treatments-xml/data/7A/DB/E6/7ADBE6F65D695409BC6E9468436F2166.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.137.46544" ID-PMC="PMC6962254" ID-Pensoft-Pub="1314-2003-137-1" ID-Pensoft-UUID="FB2C14EFA257564AA84CD47711BBC273" ID-PubMed="31969792" ModsDocID="1314-2003-137-1" checkinTime="1578614985992" checkinUser="pensoft" docAuthor="Zaika, Maxim A., Kilian, Norbert, Jones, Katy, Krinitsina, Anastasiya A., Nilova, Maya V., Speranskaya, Anna S. &amp; Sukhorukov, Alexander P." docDate="2020" docId="7ADBE6F65D695409BC6E9468436F2166" docLanguage="en" docName="PhytoKeys 137: 1-85" docOrigin="PhytoKeys 137" docSource="http://dx.doi.org/10.3897/phytokeys.137.46544" docTitle="Takhtajaniantha Nazarova, Biol. Zhurn. Armenii 43: 179. 1990" docType="treatment" docVersion="5" id="FB2C14EFA257564AA84CD47711BBC273" lastPageNumber="1" masterDocId="FB2C14EFA257564AA84CD47711BBC273" masterDocTitle="Scorzonera sensu lato (Asteraceae, Cichorieae) - taxonomic reassessment in the light of new molecular phylogenetic and carpological analyses" masterLastPageNumber="85" masterPageNumber="1" pageNumber="1" updateTime="1668139594603" updateUser="ExternalLinkService">
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<mods:title>Scorzonera sensu lato (Asteraceae, Cichorieae) - taxonomic reassessment in the light of new molecular phylogenetic and carpological analyses</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zaika, Maxim A.</mods:namePart>
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<mods:namePart>Kilian, Norbert</mods:namePart>
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<mods:namePart>Jones, Katy</mods:namePart>
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<mods:namePart>Krinitsina, Anastasiya A.</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nilova, Maya V.</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Speranskaya, Anna S.</mods:namePart>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sukhorukov, Alexander P.</mods:namePart>
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<mods:date>2020</mods:date>
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<mods:number>137</mods:number>
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<treatment ID-GBIF-Taxon="161671725" LSID="urn:lsid:plazi:treatment:7ADBE6F65D695409BC6E9468436F2166" httpUri="http://treatment.plazi.org/id/7ADBE6F65D695409BC6E9468436F2166" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="7ADBE6F6-5D69-5409-BC6E-9468436F2166" authority="Nazarova, Biol. Zhurn. Armenii 43: 179. 1990." class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Takhtajaniantha Nazarova, Biol. Zhurn. Armenii 43: 179. 1990.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
= 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Fibrillosae">Scorzonera sect. Fibrillosae</taxonomicName>
Nakai, Rep. Inst. Sci. Res. Manchoukuo, ser. 1, 6: 171. 1937. Type: 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera glabra" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="glabra">Scorzonera glabra</taxonomicName>
Rupr. [= 
<taxonomicName family="Asteraceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="S. austriaca" order="Asterales" pageId="0" pageNumber="1" rank="species" species="austriaca">S. austriaca</taxonomicName>
Willd.]
</paragraph>
<paragraph pageId="0" pageNumber="1">
= 
<taxonomicName authority="subsect. Egregia" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Scorzonera subsect. Egregia</taxonomicName>
Kult., Tau-Sagyz Ekol. Osnovy Vvedeniya Ego 
<normalizedToken originalValue="Kul’t">Kul't</normalizedToken>
.: 108. 1938.
</paragraph>
<paragraph pageId="0" pageNumber="1">
≡ 
<taxonomicName authority="subg. Egregia" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Scorzonera subg. Egregia</taxonomicName>
(Kult.) Ovcz., Soobshch. Tadzhiksk. Fil. Akad. Nauk SSSR 20: 54. 1950.
