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<mods:title id="E4C1FA509DABE4E9CA35CF28622F04AA">A New Species of Forstercooperia (Perissodactyla: Paraceratheriidae) from Northern China with a Systematic Revision of Forstercooperiines</mods:title>
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<heading id="ED0581FCFFF12916A1CD1F84FC9ED519" box="[497,823,1532,1558]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="5">
<taxonomicName id="71F24D13FFF12916A1CD1F84FC9ED519" authority="Wood, 1938" authorityName="Wood" authorityYear="1938" box="[497,823,1532,1558]" class="Mammalia" family="Hyracodontidae" genus="Forstercooperia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
<emphasis id="8486EA82FFF12916A1CD1F84FD08D519" box="[497,673,1533,1558]" italics="true" pageId="3" pageNumber="4">Forstercooperia</emphasis>
<bibRefCitation id="D2634B61FFF12916A2951F85FC9ED519" author="Wood, H. E." box="[681,823,1532,1558]" pageId="3" pageNumber="4" pagination="1 - 20" refId="ref20185" refString="Wood, H. E. 1938. Cooperia totadentata, a remarkable rhinoceros from the Eocene of Mongolia. American Museum Novitates 1012: 1 - 20." type="journal article" year="1938">Wood, 1938</bibRefCitation>
</taxonomicName>
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<typeStatus id="69498832FFF12916A0A01C4CFF7ED540" box="[156,215,1589,1615]" pageId="3" pageNumber="4">TYPE</typeStatus>
SPECIES:
<taxonomicName id="71F24D13FFF12916A1741C4CFCB7D541" authority="Wood, 1938" authorityName="Wood" authorityYear="1938" box="[328,798,1588,1614]" class="Mammalia" family="Hyracodontidae" genus="Forstercooperia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="totadentata">
<emphasis id="8486EA82FFF12916A1741C4CFD2CD541" box="[328,645,1588,1614]" italics="true" pageId="3" pageNumber="4">Forstercooperia totadentata</emphasis>
<bibRefCitation id="D2634B61FFF12916A2B11C4DFCB7D541" author="Wood, H. E." box="[653,798,1588,1614]" pageId="3" pageNumber="4" pagination="1 - 20" refId="ref20185" refString="Wood, H. E. 1938. Cooperia totadentata, a remarkable rhinoceros from the Eocene of Mongolia. American Museum Novitates 1012: 1 - 20." type="journal article" year="1938">Wood, 1938</bibRefCitation>
</taxonomicName>
.
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INCLUDED SPECIES: The
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species,
<taxonomicName id="71F24D13FFF12916A2681C23FB11D57C" authority="Qiu and Wang, 2007" authorityName="Qiu and Wang" authorityYear="2007" box="[596,1208,1625,1652]" class="Mammalia" family="Hyracodontidae" genus="Forstercooperia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="mazhuangensis">
<emphasis id="8486EA82FFF12916A2681C23FC12D57C" box="[596,955,1625,1651]" italics="true" pageId="3" pageNumber="4">Forstercooperia mazhuangensis</emphasis>
<bibRefCitation id="D2634B61FFF12916A3FF1C20FB11D57C" author="Qiu, Z. X. &amp; B. Y. Wang" box="[963,1208,1625,1652]" pageId="3" pageNumber="4" refId="ref19615" refString="Qiu, Z. X., and B. Y. Wang. 2007. Paracerathere fossils of China. Beijing: Science Press." type="book" year="2007">Qiu and Wang, 2007</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="71F24D13FFF12916A09C1C06FDAED596" authority="Wang &amp; Bai &amp; Meng &amp; Wang, 2018" authorityName="Wang &amp; Bai &amp; Meng &amp; Wang" authorityYear="2018" box="[160,519,1663,1689]" class="Mammalia" family="Hyracodontidae" genus="Forstercooperia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ulanshirehensis" status="sp. nov.">
<emphasis id="8486EA82FFF12916A09C1C06FDAED596" box="[160,519,1663,1689]" italics="true" pageId="3" pageNumber="4">Forstercooperia ulanshirehensis</emphasis>
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,
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REVISED DIAGNOSIS: Large forstercooperiines with skull length about
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and length of M1M3/m1m3 about
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(
<tableCitation id="FB70032BFFF12916A2931CB0FC8DD5EC" box="[687,804,1737,1763]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="5.[108,182,224,248]" captionStartId-1="6.[108,182,224,248]" captionStartId-2="7.[108,182,224,248]" captionText-0="TABLE 1. Measurements of cranium of Forstercooperia ulanshirehensis (mm). H = height; L = length; Max = maximum; W = width." captionText-1="TABLE 2. Measurements of upper dentition of Forstercooperia ulanshirehensis (mm). L = length; W = width." captionText-2="TABLE 3. Measurements of mandible and lower dentition of Forstercooperia ulanshirehensis (mm). H = height; L = length; W = width." pageId="3" pageNumber="4">table 13</tableCitation>
). Differs from
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<emphasis id="8486EA82FFF12916A3E61CB3FBF6D5EC" box="[986,1119,1738,1763]" italics="true" pageId="3" pageNumber="4">Pappaceras</emphasis>
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in having a larger body size (the length of molar series greater than
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), a transversely narrower postglenoid fossa with a well-developed median ridge separating two articular facets, sagittal crest distinct, occipital condyle comparatively more enlarged, dentition complete; I2 slightly larger than I1 and I3, canines stout and rounded at base; P2 premolarized with protocone and hypocone slightly separated; M3 triangular in occlusal outline along with the absence of metacone and ectoloph; tooth crowns of lower incisors nearly symmetric with distinct mesial and distal flanges, i2 larger than i1 and i3; and talonids of p34 elongate with the presence of entoconid.
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