189 lines
20 KiB
XML
189 lines
20 KiB
XML
<document id="4046A0B99A6B4CC529BCBA28825821B0" ID-CLB-Dataset="20903" ID-DOI="10.11646/zootaxa.4980.1.9" ID-GBIF-Dataset="c6aedffd-0196-45d9-acc3-071c4e886e66" ID-ISSN="1175-5326" ID-Zenodo-Dep="4883181" ID-ZooBank="E4D00067-F4F3-4BC5-9CDA-6532773B56B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622464354611" checkinUser="plazi" docAuthor="Makarkin, Vladimir N., Archibald, S. Bruce & Mathewes, Rolf W." docDate="2021" docId="03F07D0C043DFFF3F09578B2FCB2059E" docLanguage="en" docName="zootaxa.4980.1.9.pdf" docOrigin="Zootaxa 4980 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Osmylidia taliae Makarkin & Archibald & Mathewes 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="148" masterDocId="FFC905740438FFF5F0027825FF9D0753" masterDocTitle="New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks" masterLastPageNumber="156" masterPageNumber="142" pageNumber="147" updateTime="1698995111136" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="96F904CE352B6D4A697960C4CAD6774D">
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<mods:title id="7177944526B63027617581B6468DE7FA">New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks</mods:title>
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<mods:name id="A401511C9565435673999ADE9DE25A1E" type="personal">
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<mods:namePart id="E75F35A4CD8AC0F4364E3B9F5477D707">Makarkin, Vladimir N.</mods:namePart>
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<mods:nameIdentifier id="BBF317AF4A975690BEC2581392347CC6" type="ORCID">0000-0002-1304-0461</mods:nameIdentifier>
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<mods:affiliation id="3EB3031C63E3D9A24FAACA251C472A5B">Federal Scientific Centre of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok 690022, Russia.</mods:affiliation>
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<mods:nameIdentifier id="3AE762B8622177600209C761991E3F0A" type="email">vnmakarkin@mail.ru</mods:nameIdentifier>
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<mods:namePart id="4364F6C0A81AF1E7C32C664FB1AB31F8">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="F58D4075610BCCA7F60A051C4C9FADF1">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, BC, V 5 A 1 S 6, Canada. & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America; and Royal British Columbia Museum, 675 Belleville Street, Victoria, BC, V 8 W 9 W 2, Canada.</mods:affiliation>
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<mods:namePart id="D7E683E66165CD06EE961D474AFC8F77">Mathewes, Rolf W.</mods:namePart>
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<mods:affiliation id="FFFFAC0FA44AF3CC3A40DDC7F6837064">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, BC, V 5 A 1 S 6, Canada.</mods:affiliation>
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<mods:title id="4969B80F2F214AF3AEC51A16816EC659">Zootaxa</mods:title>
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<mods:part id="093232C32498F4117D8C771838259060">
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<mods:date id="B994CB14B83393EAA4FA4FD80931CD00">2021</mods:date>
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<treatment id="03F07D0C043DFFF3F09578B2FCB2059E" ID-DOI="http://doi.org/10.5281/zenodo.4883201" ID-GBIF-Taxon="192720196" ID-Zenodo-Dep="4883201" LSID="urn:lsid:plazi:treatment:03F07D0C043DFFF3F09578B2FCB2059E" httpUri="http://treatment.plazi.org/id/03F07D0C043DFFF3F09578B2FCB2059E" lastPageId="6" lastPageNumber="148" pageId="5" pageNumber="147">
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<subSubSection id="C3439F91043DFFF0F09578B2FE5D07E2" box="[151,448,151,178]" pageId="5" pageNumber="147" type="nomenclature">
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<paragraph id="8BE6CC1A043DFFF0F09578B2FE5D07E2" blockId="5.[151,448,151,213]" box="[151,448,151,178]" pageId="5" pageNumber="147">
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<heading id="D0AE7B76043DFFF0F09578B2FE5D07E2" bold="true" box="[151,448,151,178]" fontSize="11" level="1" pageId="5" pageNumber="147" reason="1">
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<emphasis id="B92D1008043DFFF0F09578B2FE5D07E2" bold="true" box="[151,448,151,178]" pageId="5" pageNumber="147">