</paragraph>
<paragraph pageId="0" pageNumber="1">
≡ 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Egregia">Scorzonera sect. Egregia</taxonomicName>
(Kult.) Lipsch., Fl. URSS 29: 58. 1964. Lectotype (designated here): 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera tau-saghyz" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tau-saghyz">Scorzonera tau-saghyz</taxonomicName>
Lipsch. &amp; G.G.Bosse
</paragraph>
<paragraph pageId="0" pageNumber="1">
= 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Pusillae">Scorzonera sect. Pusillae</taxonomicName>
Lipsch., Fragm. Monogr. Gen. 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Scorzonera</taxonomicName>
2: 59. 1939. Type: 
<taxonomicName authorityName="Pall" authorityYear="1773" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera pusilla" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pusilla">Scorzonera pusilla</taxonomicName>
Pall.
</paragraph>
<paragraph pageId="0" pageNumber="1">
= 
<taxonomicName authorityName="Maxim" authorityYear="1888" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Capito">Scorzonera sect. Capito</taxonomicName>
Tzvelev, Rast. Tsentral. Azii 14b: 115. 2009. Type: 
<taxonomicName authorityName="Maxim" authorityYear="1888" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera capito" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="capito">Scorzonera capito</taxonomicName>
Maxim.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha pusilla" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pusilla">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha pusilla</emphasis>
</taxonomicName>
(Pall.) Nazarova
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnostic features">
<paragraph pageId="0" pageNumber="1">Diagnostic features.</paragraph>
<paragraph pageId="0" pageNumber="1">Perennials; caudex often with dark brown fibrous leaf sheath residues and base of the rosulate leaves often lanate; pollen with 6 lacunae.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Habit, life form, subterranean parts:</emphasis>
perennial herbs or subshrublets with a taproot or a tuber.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Leaves:</emphasis>
rosulate or crowded in the basal part of the stem, rarely dispersed along the stem, sessile or petiolate, filiform or ovate, apically sometimes hooked (
<taxonomicName lsidName="T. pusilla" pageId="0" pageNumber="1" rank="species" species="pusilla">
<emphasis italics="true" pageId="0" pageNumber="1">T. pusilla</emphasis>
</taxonomicName>
), margin flat or undulate, entire or denticulate.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Stem, synflorescence:</emphasis>
stem solitary or branched, scape-like or leafy, capitula terminal.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pollen:</emphasis>
echinolophate, tricolporate and each colpus divided into 2 lacunae, with 6 abporal lacunae (sometimes with spineless areas resembling rudimentary lacunae (
<bibRefCitation DOI="https://doi.org/10.1080/00173138209427691" author="Blackmore, S" journalOrPublisher="Grana" pageId="0" pageNumber="1" pagination="149 - 160" refId="B9" refString="Blackmore, S, 1982. Palynology of subtribe Scorzonerinae (Compositae. Lactuceae) and its taxonomic significance. Grana 21 (3): 149 - 160, DOI: https://doi.org/10.1080/00173138209427691" title="Palynology of subtribe Scorzonerinae (Compositae. Lactuceae) and its taxonomic significance." url="https://doi.org/10.1080/00173138209427691" volume="21" year="1982">Blackmore 1982</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Capitula:</emphasis>
glabrous to arachnoid hairy or tomentose, phyllaries in several series, outer phyllaries smaller than inner ones, triangular-ovate, inner phyllaries linear to ovate, receptacle glabrous, flat, with many or rarely few (up to 12: 
<taxonomicName lsidName="T. pseudodivaricata" pageId="0" pageNumber="1" rank="species" species="pseudodivaricata">
<emphasis italics="true" pageId="0" pageNumber="1">T. pseudodivaricata</emphasis>
</taxonomicName>
) florets; florets yellow (Fig. 
