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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.42.8455" ID-PMC="PMC4225075" ID-Pensoft-Pub="1314-2003-42-57" ID-Pensoft-UUID="220A99134B5CFD1E5E39D053FFB6757D" ID-PubMed="25383012" ID-Zenodo-Dep="576251" ModsDocID="1314-2003-42-57" checkinTime="1451251668893" checkinUser="pensoft" docAuthor="Dorr, Laurence J. &amp; Stergios, Basil" docDate="2014" docId="93F102F3CD8B1C98BE89EC20B1632B43" docLanguage="en" docName="PhytoKeys 42: 57-76" docOrigin="PhytoKeys 42" docPubDate="2014-10-24" docSource="http://dx.doi.org/10.3897/phytokeys.42.8455" docTitle="Pilea nidiae Dorr &amp; Stergios 2014, sp. nov." docType="treatment" docVersion="7" id="220A99134B5CFD1E5E39D053FFB6757D" lastPageNumber="65" masterDocId="220A99134B5CFD1E5E39D053FFB6757D" masterDocTitle="Four new species of Andean Pilea (Urticaceae), with additional notes on the genus in Venezuela" masterLastPageNumber="76" masterPageNumber="57" pageNumber="62" updateTime="1668141252568" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Four new species of Andean Pilea (Urticaceae), with additional notes on the genus in Venezuela</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dorr, Laurence J.</mods:namePart>
<mods:affiliation>Department of Botany, MRC- 166, National Museum of Natural History, Smithsonian Institution, P. O. Box 37012, Washington, DC 20013 - 7012, USA</mods:affiliation>
<mods:nameIdentifier type="email">dorrl@si.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stergios, Basil</mods:namePart>
<mods:affiliation>Universidad Nacional Experimental de los Llanos Occidentales &quot; Ezequiel Zamora &quot; (UNELLEZ), Mesa de Cavacas, Guanare, Estado Portuguesa 3323, Venezuela</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-10-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>42</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>57</mods:start>
<mods:end>76</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.42.8455</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.42.8455</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-42-57</mods:identifier>
<mods:identifier type="Pensoft-UUID">220A99134B5CFD1E5E39D053FFB6757D</mods:identifier>
<mods:identifier type="Zenodo-Dep">576251</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152025196" LSID="urn:lsid:plazi:treatment:93F102F3CD8B1C98BE89EC20B1632B43" httpUri="http://treatment.plazi.org/id/93F102F3CD8B1C98BE89EC20B1632B43" lastPageId="8" lastPageNumber="65" pageId="5" pageNumber="62">
<subSubSection pageId="5" pageNumber="62" type="nomenclature">
<paragraph pageId="5" pageNumber="62">
<taxonomicName LSID="93F102F3-CD8B-1C98-BE89-EC20B1632B43" authority="Dorr &amp; Stergios" authorityName="Dorr &amp; Stergios" authorityYear="2014" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea nidiae" order="Rosales" pageId="5" pageNumber="62" phylum="Tracheophyta" rank="species" species="nidiae" status="sp. nov.">Pilea nidiae Dorr &amp; Stergios</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="62">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pilea nidiae. A Habit; note the unequal leaf laminae at each node B Leaf detail (major lamina upper surface) showing cystoliths C Stipules and stem covered with cystoliths D Staminate inflorescence E Infructescence F, G Pistillate flowers with developing achenes. (A, D from J. L. Luteyn &amp; E. Cotton 9705 (NY); B, C from J. A. Steyermark 55533 (US); E-G from B. Stergios &amp; R. Caracas 19810 (US ))." pageId="5" pageNumber="62">Figure 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="63" type="reference_group">
<paragraph pageId="6" pageNumber="63">
<taxonomicName authority="Dorr &amp; Stergios, 2014" authorityName="Dorr &amp; Stergios" authorityYear="2014" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea" order="Rosales" pageId="6" pageNumber="63" phylum="Tracheophyta" rank="genus" status="sp. nov.">
<pageBreakToken pageId="6" pageNumber="63" start="start">Pilea</pageBreakToken>
</taxonomicName>
sp. D, Dorr et al., Contr. U.S. Natl. Herb. 40: 147. 2000 [2001].
