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<document ID-DOI="http://dx.doi.org/10.3897/asp.79.e67185" ID-GBIF-Dataset="0f8f850f-4b02-4477-b780-bf054d7c6131" ID-GBIF-Taxon="182075318" ID-Pensoft-Pub="1864-8312-79-115" ID-Pensoft-UUID="A4EA6692C24753F7B3CB5DFCAF1EFFAD" ID-ZooBank="6D857849A9ED4DA3A5B834FDAAB0A318" ModsDocID="1864-8312-79-115" checkinTime="1621063390318" checkinUser="pensoft" docAuthor="da Silveira, Luiz Felipe Lima, Roza, Andre Silva, Vaz, Stephanie &amp; Mermudes, Jose Ricardo M." docDate="2021" docId="7A66CDD6376A5D02B26DDF7AEB21325C" docLanguage="en" docName="ArthropodSystPhyl 79: 115-150" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 79" docPubDate="2021-05-14" docSource="http://dx.doi.org/10.3897/asp.79.e67185" docTitle="Costalampys pauper da Silveira &amp; Roza &amp; Vaz &amp; Mermudes 2021, comb. nov." docType="treatment" docVersion="3" id="A4EA6692C24753F7B3CB5DFCAF1EFFAD" lastPageNumber="115" masterDocId="A4EA6692C24753F7B3CB5DFCAF1EFFAD" masterDocTitle="Description and phylogenetic analysis of a new firefly genus from the Atlantic Rainforest, with five new species and new combinations (Coleoptera: Lampyridae: Lampyrinae)" masterLastPageNumber="150" masterPageNumber="115" pageId="0" pageNumber="115" updateTime="1632022650993" updateUser="ExternalLinkService">
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<mods:title>Description and phylogenetic analysis of a new firefly genus from the Atlantic Rainforest, with five new species and new combinations (Coleoptera: Lampyridae: Lampyrinae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>da Silveira, Luiz Felipe Lima</mods:namePart>
<mods:affiliation>Biology Department, Western Carolina University, 122 Central Dr, Cullowhee, NC 28723, United States of America; Luiz Felipe Lima da Silveira [silveira. lfl @ gmail. com]</mods:affiliation>
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<mods:namePart>Roza, Andre Silva</mods:namePart>
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<mods:affiliation>Laboratorio de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, A 1 - 107, Bloco A, Av. Carlos Chagas Filho, 373, Cidade Universitaria, Ilha do Fundao, Rio de Janeiro, Brasil; Andre Silva Roza [andreroza 1993 @ gmail. com]; Jose Ricardo Miras Mermudes [jrmermudes @ gmail. com]</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vaz, Stephanie</mods:namePart>
<mods:affiliation>Laboratorio de Ecologia de Insetos, Departamento de Ecologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, A 0 - 111, Bloco A, Av. Carlos Chagas Filho, 373, Cidade Universitaria, Ilha do Fundao, Rio de Janeiro, Brasil; Stephanie Vaz [anievaz @ gmail. com]</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Mermudes, Jose Ricardo M.</mods:namePart>
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<mods:affiliation>Laboratorio de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, A 1 - 107, Bloco A, Av. Carlos Chagas Filho, 373, Cidade Universitaria, Ilha do Fundao, Rio de Janeiro, Brasil; Andre Silva Roza [andreroza 1993 @ gmail. com]; Jose Ricardo Miras Mermudes [jrmermudes @ gmail. com]</mods:affiliation>
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<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date>2021</mods:date>
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<paragraph pageId="0" pageNumber="115">
<taxonomicName LSID="7A66CDD6-376A-5D02-B26D-DF7AEB21325C" authority="(Olivier, 1899)" authorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" authorityYear="2021" baseAuthorityName="Olivier" baseAuthorityYear="1899" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys pauper" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="pauper" status="comb. nov.">Costalampys pauper (Olivier, 1899)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="115">comb. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">Figs 19A-M</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Distribution map of Costalampys gen. nov. species." figureDoi="10.3897/asp.79.e67185.figure22" httpUri="https://binary.pensoft.net/fig/543225" pageId="0" pageNumber="115">, 22</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="115" type="reference_group">
<paragraph pageId="0" pageNumber="115">
<taxonomicName authorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" authorityYear="2021" baseAuthorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" baseAuthorityYear="2021" class="Insecta" family="Lampyridae" genus="Cladodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladodes pauper" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="pauper" status="comb. nov.">Cladodes pauper</taxonomicName>
Olivier, 1899:90 (desc.); 
