima montenegrina subsp. semilabiata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="montenegrina" subSpecies="semilabiata">
<emphasis italics="true" pageId="0" pageNumber="1">Delima montenegrina semilabiata</emphasis>
</taxonomicName>
(Walderdorff, 1864).
</paragraph>
<paragraph pageId="0" pageNumber="1">
The Friulian populations with translucent shell and reduced shell dimensions are here named 
<taxonomicName lsidName="C. itala subsp. zalloti" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="zalloti">
<emphasis italics="true" pageId="0" pageNumber="1">C. itala zalloti</emphasis>
</taxonomicName>
nom. nov.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="https://hnords.de/wordpress/charpentieria-itala/" refId="B67" refString="Nordsieck, H, 2011. Charpentieria itala: subspecies, species, superspecies. https://hnords.de/wordpress/charpentieria-itala/" title="Charpentieria itala: subspecies, species, superspecies." url="https://hnords.de/wordpress/charpentieria-itala/" year="2011">Nordsieck (2011</bibRefCitation>
, supplemented May 2021) in chapter 4 clearly referred to our data and manuscript when he felt the need to supplement his web article. He cited 
<bibRefCitation author="Pirona, A" journalOrPublisher="Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti" pageId="0" pageNumber="1" pagination="675 - 708" refId="B79" refString="Pirona, A, 1865. Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti 10: 675 - 708" title="Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli." volume="10" year="1865">Pirona (1865)</bibRefCitation>
stating that &quot;The form from Udine, Friuli (
<normalizedToken originalValue="…">...</normalizedToken>
) is strikingly similar to that from Verona. (
<normalizedToken originalValue="…">...</normalizedToken>
) Because of this similarity it is deemed to have been introduced.&quot; We disagree with his rushed conclusion, as the presence of this taxon also in many natural spots along the low Prealps makes an introduction quite unlikely or, at least, deserving additional research.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. zalloti" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="zalloti">
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala zalloti</emphasis>
</taxonomicName>
nom. nov. is found on limestone stonewalls (Cividale) or natural limestone small cliffs (Udine castle hill, Tricesimo and Julian Prealps), usually in shady spots. The taxon is found in the Friuli plain and in natural habitats along the low Prealps. 
<bibRefCitation author="Pirona, A" journalOrPublisher="Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti" pageId="0" pageNumber="1" pagination="675 - 708" refId="B79" refString="Pirona, A, 1865. Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti 10: 675 - 708" title="Prospetto dei molluschi terrestri e fluviatili finora raccolti nel Friuli." volume="10" year="1865">Pirona (1865</bibRefCitation>
: 692) provides a wider distribution including territories in the west, as far as Sacile, and in the east as far as Dolegna.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="L. Anzil" country="Italy" determinerName="W. De Mattia" elevation="140" latitude="46.064823" location="Udine" longLatPrecision="1" longitude="13.236764" specimenCount="1" stateProvince="Friuli-Venezia Giulia">
<collectingCountry name="Italy">Italy</collectingCountry>
, 
<collectingRegion country="Italy" name="Friuli-Venezia Giulia">Friuli-Venezia Giulia</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:47A137B854F75944B6ECCCB9896B43AB:B7F73F29941CB8ADE03A49D598335074" country="Italy" latitude="46.064823" longLatPrecision="1" longitude="13.236764" name="Udine" stateProvince="Friuli-Venezia Giulia">Udine</location>
, castle hill. 
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.4" unit="m" value="140.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.4" unit="m" value="140.0">140 m</quantity>
asl
</elevation>
, 
<geoCoordinate degrees="46" direction="north" minutes="03" orientation="latitude" precision="1" seconds="53.37" value="46.064823">46°03'53.37&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="13" direction="east" minutes="14" orientation="longitude" precision="1" seconds="12.35" value="13.236764">13°14'12.35&quot;E</geoCoordinate>
, 
<collectorName>L. Anzil</collectorName>
leg., 
<determinerName>W. De Mattia</determinerName>
det., 3 dissected spm
</materialsCitation>
. 