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="63" type="diagnosis">
<paragraph pageId="6" pageNumber="63">Diagnosis.</paragraph>
<paragraph pageId="6" pageNumber="63">
Similar to
<taxonomicName authorityName="Wedd" authorityYear="1852" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea flexuosa" order="Rosales" pageId="6" pageNumber="63" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis italics="true" pageId="6" pageNumber="63">Pilea flexuosa</emphasis>
</taxonomicName>
Wedd. from which it differs by its asymmetrically elliptic to narrowly-elliptic or obovate (versus broadly ovate) laminae that are asymmetrically cuneate (versus rounded or cordate) at the base.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="63" type="type">
<paragraph pageId="6" pageNumber="63">Type.</paragraph>
<paragraph pageId="6" pageNumber="63">
<emphasis bold="true" pageId="6" pageNumber="63">VENEZUELA.</emphasis>
Trujillo: Mpio.
<normalizedToken originalValue="Boconó">Bocono</normalizedToken>
: Parque Nacional Guaramacal, sector vertiente sur, 2800-2900 m, 25-30 Jul 2002,
<emphasis italics="true" pageId="6" pageNumber="63">B. Stergios &amp; R. Caracas 19810</emphasis>
(holotype: PORT [85861]: isotypes: K, MO, US [00772535]).
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="64" pageId="6" pageNumber="63" type="description">
<paragraph pageId="6" pageNumber="63">Description.</paragraph>
<paragraph lastPageId="7" lastPageNumber="64" pageId="6" pageNumber="63">
Herb, to 1.25 m tall; terrestrial; dioecious. Stems erect or prostrate (fide
<emphasis italics="true" pageId="7" pageNumber="64">
<pageBreakToken pageId="7" pageNumber="64" start="start">Stergios</pageBreakToken>
&amp; Caracas 19810
</emphasis>
), succulent, branched, drying reddish-brown, dull purple (fide
<emphasis italics="true" pageId="7" pageNumber="64">Steyermark 55533</emphasis>
) or almost black, glabrous, cystoliths punctiform or short fusiform, often clustered at nodes, internodes 0.8-3.5
<normalizedToken originalValue="×">x</normalizedToken>
1-4 mm (shorter distally), terete in cross-section, angulate when dry, nodes constricted (at least when dry). Stipules 6-11 mm long, narrowly triangular, drying pale brown or tan, persistent. Leaves petiolate, distichous; petioles at the same node unequal by a ratio of 1:11.5-13.5 (-23), canaliculate above, glabrous; major petioles 2.3-2.7 cm long; minor petioles ca 1-2 mm long; laminae of leaves at each node unequal by a ratio of 1:1.7-2.2, major laminae in a pair 6.5-9.5
<normalizedToken originalValue="×">x</normalizedToken>
1.5-3.2 cm, asymmetrically elliptic to narrowly-elliptic or obovate, membranous, 3-nerved with lateral nerves diverging from midrib 1-6 mm above the base, forming pocket domatia where the 3 nerves join, midrib and lateral nerves prominent below, slightly impressed (or not) above, lateral nerves visible almost the entire lamina length but disappearing below the apex, secondary nerves 8-14 pair, borne 70-90° to the midrib and then curved distally, upper surface dark green, drying dark brown, glabrous except for scattered, minute peltate scales, cystoliths fusiform or absent, lower surface pale green drying dark brown, glabrous except for scattered, minute peltate scales, base cuneate, asymmetrical, margin coarsely toothed entire length, apex long acuminate; minor laminae in a pair 3-5
<normalizedToken originalValue="×">x</normalizedToken>
0.8-1.5 cm, otherwise as major laminae. Inflorescences 1-10 per stem, unisexual, green suffused with maroon; bracts ca 2 mm long; bracteoles ca 1 mm long. Staminate inflorescences 1 per axil, 2.8-3.5 cm long, bearing ca 50 flowers in a compact head-like cyme; peduncles 2-3 cm long, glabrous with minute, scattered peltate scales, occasionally cystoliths present; pedicels ca 0.25 mm long. Staminate flowers ca 1.5
<normalizedToken originalValue="×">x</normalizedToken>
1-1.25 mm (mature flowers not seen); tepals 4, ca 2 mm long, notched inside; stamens 4. Pistillate inflorescences 1 or 2 per axil, ca 5 mm long, bearing ca 50 flowers in a
<normalizedToken originalValue="±">+/-</normalizedToken>
loose cyme; peduncles ca 2 mm long, glabrous; pedicels minute. Pistillate flowers ca 0.5-0.75 mm long, cucullate tepal ca 0.5 mm long,
<normalizedToken originalValue="±">+/-</normalizedToken>
lanceolate; lateral tepals minute. Infructescences 1-2.5 cm long, frequently including receptive pistillate flowers; peduncles 0.7-1.8 cm long; achenes ca 1.25
<normalizedToken originalValue="×">x</normalizedToken>
1 mm, compressed, asymmetrically ellipsoid or lachrymiform, verrucose, margin narrowly thickened with a very narrow hyaline wing.