<bibRefCitation author="Blackwelder, RE" journalOrPublisher="Bulletin of the United States National Museum" pageId="0" pageNumber="115" pagination="343 - 550" refId="B4" refString="Blackwelder, RE, 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America Part 3. Bulletin of the United States National Museum 185: 343 - 550" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America Part 3." volume="185" year="1945">Blackwelder 1945</bibRefCitation>
:352 (cat.); 
<bibRefCitation author="McDermott, FA" editor="Steel, WO" journalOrPublisher="Pars 9 (editio secunda). W. Junk, S'Gravenhage" pageId="0" pageNumber="115" pagination="1 - 149" refId="B34" refString="McDermott, FA, 1966. Lampyridae. In: Steel, WO, Ed., , Coleopterorum Catalogus Supplementa. Pars 9 (editio secunda). W. Junk, S'Gravenhage: 1 - 149" title="Lampyridae." volumeTitle=", Coleopterorum Catalogus Supplementa." year="1966">McDermott 1966</bibRefCitation>
:82 (cat.).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="description">
<paragraph pageId="0" pageNumber="115">Diagnostic description.</paragraph>
<paragraph pageId="0" pageNumber="115">
Overall dark brown to black, except abdominal terga and sterna (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19B</figureCitation>
), which are dark brown. Pronotal disc (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19F</figureCitation>
) entirely black, pronotal expansions dark brown to black, often with anterior corners slightly lighter. Elytron (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19A</figureCitation>
) dark brown to black, with a pale-yellow longitudinal stripe about 4/5
<normalizedToken originalValue="×">x</normalizedToken>
as long as elytra. Legs dark brown to black. Sternum VIII (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19G</figureCitation>
) entirely dark brown, without lateral vitreous spots. Pygidium (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19H</figureCitation>
) entirely dark brown.
</paragraph>
<caption doi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="115">
<emphasis bold="true" pageId="0" pageNumber="115">Figure 19.</emphasis>
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" authorityYear="2021" baseAuthorityName="Olivier" baseAuthorityYear="1899" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys pauper" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="pauper">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys pauper</emphasis>
</taxonomicName>
(Olivier) 
<emphasis bold="true" pageId="0" pageNumber="115">comb. nov. A</emphasis>
, Male habitus, dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="115">B</emphasis>
, Male habitus, ventral view; 
<emphasis bold="true" pageId="0" pageNumber="115">C</emphasis>
, Female habitus, dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="115">D</emphasis>
, Female habitus, ventral view; 
<emphasis bold="true" pageId="0" pageNumber="115">E</emphasis>
, Male antenna, lateral view; 
<emphasis bold="true" pageId="0" pageNumber="115">F</emphasis>
, Male pronotum, dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="115">G</emphasis>
, Sternum IX, ventral view; 
<emphasis bold="true" pageId="0" pageNumber="115">H</emphasis>
, Pygidium, dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="115">I</emphasis>
, Syntergite, dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="115">J</emphasis>
, Sternum IX, Ventral view; 
<emphasis bold="true" pageId="0" pageNumber="115">K</emphasis>
, Aedeagus, dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="115">L</emphasis>
, Aedeagus, lateral view; 
<emphasis bold="true" pageId="0" pageNumber="115">M</emphasis>
, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="115">
Pronotum (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19F</figureCitation>
) with sides rounded, divergent posteriorly. Sternum VI without lanterns. 
<emphasis bold="true" pageId="0" pageNumber="115">Male.</emphasis>
Total length = 8.9-11.3 mm (aver. 10.1 mm); Pronotal length = 2.3-2.7 mm (aver. 2.5 mm); Pronotal width = 1.8-3.3 mm (aver. 2.55 mm); Elytral length = 7.3-8.8 mm (aver. 8.05 mm); Elytral width = 1.8-2.2 mm (aver. 2.0 mm). Antennomere III (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19E</figureCitation>
) with lamella 3
<normalizedToken originalValue="×">x</normalizedToken>
longer than core antennomere. Sternum VIII with posterior margin mucronate. Pygidium (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19H</figureCitation>
) with sides rounded, posterior corners barely visible, posterior margin rounded to almost straight. Syntergite (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19I,J</figureCitation>
) boomerang-shaped (with anterior margin strongly curved). Phallus (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19K-M</figureCitation>
) with dorsal plate curved dorsally in lateral view. Paramere apex almost straight, ventral projection (lateral view) moderately developed, almost right-angled. Paramere apex curved ventrally, ventral projection (lateral view) moderately developed, rounded to almost right-angled. 