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="W. De Mattia, J. Macor" country="Italy" elevation="600" latitude="46.14071" location="Tribil Superiore." longLatPrecision="1" longitude="13.617561" municipality="Udine" specimenCount="1" stateProvince="Friuli-Venezia Giulia">
<collectingCountry name="Italy">Italy</collectingCountry>
, 
<collectingRegion country="Italy" name="Friuli-Venezia Giulia">Friuli-Venezia Giulia</collectingRegion>
, 
<collectingMunicipality>Udine</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:47A137B854F75944B6ECCCB9896B43AB:FA4A93948CF9A64EFE95B2B33FB07B57" country="Italy" latitude="46.14071" longLatPrecision="1" longitude="13.617561" municipality="Udine" name="Tribil Superiore." stateProvince="Friuli-Venezia Giulia">Tribil Superiore.</location>
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
asl
</elevation>
, 
<geoCoordinate degrees="46" direction="north" minutes="08" orientation="latitude" precision="1" seconds="26.55" value="46.14071">46°08'26.55&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="13" direction="east" minutes="37" orientation="longitude" precision="1" seconds="3.22" value="13.617561">13°37'3.22&quot;E</geoCoordinate>
, 
<collectorName>W. De Mattia</collectorName>
and 
<collectorName>J. Macor</collectorName>
leg. and det., 2 dissected spm
</materialsCitation>
. 
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="W. De Mattia, J. Macor" country="Italy" elevation="340" latitude="46.132034" location="Lesizza." longLatPrecision="1" longitude="13.565833" municipality="Udine" specimenCount="1" stateProvince="Friuli-Venezia Giulia">
<collectingCountry name="Italy">Italy</collectingCountry>
, 
<collectingRegion country="Italy" name="Friuli-Venezia Giulia">Friuli-Venezia Giulia</collectingRegion>
, 
<collectingMunicipality>Udine</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:47A137B854F75944B6ECCCB9896B43AB:DF5B7333BFCEE6BF45A50BD159239547" country="Italy" latitude="46.132034" longLatPrecision="1" longitude="13.565833" municipality="Udine" name="Lesizza." stateProvince="Friuli-Venezia Giulia">Lesizza.</location>
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.4" unit="m" value="340.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.4" unit="m" value="340.0">340 m</quantity>
asl
</elevation>
, 
<geoCoordinate degrees="46" direction="north" minutes="7" orientation="latitude" precision="1" seconds="55.33" value="46.132034">46°7'55.33&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="13" direction="east" minutes="33" orientation="longitude" precision="1" seconds="57.00" value="13.565833">13°33'57.00&quot;E</geoCoordinate>
, 
<collectorName>W. De Mattia</collectorName>
and 
<collectorName>J. Macor</collectorName>
leg. and det
</materialsCitation>
. 
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="W. De Mattia, J. Macor" country="Italy" elevation="730" latitude="46.187527" location="Stermizza." longLatPrecision="1" longitude="13.525355" municipality="Udine" specimenCount="1" stateProvince="Friuli-Venezia Giulia">
<collectingCountry name="Italy">Italy</collectingCountry>
, 
<collectingRegion country="Italy" name="Friuli-Venezia Giulia">Friuli-Venezia Giulia</collectingRegion>
, 
<collectingMunicipality>Udine</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:47A137B854F75944B6ECCCB9896B43AB:3639332AC7E17C8875C37111783B6EEE" country="Italy" latitude="46.187527" longLatPrecision="1" longitude="13.525355" municipality="Udine" name="Stermizza." stateProvince="Friuli-Venezia Giulia">Stermizza.</location>
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">730 m</quantity>
asl
</elevation>
, 
<geoCoordinate degrees="46" direction="north" minutes="11" orientation="latitude" precision="1" seconds="15.09" value="46.187527">46°11'15.09&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="13" direction="east" minutes="31" orientation="longitude" precision="1" seconds="31.28" value="13.525355">13°31'31.28&quot;E</geoCoordinate>
, 
<collectorName>W. De Mattia</collectorName>
and 
<collectorName>J. Macor</collectorName>
leg. and det
</materialsCitation>
. 