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Images of living plants. A, B Ramaliella tortuosissima A general view B close up view of the capitula (Jordan, Nakeb, 1250 m alt., 8 May 2019). Photograph by O. Fragman-Sapir C Pseudopodospermum papposum (Jordan, Nakeb, 8 May 2019). Photograph by O. Fragman-Sapir D Takhtajaniantha austriaca (Italy, South Tyrol, Schlanders, 6 May 2010). Photograph by R. Hand E Scorzonera humilis (Germany, Ammersee, 15 May 2009). Photograph by C. Niederbichler F Scorzonera meyeri (Russia, Karachaevo-Cherkessiya, Teberda, 14 July 2005). Photograph by A. S. Zernov." figureDoi="10.3897/phytokeys.137.46544.figure16" httpUri="https://binary.pensoft.net/fig/370364" pageId="0" pageNumber="1">16D</figureCitation>
) or with dark red veins, usually clearly exeeding the incolucre.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Achenes:</emphasis>
5-15 mm, without carpopodium, straight or slightly curved, often with 10 ribs or rarely roundish, glabrous or hairy, sometimes (as in 
<taxonomicName lsidName="T. pusilla" pageId="0" pageNumber="1" rank="species" species="pusilla">
<emphasis italics="true" pageId="0" pageNumber="1">T. pusilla</emphasis>
</taxonomicName>
) with stout conglomerations; achene wall with parenchyma of two types, with thick- and thin-walled cells, present only as subepidermal continuous layer(s) or only with thick-walled cells, air cavities absent, sclerenchyma with continuous layer with a narrow or wider hunch on either side of the principal rib or, as in 
<taxonomicName lsidName="T. pseudodivaricata" pageId="0" pageNumber="1" rank="species" species="pseudodivaricata">
<emphasis italics="true" pageId="0" pageNumber="1">T. pseudodivaricata</emphasis>
</taxonomicName>
, without hunches, but discontinuous forming 5 bundles (with a gap in the principal ribs), or continuous, its fibres orientated parallel to the fruit axis, tannins absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pappus:</emphasis>
7-28 mm, ice white or dirty white, bristles scabrid in upper part.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Chromosome number:</emphasis>
x = 7 (diploids); 2n = 28 in 
<taxonomicName lsidName="T. pusilla" pageId="0" pageNumber="1" rank="species" species="pusilla">
<emphasis italics="true" pageId="0" pageNumber="1">T. pusilla</emphasis>
</taxonomicName>
(amphiploid); in 
<taxonomicName lsidName="T. tau-saghyz" pageId="0" pageNumber="1" rank="species" species="tau-saghyz">
<emphasis italics="true" pageId="0" pageNumber="1">T. tau-saghyz</emphasis>
</taxonomicName>
polyploid cytoraces 2n = 14, 21, 28, 42 and aneuploids with chromosome numbers 15, 16, 17, 18, 22 and 24, 28 and 42 are known (
<bibRefCitation author="Krayevoy, SU" journalOrPublisher="Botanicheskii Zhurnal" pageId="0" pageNumber="1" pagination="367 - 375" refId="B53" refString="Krayevoy, SU, 1934. K voprosu o khromosomnykh variatsiyah v Scorzonera tau-saghyz. Botanicheskii Zhurnal 19: 367 - 375" title="K voprosu o khromosomnykh variatsiyah v Scorzonera tau-saghyz." volume="19" year="1934">Krayevoy 1934</bibRefCitation>
; 
<bibRefCitation author="Poddubnaya-Arnoldi, VA" journalOrPublisher="Botanicheskii Zhurnal" pageId="0" pageNumber="1" pagination="338 - 366" refId="B80" refString="Poddubnaya-Arnoldi, VA, Steshina, N, Sosnovets, A, 1934. Razvitiye i zhiznesposobnost' pyltsy Scorzonera tau-saghyz Lipsch et Bosse. Botanicheskii Zhurnal 19: 338 - 366" title="Razvitiye i zhiznesposobnost' pyltsy Scorzonera tau-saghyz Lipsch et Bosse." volume="19" year="1934">Poddubnaya-Arnoldi et al. 1934</bibRefCitation>
; 
<bibRefCitation author="Sosnovets, AP" journalOrPublisher="Botanicheskii Zhurnal" pageId="0" pageNumber="1" pagination="1813 - 1815" refId="B91" refString="Sosnovets, AP, 1960. Tsitologiya roda Scorzonera L. Botanicheskii Zhurnal 45: 1813 - 1815" title="Tsitologiya roda Scorzonera L." volume="45" year="1960">Sosnovets 1960</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Species.</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="1">
(1) 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha austriaca" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="austriaca">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha austriaca</emphasis>
</taxonomicName>
(Willd.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. (Fig. 