</paragraph>
<caption pageId="7" pageNumber="64" start="Figure 4" startId="F4">
<paragraph pageId="7" pageNumber="64">
<emphasis bold="true" pageId="7" pageNumber="64">Figure 4.</emphasis>
<taxonomicName authorityName="Dorr &amp; Stergios" authorityYear="2014" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea nidiae" order="Rosales" pageId="7" pageNumber="64" phylum="Tracheophyta" rank="species" species="nidiae">
<emphasis italics="true" pageId="7" pageNumber="64">Pilea nidiae</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="7" pageNumber="64">A</emphasis>
Habit; note the unequal leaf laminae at each node
<emphasis bold="true" pageId="7" pageNumber="64">B</emphasis>
Leaf detail (major lamina upper surface) showing cystoliths
<emphasis bold="true" pageId="7" pageNumber="64">C</emphasis>
Stipules and stem covered with cystoliths
<emphasis bold="true" pageId="7" pageNumber="64">D</emphasis>
Staminate inflorescence
<emphasis bold="true" pageId="7" pageNumber="64">E</emphasis>
Infructescence
<emphasis bold="true" pageId="7" pageNumber="64">F, G</emphasis>
Pistillate flowers with developing achenes. (
<emphasis bold="true" pageId="7" pageNumber="64">A, D</emphasis>
from
<emphasis italics="true" pageId="7" pageNumber="64">J.L. Luteyn &amp; E. Cotton 9705</emphasis>
(NY);
<emphasis bold="true" pageId="7" pageNumber="64">B, C</emphasis>
from
<emphasis italics="true" pageId="7" pageNumber="64">J.A. Steyermark 55533</emphasis>
(US);
<emphasis bold="true" pageId="7" pageNumber="64">E-G</emphasis>
from
<emphasis italics="true" pageId="7" pageNumber="64">B. Stergios &amp; R. Caracas 19810</emphasis>
(US)).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="7" pageNumber="64" type="distribution and ecology">
<paragraph pageId="7" pageNumber="64">Distribution and ecology.</paragraph>
<paragraph pageId="7" pageNumber="64">Known only from the Andes of Venezuela (Lara and Trujillo states) where it is found in the moist, shaded understory of montane forest; 2285-2900 m.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="64" type="etymology">
<paragraph pageId="7" pageNumber="64">Etymology.</paragraph>
<paragraph pageId="7" pageNumber="64">This species is named in honor of Nidia Cuello, Director of Herbario PORT, UNELLEZ, Guanare, and expert on the vegetative ecology of Guaramacal National Park.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="64" type="specimens examined">
<paragraph pageId="7" pageNumber="64">Specimens examined.</paragraph>
<paragraph pageId="7" pageNumber="64">
<emphasis bold="true" pageId="7" pageNumber="64">VENEZUELA.</emphasis>
<emphasis bold="true" pageId="7" pageNumber="64">Lara:</emphasis>
Mpio.
<normalizedToken originalValue="Morán">Moran</normalizedToken>
: Trail from Humocaro to Buenos Aires (
<normalizedToken originalValue="caserío">caserio</normalizedToken>
) below
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
Los Rosas (ca
<geoCoordinate degrees="09" direction="north" minutes="40" orientation="latitude" precision="925" value="9.666667">09°40'N</geoCoordinate>
,
<geoCoordinate degrees="070" direction="west" minutes="05" orientation="longitude" precision="925" value="-70.083336">070°05'W</geoCoordinate>
), 2750 m, 25 Jul 1979,
<emphasis italics="true" pageId="7" pageNumber="64">R. Liesner et al. 8158</emphasis>
(MO, VEN); Between Buenos Aires to Canyon of El Callado, above Humocaro Alto, 2285-2740 m, 12 Feb 1944,
<emphasis italics="true" pageId="7" pageNumber="64">J.A. Steyermark 55533</emphasis>
(NY, US).