<emphasis bold="true" pageId="0" pageNumber="115">Female</emphasis>
(Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19C, D</figureCitation>
). Total length = 11.0-11.3 mm (aver. 11.15 mm); Pronotal length = 1.7-1.8 mm (aver. 1.75 mm); Pronotal width = 2.7-2.8 mm (aver. 2.75 mm); Elytral length = 7.2-8.7 mm (aver. 7.95 mm); Elytral width = 2.0-2.5 mm (aver. 2.25 mm). Antennomeres (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">19C, D</figureCitation>
) III-X with apical corners almost right-angled, not projected and pointed.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="remarks">
<paragraph pageId="0" pageNumber="115">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="115">
<bibRefCitation author="Olivier, E" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="115" pagination="86 - 93" refId="B43" refString="Olivier, E, 1899. Contribution a l'etude des Lampyrides [Col.], descriptions et observations. Bulletin de la Societe Entomologique de France 4: 86 - 93" title="Contribution a l'etude des Lampyrides [Col.], descriptions et observations." volume="4" year="1899">Olivier (1899)</bibRefCitation>
described 
<taxonomicName authorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" authorityYear="2021" baseAuthorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" baseAuthorityYear="2021" class="Insecta" family="Lampyridae" genus="Cladodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladodes pauper" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="pauper">
<emphasis italics="true" pageId="0" pageNumber="115">Cladodes pauper</emphasis>
</taxonomicName>
from Brazil - the syntype we studied is from 
<normalizedToken originalValue="Teresópolis">Teresopolis</normalizedToken>
, Rio de Janeiro, Brazil. In addition to 
<normalizedToken originalValue="Teresópolis">Teresopolis</normalizedToken>
, in the 
<emphasis italics="true" pageId="0" pageNumber="115">
Serra dos 
<normalizedToken originalValue="Órgãos">Orgaos</normalizedToken>
</emphasis>
mountain range, we found previously unreported populations at the 
<emphasis italics="true" pageId="0" pageNumber="115">Serra da Mantiqueira</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="115">
<normalizedToken originalValue="Maciço">Macico</normalizedToken>
da Pedra Branca
</emphasis>
mountain ranges, in the states of Rio de Janeiro and Minas Gerais. Our study provides detailed geographic information for the first time, and expands considerably the known range of the species. After our phylogenetic analysis (see section 3.2), we transfer 
<taxonomicName authorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" authorityYear="2021" baseAuthorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" baseAuthorityYear="2021" class="Insecta" family="Lampyridae" genus="Cladodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladodes pauper" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="pauper">
<emphasis italics="true" pageId="0" pageNumber="115">Cladodes pauper</emphasis>
</taxonomicName>
to 
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">gen. nov.</emphasis>
, therefore 
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" authorityYear="2021" baseAuthorityName="Olivier" baseAuthorityYear="1899" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys pauper" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="pauper">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys pauper</emphasis>
</taxonomicName>
(Olivier, 1899). 
<bibRefCitation author="Olivier, E" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="115" pagination="86 - 93" refId="B43" refString="Olivier, E, 1899. Contribution a l'etude des Lampyrides [Col.], descriptions et observations. Bulletin de la Societe Entomologique de France 4: 86 - 93" title="Contribution a l'etude des Lampyrides [Col.], descriptions et observations." volume="4" year="1899">Olivier (1899)</bibRefCitation>
<normalizedToken originalValue="didn’t">didn't</normalizedToken>
provide an etymology for 
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" authorityYear="2021" baseAuthorityName="Olivier" baseAuthorityYear="1899" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys pauper" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="pauper">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys pauper</emphasis>
</taxonomicName>
, but we assume it was based on the Latin word for 
<normalizedToken originalValue="“poor”">&quot;poor&quot;</normalizedToken>
. Because 
<bibRefCitation author="Olivier, E" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="115" pagination="86 - 93" refId="B43" refString="Olivier, E, 1899. Contribution a l'etude des Lampyrides [Col.], descriptions et observations. Bulletin de la Societe Entomologique de France 4: 86 - 93" title="Contribution a l'etude des Lampyrides [Col.], descriptions et observations." volume="4" year="1899">Olivier (1899)</bibRefCitation>
originally placed it in 
<taxonomicName class="Insecta" family="Lampyridae" genus="Cladodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladodes" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Cladodes</emphasis>
</taxonomicName>
Solier, 1849, 
<taxonomicName authorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" authorityYear="2021" baseAuthorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" baseAuthorityYear="2021" class="Insecta" family="Lampyridae" genus="Cladodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladodes pauper" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="pauper">
<emphasis italics="true" pageId="0" pageNumber="115">Cladodes pauper</emphasis>
</taxonomicName>
would be relatively smaller than the remaining species in this genus, therefore 
<normalizedToken originalValue="“poor”">&quot;poor&quot;</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="da Silveira &amp; Roza &amp; Vaz &amp; Mermudes" authorityYear="2021" baseAuthorityName="Olivier" baseAuthorityYear="1899" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys pauper" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="pauper">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys pauper</emphasis>
</taxonomicName>
(Olivier, 1899) is unique among 
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
spp. by the pronotum entirely dark brown to black and sternum VI without lantern, in both sexes. 