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Images of living plants. A, B Ramaliella tortuosissima A general view B close up view of the capitula (Jordan, Nakeb, 1250 m alt., 8 May 2019). Photograph by O. Fragman-Sapir C Pseudopodospermum papposum (Jordan, Nakeb, 8 May 2019). Photograph by O. Fragman-Sapir D Takhtajaniantha austriaca (Italy, South Tyrol, Schlanders, 6 May 2010). Photograph by R. Hand E Scorzonera humilis (Germany, Ammersee, 15 May 2009). Photograph by C. Niederbichler F Scorzonera meyeri (Russia, Karachaevo-Cherkessiya, Teberda, 14 July 2005). Photograph by A. S. Zernov." figureDoi="10.3897/phytokeys.137.46544.figure16" httpUri="https://binary.pensoft.net/fig/370364" pageId="0" pageNumber="1">16D</figureCitation>
) ≡ 
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera austriaca" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="austriaca">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera austriaca</emphasis>
</taxonomicName>
Willd., Sp. Pl. 3(3): 1498. 1803 (incl. 
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. glabra" order="Asterales" pageId="0" pageNumber="1" rank="species" species="glabra">
<emphasis italics="true" pageId="0" pageNumber="1">S. glabra</emphasis>
</taxonomicName>
Rupr.). urn:lsid:ipni.org:names:77204033-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(2) 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha capito" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="capito">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha capito</emphasis>
</taxonomicName>
(Maxim.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡ 
<taxonomicName authorityName="Maxim" authorityYear="1888" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera capito" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="capito">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera capito</emphasis>
</taxonomicName>
Maxim., Bull. Acad. Imp. Sci. 
<normalizedToken originalValue="Saint-Pétersb">Saint-Petersb</normalizedToken>
. 32: 491. 1888. urn:lsid:ipni.org:names:77204030-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(3) 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha crispa" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="crispa">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha crispa</emphasis>
</taxonomicName>
(M.Bieb.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡ 
<taxonomicName authorityName="Bieb" authorityYear="1808" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera crispa" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="crispa">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera crispa</emphasis>
</taxonomicName>
M.Bieb., Fl. Taur.-Caucas. 2: 234. 1808. urn:lsid:ipni.org:names:77204028-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(4) 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha ikonnikovii" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="ikonnikovii">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha ikonnikovii</emphasis>
</taxonomicName>
(Krasch. &amp; Lipsch.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡ 
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera ikonnikovii" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="ikonnikovii">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera ikonnikovii</emphasis>
</taxonomicName>
Krasch. &amp; Lipsch. in Lipschitz, Fragm. Monogr. Gen. 
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Scorzonera</taxonomicName>
1: 109. 1935. urn:lsid:ipni.org:names:77204024-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(5) 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha mongolica" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="mongolica">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha mongolica</emphasis>
</taxonomicName>
(Maxim.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡ 
<taxonomicName authorityName="Maxim" authorityYear="1888" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera mongolica" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="mongolica">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera mongolica</emphasis>
</taxonomicName>
Maxim., Bull. Acad. Imp. Sci. 