<emphasis bold="true" pageId="7" pageNumber="64">Trujillo:</emphasis>
Mpio.
<normalizedToken originalValue="Boconó">Bocono</normalizedToken>
:
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
Guaramacal, 20-21 km beyond jct. with hwy. NE of
<normalizedToken originalValue="Boconó">Bocono</normalizedToken>
, ca
<geoCoordinate degrees="09" direction="north" minutes="13" orientation="latitude" precision="925" value="9.216666">09°13'N</geoCoordinate>
,
<geoCoordinate degrees="070" direction="west" minutes="13" orientation="longitude" precision="925" value="-70.21667">070°13'W</geoCoordinate>
, 2640-2700 m, 14 Mar 1984,
<emphasis italics="true" pageId="7" pageNumber="64">J.L Luteyn &amp; E. Cotton 9705</emphasis>
(MO, NY, PORT, VEN).
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="65" pageId="7" pageNumber="64" type="discussion">
<paragraph pageId="7" pageNumber="64">Discussion.</paragraph>
<paragraph pageId="7" pageNumber="64">
<taxonomicName authorityName="Dorr &amp; Stergios" authorityYear="2014" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea nidiae" order="Rosales" pageId="7" pageNumber="64" phylum="Tracheophyta" rank="species" species="nidiae">
<emphasis italics="true" pageId="7" pageNumber="64">Pilea nidiae</emphasis>
</taxonomicName>
belongs in the Heterophyllae group of
<bibRefCitation author="Weddell, HA" journalOrPublisher="Prodromus" pageId="11" pageNumber="68" pagination="104 - 163" refId="B11" refString="Weddell, HA, 1869. Pilea. Prodromus 16 (1): 104 - 163" title="Pilea." volume="16" year="1869">Weddell (1869)</bibRefCitation>
. Its 3-nerved, toothed leaves that are unequal in size at each node and conspicuous, persistent stipules place it in the Flexuosae group of
<bibRefCitation author="Killip, EP" journalOrPublisher="Contributions from the United States National Herbarium" pageId="10" pageNumber="67" pagination="367 - 394" refId="B4" refString="Killip, EP, 1936. New species of Pilea from the Andes. Contributions from the United States National Herbarium 26 (8): 367 - 394" title="New species of Pilea from the Andes." volume="26" year="1936">Killip (1936</bibRefCitation>
,
<bibRefCitation author="Killip, EP" journalOrPublisher="Contributions from the United States National Herbarium" pageId="10" pageNumber="67" pagination="475 - 530" refId="B5" refString="Killip, EP, 1939. The Andean species of Pilea. Contributions from the United States National Herbarium 26 (10): 475 - 530" title="The Andean species of Pilea." volume="26" year="1939">1939</bibRefCitation>
). We have not encountered any other Andean species from Venezuela with stipules that are as large as those of
<taxonomicName authorityName="Dorr &amp; Stergios" authorityYear="2014" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea nidiae" order="Rosales" pageId="7" pageNumber="64" phylum="Tracheophyta" rank="species" species="nidiae">
<emphasis italics="true" pageId="7" pageNumber="64">Pilea nidiae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph lastPageId="8" lastPageNumber="65" pageId="7" pageNumber="64">
The leaf laminae of one of the paratype collections (
<emphasis italics="true" pageId="8" pageNumber="65">
<pageBreakToken pageId="8" pageNumber="65" start="start">Steyermark</pageBreakToken>
55533
</emphasis>
) are narrower than in the type of
<taxonomicName authorityName="Dorr &amp; Stergios" authorityYear="2014" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea nidiae" order="Rosales" pageId="8" pageNumber="65" phylum="Tracheophyta" rank="species" species="nidiae">
<emphasis italics="true" pageId="8" pageNumber="65">Pilea nidiae</emphasis>
</taxonomicName>
and densely covered in cystoliths above and below (the type mostly lacks cystoliths). All other characters (leaf shape, venation, toothing, etc.) agree with our concept of this new species.