<taxonomicName genus="C." lsidName="C. pauper" pageId="0" pageNumber="115" rank="species" species="pauper">
<emphasis italics="true" pageId="0" pageNumber="115">C. pauper</emphasis>
</taxonomicName>
is also unique in being restricted to relatively cooler climates, either at sites of relatively higher elevation (&gt;1200m) during the Austral Spring season, or at lower elevations (~600m), during Austral Winter. These observations suggest that this species might be adapted to climates substantially cooler than its congenerics, which deserves further scrutiny.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="types">
<paragraph pageId="0" pageNumber="115">Types.</paragraph>
<paragraph pageId="0" pageNumber="115">
<emphasis italics="true" pageId="0" pageNumber="115">Syntype</emphasis>
: BRAZIL: 1 male (MNHN, col. E. Olivier). Label is written 
<normalizedToken originalValue="“Brésil">&quot;Bresil</normalizedToken>
<normalizedToken originalValue="intér.”">inter.&quot;</normalizedToken>
, which we interpret as 
<normalizedToken originalValue="“interior”">&quot;interior&quot;</normalizedToken>
, &quot;country side&quot; of BRAZIL.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="materials_examined">
<paragraph pageId="0" pageNumber="115">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="115">
<materialsCitation ID-GBIF-Occurrence="3111796342" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="BRAZIL" location="Minas Gerais" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais">
<collectingCountry name="Brazil">BRAZIL</collectingCountry>
: 
<collectingRegion country="Brazil" name="Minas Gerais">Minas Gerais</collectingRegion>
: 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, 
<materialsCitation ID-GBIF-Occurrence="3111796313" collectingDate="2010-11-25" collectingDateMax="2010-11-29" collectingDateMin="2010-11-25" collectorName="Carvalho" country="Brazil" county="Itamonte" location="Maringa" municipality="Travessia de Itatiaia" specimenCount="1" specimenCount-male="1">
<collectingCounty>Itamonte</collectingCounty>
, 
<collectingMunicipality>Travessia de Itatiaia</collectingMunicipality>
para 
<normalizedToken originalValue="Maringá">
<location LSID="urn:lsid:plazi:treatment:7A66CDD6376A5D02B26DDF7AEB21325C:A3EED6132A059FB17EAB7EFBE61FAD73" country="Brazil" county="Itamonte" municipality="Travessia de Itatiaia" name="Maringa">Maringa</location>
</normalizedToken>
, 
<collectingDate value="2010-11-25" valueMax="2010-11-29" valueMin="2010-11-25">25-29.XI.2010</collectingDate>
, 
<collectorName>Carvalho</collectorName>
col. (DZRJ); 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, 
<materialsCitation ID-GBIF-Occurrence="3111796324" collectingDate="2009-11-19" collectingDateMax="2009-11-21" collectingDateMin="2009-11-19" collectorName="L. Silveira" country="Brazil" location="Itamonte" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:7A66CDD6376A5D02B26DDF7AEB21325C:E440ACDE76652EEB3CD0D27955203216" country="Brazil" name="Itamonte">Itamonte</location>
, gramado [on grass], 
<collectingDate value="2009-11-19" valueMax="2009-11-21" valueMin="2009-11-19">19-21.XI.2009</collectingDate>
, active search [afternoon], 
<collectorName>L. Silveira</collectorName>
col. (DZRJ)
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3111796335" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Brazil" location="Rio de Janeiro" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rio de Janeiro">
<collectingRegion country="Brazil" name="Rio de Janeiro">Rio de Janeiro</collectingRegion>
: 
<specimenCount type="male">1♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, 
<materialsCitation ID-GBIF-Occurrence="3111796338" collectingDate="2013-12" collectingMethod="Malaise trap" collectorName="Itatiaia, P. N, R. Monteiro" country="Brazil" elevation="1846" latitude="-22.416945" location="Itatiaia" longLatPrecision="1" longitude="-44.642475" specimenCount="1" specimenCount-male="1">
<collectorName>Itatiaia, P. N</collectorName>
. 