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<taxonomicName id="4C59B799043DFFF0F09578B2FEC107E1" authority="Makarkin & Archibald & Mathewes, 2021" authorityName="Makarkin & Archibald & Mathewes" authorityYear="2021" box="[151,348,151,178]" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="147" phylum="Arthropoda" rank="species" species="taliae" status="sp. nov.">
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<emphasis id="B92D1008043DFFF0F09578B2FEC107E1" bold="true" box="[151,348,151,178]" italics="true" pageId="5" pageNumber="147">Osmylidia taliae</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A21EAD73043DFFF0F16178B2FE5D07E2" box="[355,448,151,177]" pageId="5" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3439F91043DFFF0F095789EFF440786" box="[151,217,187,213]" pageId="5" pageNumber="147" type="description">
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<paragraph id="8BE6CC1A043DFFF0F095789EFF440786" blockId="5.[151,448,151,213]" box="[151,217,187,213]" pageId="5" pageNumber="147">
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<figureCitation id="1362D09F043DFFF0F095789EFF440786" box="[151,217,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1867,1891]" captionTargetBox="[151,1436,633,1835]" captionTargetId="figure-177@5.[151,1436,633,1843]" captionTargetPageId="5" captionText="FIGURE 3. Osmylidia taliae sp. nov., holotype UCM 67996 from the Green River Formation. A, photograph of the part; B, drawing of forewing venation; C, drawing of hind wing venation. Posterior branch of MA in red. Scale bar = 5 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4883189" httpUri="https://zenodo.org/record/4883189/files/figure.png" pageId="5" pageNumber="147">Fig. 3</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3439F91043DFFF0F0957926FB3206D9" pageId="5" pageNumber="147" type="materials_examined">
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<paragraph id="8BE6CC1A043DFFF0F0957926FC4F0612" blockId="5.[151,1436,259,610]" pageId="5" pageNumber="147">
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<emphasis id="B92D1008043DFFF0F0957926FED8064E" bold="true" box="[151,325,259,285]" pageId="5" pageNumber="147">Type material.</emphasis>
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<materialsCitation id="3B31C647043DFFF0F1497926FC4F0612" ID-GBIF-Occurrence="3459552304" accessionNumber="UCM 67996" collectingDate="2004-03-04" collectorName="David Kohls" pageId="5" pageNumber="147" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54E272B8043DFFF0F1497926FE2E064E" box="[331,435,259,285]" pageId="5" pageNumber="147" type="holotype">Holotype</typeStatus>
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UCM 67996, collected by
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<collectorName id="26ACA9CC043DFFF0F2E67926FCE8064E" box="[740,885,259,285]" pageId="5" pageNumber="147">David Kohls</collectorName>
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,
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<date id="FFE7EADA043DFFF0F3827926FBBD064E" box="[896,1056,259,285]" pageId="5" pageNumber="147" value="2004-03-04">
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<collectingDate id="EFA31332043DFFF0F3827926FBBD064E" box="[896,1056,259,285]" pageId="5" pageNumber="147" value="2004-03-04">4 March 2004</collectingDate>
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</date>
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; deposited in UCM. A poorly preserved incomplete specimen with at least small portions of all four wings.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE6CC1A043DFFF0F0C5796EFB3206D9" blockId="5.[151,1436,259,610]" pageId="5" pageNumber="147">
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<materialsCitation id="3B31C647043DFFF0F0C5796EFB3106D9" ID-GBIF-Occurrence="3459552301" country="United States of America" county="Garfield County" location="Parachute Creek Member" municipality="Ypresian" pageId="5" pageNumber="147" specimenCount="1" stateProvince="Colorado" typeStatus="holotype">
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<emphasis id="B92D1008043DFFF0F0C5796EFD970636" bold="true" box="[199,522,331,357]" pageId="5" pageNumber="147">
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<typeStatus id="54E272B8043DFFF0F0C5796EFE9F0636" box="[199,258,331,357]" pageId="5" pageNumber="147">Type</typeStatus>
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locality and horizon.