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="W. De Mattia, J. Macor" country="Italy" elevation="670" latitude="46.153507" location="Rucchin." longLatPrecision="1" longitude="13.63673" municipality="Udine" specimenCount="1" stateProvince="Friuli-Venezia Giulia">
<collectingCountry name="Italy">Italy</collectingCountry>
, 
<collectingRegion country="Italy" name="Friuli-Venezia Giulia">Friuli-Venezia Giulia</collectingRegion>
, 
<collectingMunicipality>Udine</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:47A137B854F75944B6ECCCB9896B43AB:DAF77EA35012B33DE962232FBAB833D5" country="Italy" latitude="46.153507" longLatPrecision="1" longitude="13.63673" municipality="Udine" name="Rucchin." stateProvince="Friuli-Venezia Giulia">Rucchin.</location>
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">670 m</quantity>
asl
</elevation>
, 
<geoCoordinate degrees="46" direction="north" minutes="9" orientation="latitude" precision="1" seconds="12.62" value="46.153507">46°9'12.62&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="13" direction="east" minutes="38" orientation="longitude" precision="1" seconds="12.23" value="13.63673">13°38'12.23&quot;E</geoCoordinate>
, 
<collectorName>W. De Mattia</collectorName>
and 
<collectorName>J. Macor</collectorName>
leg. and det
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="external morphology of the genital organs">
<paragraph pageId="0" pageNumber="1">External morphology of the genital organs</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Fig. 
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Charpentieria itala zalloti nom. nov., Udine 48.1 whole distal genital organs 48.2 internal distal part of genital organs. Charpentieria itala serravalensis (Nordsieck, 1963), Serravalle, Vittorio Veneto 48.3 whole distal genital organs 48.4 internal distal part of genital organs. Charpentieria itala trepida (Kaeufel, 1928), Cima Ora, Anfo 48.5 whole distal genital organs 48.6 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure48" httpUri="https://binary.pensoft.net/fig/622840" pageId="0" pageNumber="1">48</figureCitation>
.1).
</emphasis>
The V is slightly shorter than the FO. The FDBC is as long as the SDBC+BC. The BC+SDBC is club-like, with a more or less clear distinction between the SDBC and the BC. The D is slightly shorter than the BC+SDBC. The V is cylindrical and wide in diameter. The PC is only slightly longer than the V. The P is short but wide in diameter and the transition between P and EP is clearly visible. The PR is short and robust. The E is almost as long as the P but small in diameter.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="internal morphology of the genital organs">
<paragraph pageId="0" pageNumber="1">Internal morphology of the genital organs</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Fig. 
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Charpentieria itala zalloti nom. nov., Udine 48.1 whole distal genital organs 48.2 internal distal part of genital organs. Charpentieria itala serravalensis (Nordsieck, 1963), Serravalle, Vittorio Veneto 48.3 whole distal genital organs 48.4 internal distal part of genital organs. Charpentieria itala trepida (Kaeufel, 1928), Cima Ora, Anfo 48.5 whole distal genital organs 48.6 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure48" httpUri="https://binary.pensoft.net/fig/622840" pageId="0" pageNumber="1">48</figureCitation>
.2).
</emphasis>
The V is smooth. The P presents two strong irregular longitudinal folded pleats that reach the A. The PP is folded and elongated with a pointed tip. It occupies almost the whole penial volume. The PP originates from the ER which is connected to the ELP. The two ELP are weak. The epiphallar formula is: PP(ER(ELP)).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. zalloti" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="zalloti">
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala zalloti</emphasis>
</taxonomicName>
nom. nov. was named after Enrico Zallot (Feltre, Italy-Monster, The Netherlands), skilled malacologist, as a token of friendship and esteem.
</paragraph>
</subSubSection>
</treatment>
</document>