<location LSID="urn:lsid:plazi:treatment:7A66CDD6376A5D02B26DDF7AEB21325C:493F5EFC918E0FE8A0F478133BEC8101" country="Brazil" latitude="-22.416945" longLatPrecision="1" longitude="-44.642475" name="Itatiaia">Itatiaia</location>
, P5 (
<geoCoordinate degrees="22" direction="south" minutes="25" orientation="latitude" precision="1" seconds="01,0" value="-22.416945">
22°25
<normalizedToken originalValue="’01,0”">'01,0&quot;</normalizedToken>
S
</geoCoordinate>
, 
<geoCoordinate degrees="44" direction="west" minutes="38" orientation="longitude" precision="1" seconds="32,9" value="-44.642475">
44°38
<normalizedToken originalValue="’32,9”">'32,9&quot;</normalizedToken>
W
</geoCoordinate>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.846" unit="m" value="1846.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.846" unit="m" value="1846.0">1846 m</elevation>
</quantity>
), 
<collectingMethod>Malaise trap</collectingMethod>
, 
<collectingDate value="2013-12">XII.2013</collectingDate>
, 
<collectorName>R. Monteiro</collectorName>
col. (DZRJ); 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, 
<materialsCitation ID-GBIF-Occurrence="3111796304" collectingDate="2013-11-21" collectorName="P. N. Itatiaia, L. Silveira" country="Brazil" elevation="2200" location="Travessia Rui Braga" specimenCount="1" specimenCount-male="1">
<collectorName>P. N. Itatiaia</collectorName>
, 
<location LSID="urn:lsid:plazi:treatment:7A66CDD6376A5D02B26DDF7AEB21325C:1CE347A04CD32BEC84CB755D4BB82584" country="Brazil" name="Travessia Rui Braga">Travessia Rui Braga</location>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.2" unit="m" value="2200.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.2" unit="m" value="2200.0">2200 m</elevation>
</quantity>
, 
<collectingDate value="2013-11-21">21.XI.2013</collectingDate>
, 
<collectorName>L. Silveira</collectorName>
col. (DZRJ); 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, 
<materialsCitation ID-GBIF-Occurrence="3111796323" collectingDate="2013-11-21" collectingMethod="Malaise trap" collectorName="R. Monteiro" country="Brazil" elevation="2255" latitude="-22.394138" location="Itatiaia" longLatPrecision="1" longitude="-44.666584" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
P.N. 
<location LSID="urn:lsid:plazi:treatment:7A66CDD6376A5D02B26DDF7AEB21325C:4D901220233C4AD7A87263BA156191AC" country="Brazil" latitude="-22.394138" longLatPrecision="1" longitude="-44.666584" name="Itatiaia">Itatiaia</location>
, P7 (
<geoCoordinate degrees="22" direction="south" minutes="23" orientation="latitude" precision="1" seconds="38,9" value="-22.394138">
22°23
<normalizedToken originalValue="’38,9”">'38,9&quot;</normalizedToken>
S
</geoCoordinate>
, 
<geoCoordinate degrees="44" direction="west" minutes="39" orientation="longitude" precision="1" seconds="59,7" value="-44.666584">
44°39
<normalizedToken originalValue="’59,7”">'59,7&quot;</normalizedToken>
W
</geoCoordinate>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.255" unit="m" value="2255.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.255" unit="m" value="2255.0">2255 m</elevation>
</quantity>
), 
<collectingMethod>Malaise trap</collectingMethod>
, 
<collectingDate value="2013-11-21">21.XI.2013</collectingDate>
, 
<collectorName>R. Monteiro</collectorName>
col. (DZRJ); 
<specimenCount type="male">2♂</specimenCount>
, 
<specimenCount type="female">3♀</specimenCount>
</materialsCitation>
, 
<materialsCitation ID-GBIF-Occurrence="3111796310" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectingMethod="Malaise trap" collectorName="R. Monteiro" country="Brazil" elevation="1846" latitude="-22.416945" location="Itatiaia" longLatPrecision="1" longitude="-44.642475" specimenCount="1" specimenCount-male="1">
P.N. 