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</emphasis>
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UCM locality 2005025 (Anvil Points Kohls site);
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<collectingCounty id="6287B496043DFFF0F460796EFA830636" box="[1122,1310,331,357]" pageId="5" pageNumber="147">Garfield County</collectingCounty>
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,
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<collectingRegion id="499D02F8043DFFF0F52F796EFA050636" box="[1325,1432,331,357]" country="United States of America" name="Colorado" pageId="5" pageNumber="147">Colorado</collectingRegion>
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,
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||
<collectingCountry id="F34E8C8A043DFFF0F095794AFF7906DA" box="[151,228,367,393]" name="United States of America" pageId="5" pageNumber="147">U.S.A.</collectingCountry>
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<location id="8E869AC1043DFFF0F0E9794AFD9406DA" LSID="urn:lsid:plazi:treatment:03F07D0C043DFFF3F09578B2FCB2059E:8E869AC1043DFFF0F0E9794AFD9406DA" box="[235,521,367,393]" country="United States of America" county="Garfield County" municipality="Ypresian" name="Parachute Creek Member" pageId="5" pageNumber="147" stateProvince="Colorado">
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<location id="8E869AC1043DFFF0F0E9794AFE3906DA" LSID="urn:lsid:plazi:treatment:03F07D0C043DFFF3F09578B2FCB2059E:8E869AC1043DFFF0F0E9794AFE3906DA" box="[235,420,367,393]" country="United States of America" county="Garfield County" municipality="Ypresian" name="Parachute Creek" pageId="5" pageNumber="147" stateProvince="Colorado">Parachute Creek</location>
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Member
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</location>
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of the
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<location id="8E869AC1043DFFF0F25A794AFCFD06DA" LSID="urn:lsid:plazi:treatment:03F07D0C043DFFF3F09578B2FCB2059E:8E869AC1043DFFF0F25A794AFCFD06DA" box="[600,864,367,393]" country="United States of America" county="Garfield County" municipality="Ypresian" name="Green River Formation" pageId="5" pageNumber="147" stateProvince="Colorado">
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<location id="8E869AC1043DFFF0F25A794AFD7C06DA" LSID="urn:lsid:plazi:treatment:03F07D0C043DFFF3F09578B2FCB2059E:8E869AC1043DFFF0F25A794AFD7C06DA" box="[600,737,367,393]" country="United States of America" county="Garfield County" municipality="Ypresian" name="Green River" pageId="5" pageNumber="147" stateProvince="Colorado">
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Green
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<location id="8E869AC1043DFFF0F2A1794AFD7C06DA" LSID="urn:lsid:plazi:treatment:03F07D0C043DFFF3F09578B2FCB2059E:8E869AC1043DFFF0F2A1794AFD7C06DA" box="[675,737,367,393]" country="United States of America" county="Garfield County" municipality="Ypresian" name="River Formation" pageId="5" pageNumber="147" stateProvince="Colorado">River</location>
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</location>
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Formation
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</location>
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; late
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<collectingMunicipality id="6B825660043DFFF0F398794AFC6006DA" box="[922,1021,367,393]" pageId="5" pageNumber="147">Ypresian</collectingMunicipality>
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(early Eocene)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3439F91043DFFF0F0C579B6FAE506FE" box="[199,1400,403,430]" pageId="5" pageNumber="147" type="etymology">
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<paragraph id="8BE6CC1A043DFFF0F0C579B6FAE506FE" blockId="5.[151,1436,259,610]" box="[199,1400,403,430]" pageId="5" pageNumber="147">
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||
<emphasis id="B92D1008043DFFF0F0C579B6FED506FE" bold="true" box="[199,328,403,429]" pageId="5" pageNumber="147">Etymology</emphasis>
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. From the first name of Talia Karim, for her long-standing helpful support of paleoentomology.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3439F91043DFFF0F0C57992FDFA056E" pageId="5" pageNumber="147" type="diagnosis">
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||
<paragraph id="8BE6CC1A043DFFF0F0C57992FDFA056E" blockId="5.[151,1436,259,610]" pageId="5" pageNumber="147">
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||
<emphasis id="B92D1008043DFFF0F0C57992FEA70682" bold="true" box="[199,314,439,465]" pageId="5" pageNumber="147">Diagnosis</emphasis>
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. May be distinguished from other species of the genus by forewing stem of RP distally pectinately branched [other species: dichotomously branched]; MP not branched at outer gradate series [other species: branched]; hind wing posterior branch of MA fused with MP for short distance [other species (except some specimens of
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<taxonomicName id="4C59B799043DFFF0F5837A25FF52056E" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="147" phylum="Arthropoda" rank="species" species="picta">
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<emphasis id="B92D1008043DFFF0F5837A25FF52056E" italics="true" pageId="5" pageNumber="147">O. picta</emphasis>
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</taxonomicName>
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): separate, connected by crossvein].