</paragraph>
<caption ID-Table-UUID="4D5EC32A4205D4BA663E7678B059A7BF" httpUri="http://table.plazi.org/id/4D5EC32A4205D4BA663E7678B059A7BF" pageId="8" pageNumber="65" start="Table 4" startId="T4">
<paragraph pageId="8" pageNumber="65">
<emphasis bold="true" pageId="8" pageNumber="65">Table 4.</emphasis>
Diagnostic characters that distinguish
<taxonomicName authorityName="Dorr &amp; Stergios" authorityYear="2014" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea nidiae" order="Rosales" pageId="8" pageNumber="65" phylum="Tracheophyta" rank="species" species="nidiae">
<emphasis italics="true" pageId="8" pageNumber="65">Pilea nidiae</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Wedd" authorityYear="1852" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea flexuosa" order="Rosales" pageId="8" pageNumber="65" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis italics="true" pageId="8" pageNumber="65">Pilea flexuosa</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="8" pageNumber="65">
<table pageId="8" pageNumber="65">
<tr pageId="8" pageNumber="65">
<th colspan="1" pageId="8" pageNumber="65" rowspan="1">Characters</th>
<th colspan="1" pageId="8" pageNumber="65" rowspan="1">
<taxonomicName authorityName="Dorr &amp; Stergios" authorityYear="2014" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea nidiae" order="Rosales" pageId="8" pageNumber="65" phylum="Tracheophyta" rank="species" species="nidiae">
<emphasis italics="true" pageId="8" pageNumber="65">Pilea nidiae</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="8" pageNumber="65" rowspan="1">
<taxonomicName authorityName="Wedd" authorityYear="1852" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea flexuosa" order="Rosales" pageId="8" pageNumber="65" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis italics="true" pageId="8" pageNumber="65">Pilea flexuosa</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="8" pageNumber="65">
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">Stipule shape (length)</td>
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">narrowly triangular (6-11 mm)</td>
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">ovate orbicular (4-6 mm)</td>
</tr>
<tr pageId="8" pageNumber="65">
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">Major leaf lamina size</td>
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">
6.5-9.5
<normalizedToken originalValue="×">x</normalizedToken>
1.5-3.2 cm
</td>
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">
2-6
<normalizedToken originalValue="×">x</normalizedToken>
1.5-4 cm
</td>
</tr>
<tr pageId="8" pageNumber="65">
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">Leaf shape</td>
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">elliptic to narrowly elliptic or obovate</td>
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">broadly ovate</td>
</tr>
<tr pageId="8" pageNumber="65">
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">Leaf base</td>
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">cuneate</td>
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">rounded or cordate</td>
</tr>
<tr pageId="8" pageNumber="65">
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">Leaf apex</td>
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">long acuminate</td>
<td colspan="1" pageId="8" pageNumber="65" rowspan="1">abruptly acute to acuminate</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="65" type="conservation status">
<paragraph pageId="8" pageNumber="65">Conservation status.</paragraph>
<paragraph pageId="8" pageNumber="65">
Using IUCN criteria (
<bibRefCitation author="International Union for Conservation of Nature and Natural Resources Species Survival Commission" journalOrPublisher="International Union for Conservation of Nature and Natural Resources, Gland and Cambridge" pageId="10" pageNumber="67" refId="B3" refString="International Union for Conservation of Nature and Natural Resources Species Survival Commission, 2001. IUCN Red List Categories: Version 3.1 (second edition). International Union for Conservation of Nature and Natural Resources, Gland and Cambridge" title="IUCN Red List Categories: Version 3.1 (second edition)." year="2001">IUCN 2001</bibRefCitation>
) we tentatively consider
<taxonomicName authorityName="Dorr &amp; Stergios" authorityYear="2014" class="Magnoliopsida" family="Urticaceae" genus="Pilea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pilea nidiae" order="Rosales" pageId="8" pageNumber="65" phylum="Tracheophyta" rank="species" species="nidiae">
<emphasis italics="true" pageId="8" pageNumber="65">Pilea nidiae</emphasis>
</taxonomicName>
to be Endangered (E). The known range of the species is less than 5000 km2 (IUCN criterion B1) and there are only four known populations and of these only two are in a protected area (IUCN criterion B1(a)). We know nothing, however, about the dynamics of these populations and whether or not they are declining.
</paragraph>
</subSubSection>
</treatment>
</document>