<normalizedToken originalValue="Saint-Pétersb">Saint-Petersb</normalizedToken>
. 32: 492. 1888. urn:lsid:ipni.org:names:77204020-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(6) 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha pseudodivaricata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pseudodivaricata">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha pseudodivaricata</emphasis>
</taxonomicName>
(Lipsch.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡ 
<taxonomicName authorityName="Lipsch" authorityYear="1933" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera pseudodivaricata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pseudodivaricata">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera pseudodivaricata</emphasis>
</taxonomicName>
Lipsch., Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 42: 158. 1933. urn:lsid:ipni.org:names:77204019-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(7) 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha pusilla" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pusilla">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha pusilla</emphasis>
</taxonomicName>
(Pall.) Nazarova
</paragraph>
<paragraph pageId="0" pageNumber="1">
(8) 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha subacaulis" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="subacaulis">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha subacaulis</emphasis>
</taxonomicName>
(Regel) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡ 
<taxonomicName authorityName="Regel" authorityYear="1880" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera austriaca var. subacaulis" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="austriaca" variety="subacaulis">Scorzonera austriaca var. subacaulis</taxonomicName>
Regel, Trudy Imp. S.-Peterburgsk. Bot. Sada 6: 323. 1880. urn:lsid:ipni.org:names:77204015-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(9) 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha tau-saghyz" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tau-saghyz">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha tau-saghyz</emphasis>
</taxonomicName>
(Lipsch. &amp; G.G.Bosse) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡ 
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera tau-saghyz" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tau-saghyz">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera tau-saghyz</emphasis>
</taxonomicName>
Lipsch. &amp; G.G.Bosse, Trudy Nauchno-Issl. Lab. Tresta &quot;Kauchukonos&quot; 4: 27. 1930. urn:lsid:ipni.org:names:77204010-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(10) 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha veresczaginii" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="veresczaginii">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha veresczaginii</emphasis>
</taxonomicName>
(Kamelin &amp; S.V.Smirn.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡ 
<taxonomicName authorityName="Kamelin &amp; S. V. Smirn" authorityYear="2002" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera veresczaginii" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="veresczaginii">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera veresczaginii</emphasis>
</taxonomicName>
Kamelin &amp; S.V.Smirn., Turczaninowia 5(1): 17. 2002. urn:lsid:ipni.org:names:77204008-1
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution area.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">N Africa</emphasis>
: EG. 
<emphasis bold="true" pageId="0" pageNumber="1">Asia-Temperate</emphasis>
: AF; AR; AZ; CN; IL; IR; JO; KG; KP; KR; KZ; MN; RU (Asiatic part); SA; TJ; TM; UZ. 
<emphasis bold="true" pageId="0" pageNumber="1">Asia-Tropical</emphasis>
: PK. 
<emphasis bold="true" pageId="0" pageNumber="1">Europe</emphasis>
: AL; AT; BA; BG; BY; CH; CZ; DE; FR; GR; HR; HU; IT; MD; ME; RO; RS; RU; SI; SK; UA.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The following species may belong to 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha</emphasis>
</taxonomicName>
, based on their morphological characters (they were included neither in the carpological nor in molecular investigations):
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="N. Kilian" authorityYear="2011" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera aniana" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="aniana">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera aniana</emphasis>
</taxonomicName>
N.Kilian, 
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. grubovii" order="Asterales" pageId="0" pageNumber="1" rank="species" species="grubovii">
<emphasis italics="true" pageId="0" pageNumber="1">S. grubovii</emphasis>
</taxonomicName>
Lipsch., 
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. karataviensis" order="Asterales" pageId="0" pageNumber="1" rank="species" species="karataviensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. karataviensis</emphasis>
</taxonomicName>
Kult. (possibly a synonym of 
<taxonomicName lsidName="T. tau-saghyz" pageId="0" pageNumber="1" rank="species" species="tau-saghyz">
<emphasis italics="true" pageId="0" pageNumber="1">T. tau-saghyz</emphasis>
</taxonomicName>
), 
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. manshurica" order="Asterales" pageId="0" pageNumber="1" rank="species" species="manshurica">
<emphasis italics="true" pageId="0" pageNumber="1">S. manshurica</emphasis>
</taxonomicName>
Nakai, 
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. pamirica" order="Asterales" pageId="0" pageNumber="1" rank="species" species="pamirica">
<emphasis italics="true" pageId="0" pageNumber="1">S. pamirica</emphasis>
</taxonomicName>
C.Shih, 
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. sinensis" order="Asterales" pageId="0" pageNumber="1" rank="species" species="sinensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. sinensis</emphasis>
</taxonomicName>
(Lipsch. &amp; Krasch.) Nakai
</paragraph>
</subSubSection>
</treatment>
</document>