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3439F91043DFFF3F0C57A62FCB2059E" lastPageId="6" lastPageNumber="148" pageId="5" pageNumber="147" type="description">
|
||
<paragraph id="8BE6CC1A043DFFF0F0C57A62FBBC0532" blockId="5.[151,1436,259,610]" box="[199,1057,583,610]" pageId="5" pageNumber="147">
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||
<emphasis id="B92D1008043DFFF0F0C57A62FECC0532" bold="true" box="[199,337,583,609]" pageId="5" pageNumber="147">Description</emphasis>
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. Body poorly preserved,
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<emphasis id="B92D1008043DFFF0F26C7A6DFD150532" box="[622,648,584,609]" italics="true" pageId="5" pageNumber="147">ca</emphasis>
|
||
.
|
||
<quantity id="4CA161FF043DFFF0F2957A62FD450531" box="[663,728,583,610]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="147" unit="mm" value="9.0">9 mm</quantity>
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long; details not discernable.
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</paragraph>
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||
<caption id="DF269C92043DFFF0F0957F6EFF4400F8" ID-DOI="http://doi.org/10.5281/zenodo.4883189" ID-Zenodo-Dep="4883189" httpUri="https://zenodo.org/record/4883189/files/figure.png" pageId="5" pageNumber="147" startId="5.[151,250,1867,1891]" targetBox="[151,1436,633,1835]" targetPageId="5">
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<paragraph id="8BE6CC1A043DFFF0F0957F6EFF4400F8" blockId="5.[151,1437,1867,1963]" pageId="5" pageNumber="147">
|
||
<emphasis id="B92D1008043DFFF0F0957F6EFE890030" bold="true" box="[151,276,1867,1891]" pageId="5" pageNumber="147">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C59B799043DFFF0F11E7F6EFE5B0030" authority="Makarkin & Archibald & Mathewes, 2021" authorityName="Makarkin & Archibald & Mathewes" authorityYear="2021" box="[284,454,1867,1891]" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="147" phylum="Arthropoda" rank="species" species="taliae" status="sp. nov.">
|
||
<emphasis id="B92D1008043DFFF0F11E7F6EFE5B0030" box="[284,454,1867,1891]" italics="true" pageId="5" pageNumber="147">Osmylidia taliae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92D1008043DFFF0F1CF7F6EFD820030" bold="true" box="[461,543,1867,1891]" pageId="5" pageNumber="147">
|
||
<taxonomicNameLabel id="A21EAD73043DFFF0F1CF7F6EFD820030" box="[461,543,1867,1891]" pageId="5" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
|
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, holotype UCM 67996 from the Green River Formation. A, photograph of the part; B, drawing of forewing venation; C, drawing of hind wing venation. Posterior branch of MA in red. Scale bar = 5 mm (all to scale).