<location LSID="urn:lsid:plazi:treatment:7A66CDD6376A5D02B26DDF7AEB21325C:D21E4923EB74F3508317DD4ACB9E3FE9" country="Brazil" latitude="-22.416945" longLatPrecision="1" longitude="-44.642475" name="Itatiaia">Itatiaia</location>
, P5 (
<geoCoordinate degrees="22" direction="south" minutes="25" orientation="latitude" precision="1" seconds="01,0" value="-22.416945">
22°25
<normalizedToken originalValue="’01,0”">'01,0&quot;</normalizedToken>
S
</geoCoordinate>
, 
<geoCoordinate degrees="44" direction="west" minutes="38" orientation="longitude" precision="1" seconds="32,9" value="-44.642475">
44°38
<normalizedToken originalValue="’32,9”">'32,9&quot;</normalizedToken>
W
</geoCoordinate>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.846" unit="m" value="1846.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.846" unit="m" value="1846.0">1846 m</elevation>
</quantity>
), 
<collectingMethod>Malaise trap</collectingMethod>
, 
<collectorName>R. Monteiro</collectorName>
col (DZRJ); 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, 
<materialsCitation ID-GBIF-Occurrence="3111796311" collectingDate="2018-07-07" collectorName="P. E. Pedra Branca, A. L. D. Ferreira" country="Brazil" county="Taquara" elevation="400" location="Crepusculo" municipality="Trilha da Padaria" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rio de Janeiro">
<collectingRegion country="Brazil" name="Rio de Janeiro">Rio de Janeiro</collectingRegion>
, 
<collectorName>P.E. Pedra Branca</collectorName>
, 
<collectingCounty>Taquara</collectingCounty>
, 
<collectingMunicipality>Trilha da Padaria</collectingMunicipality>
(~
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400m</elevation>
</quantity>
), 
<normalizedToken originalValue="Crepúsculo">
<location LSID="urn:lsid:plazi:treatment:7A66CDD6376A5D02B26DDF7AEB21325C:9049F92CE2BFB208EAE1ECC8F0623328" country="Brazil" county="Taquara" municipality="Trilha da Padaria" name="Crepusculo" stateProvince="Rio de Janeiro">Crepusculo</location>
</normalizedToken>
, PiT-LED, 
<collectingDate value="2018-07-07">07.VII.2018</collectingDate>
, 
<collectorName>A. L. D. Ferreira</collectorName>
(DZRJ); 
<specimenCount type="male">1♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, 
<materialsCitation ID-GBIF-Occurrence="3111796319" collectingDate="2015-11-14" collectingDateMax="2015-11-15" collectingDateMin="2015-11-14" collectorName="Teresopolis, P. N, L. Silveira" country="Brazil" county="Serra" elevation="1775" location="Trilha do Sino" municipality="Orgaos" specimenCount="1">
<collectorName>
<normalizedToken originalValue="Teresópolis">Teresopolis</normalizedToken>
, P.N
</collectorName>
. 
<collectingCounty>Serra</collectingCounty>
dos 
<normalizedToken originalValue="Órgãos">
<collectingMunicipality>Orgaos</collectingMunicipality>
</normalizedToken>
, 
<location LSID="urn:lsid:plazi:treatment:7A66CDD6376A5D02B26DDF7AEB21325C:2982DF9506D3031141A667CE476ACC90" country="Brazil" county="Serra" municipality="Orgaos" name="Trilha do Sino">Trilha do Sino</location>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.775" metricValueMax="1.9" metricValueMin="1.65" unit="m" value="1775.0" valueMax="1900.0" valueMin="1650.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.775" metricValueMax="1.9" metricValueMin="1.65" unit="m" value="1775.0" valueMax="1900.0" valueMin="1650.0">1650-1900 m</elevation>
</quantity>
, 
<collectingDate value="2015-11-14" valueMax="2015-11-15" valueMin="2015-11-14">14-15.XI.2015</collectingDate>
, 
<collectorName>L. Silveira</collectorName>
col. (DZRJ)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>