|
||
</paragraph>
|
||
</caption>
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||
<paragraph id="8BE6CC1A043EFFF3F0C578B2FC720682" blockId="6.[151,1437,150,717]" pageId="6" pageNumber="148">
|
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Forewing
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<emphasis id="B92D1008043EFFF3F13978B2FEC807E3" box="[315,341,151,176]" italics="true" pageId="6" pageNumber="148">ca</emphasis>
|
||
.
|
||
<quantity id="4CA161FF043EFFF3F16778B2FE2B07E2" box="[357,438,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="148" unit="mm" value="14.0">14 mm</quantity>
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||
long,
|
||
<emphasis id="B92D1008043EFFF3F20278B2FD8707E3" box="[512,538,151,176]" italics="true" pageId="6" pageNumber="148">ca</emphasis>
|
||
. 4.8–5.0 mm wide. Costal space broad, with subcostal veinlets simple, closely spaced. Sc fused with RA far from wing apex. Subcostal space narrow in distal portion; crossveins not detected. RA space relatively broad medially, with two preserved crossveins. RP with 11 pectinate branches, all shallowly forked. All preserved crossveins in radial to medio-cubital spaces arranged in three gradate series: proximal series with three preserved crossveins; intermediate series with eight preserved crossveins; outer series with 11 preserved crossveins (probably
|
||
<quantity id="4CA161FF043EFFF3F187796EFE230636" box="[389,446,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="6" pageNumber="148" unit="in" value="14.0">14 in</quantity>
|
||
life). MA pectinately branched distally, probably with three branches. MP probably shallowly branched (apex not preserved), at least not branched at outer gradate series or proximad it. CuA pectinately branched, with three preserved branches. Wing colouration: faint spots at pterostigma and along apical margin; some crossveins in gradate series appear surrounded by fuscous colouration.
|
||
</paragraph>
|
||
<paragraph id="8BE6CC1A043EFFF3F0C579FEFCB2059E" blockId="6.[151,1437,150,717]" pageId="6" pageNumber="148">
|
||
Hind wing
|
||
<quantity id="4CA161FF043EFFF3F14679FFFE0406A6" box="[324,409,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="148" unit="mm" value="7.5">7.5 mm</quantity>
|
||
long as preserved (estimated complete length
|
||
<emphasis id="B92D1008043EFFF3F39979FEFC2806A7" box="[923,949,475,500]" italics="true" pageId="6" pageNumber="148">ca</emphasis>
|
||
.
|
||
<quantity id="4CA161FF043EFFF3F3C179FEFB8806A6" box="[963,1045,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="148" unit="mm" value="13.0">13 mm</quantity>
|
||
),
|
||
<emphasis id="B92D1008043EFFF3F42A79FEFBDF06A7" box="[1064,1090,475,500]" italics="true" pageId="6" pageNumber="148">ca</emphasis>
|
||
.
|
||
<quantity id="4CA161FF043EFFF3F44D79FEFB1206A6" box="[1103,1167,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="148" unit="mm" value="3.0">3 mm</quantity>
|
||
wide as preserved (estimated complete width
|
||
<emphasis id="B92D1008043EFFF3F19679DAFE33054B" box="[404,430,511,536]" italics="true" pageId="6" pageNumber="148">ca</emphasis>
|
||
.
|
||
<quantity id="4CA161FF043EFFF3F1B979DBFD89054A" box="[443,532,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="148" unit="mm" value="3.5">3.5 mm</quantity>
|
||
). Costal space moderately broad, with simple, relatively closely spaced preserved subcostal veinlets. Three closely spaced crossveins preserved in RA space medially. RP with five partially preserved branches. MA deeply forked, distally pectinately forked, with at least three branches. MP pectinately forked with two branches. CuA fragmentarily preserved. Crossveins in radial, radio-medial spaces arranged in two preserved gradate series: intermediate series with four preserved crossveins; outer series with six preserved crossveins. Two preserved crossveins between posterior branch of MA, MP.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |