291 lines
37 KiB
XML
291 lines
37 KiB
XML
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<mods:titleInfo id="1540702A8951545B189A580ECA78FC3C">
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<mods:title id="DCE817739E825804D2F04726E8883146">New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks</mods:title>
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<mods:namePart id="710DF36DD5939F0DF7B2B581B57CC83F">Makarkin, Vladimir N.</mods:namePart>
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<mods:nameIdentifier id="0DDB56A0DEB911310ECE89A9180B2675" type="ORCID">0000-0002-1304-0461</mods:nameIdentifier>
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<mods:affiliation id="452A46E1ADCFA5A049BE837903248909">Federal Scientific Centre of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok 690022, Russia.</mods:affiliation>
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<mods:nameIdentifier id="6ED2DD8B818770D72493C289914EE941" type="email">vnmakarkin@mail.ru</mods:nameIdentifier>
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<mods:namePart id="B54A935A3264A51065CF8949D08FF19E">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="90B82F5F484F9C7135F6D20AA61E75D2">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, BC, V 5 A 1 S 6, Canada. & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America; and Royal British Columbia Museum, 675 Belleville Street, Victoria, BC, V 8 W 9 W 2, Canada.</mods:affiliation>
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<mods:namePart id="B322B08F12C1127E8A625C77AE379C50">Mathewes, Rolf W.</mods:namePart>
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<mods:affiliation id="6BB869F1639D9540E71670D083EBB12E">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, BC, V 5 A 1 S 6, Canada.</mods:affiliation>
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<mods:title id="570C3DFA6852243905DA98284610EAB4">Zootaxa</mods:title>
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<mods:date id="A8F2E8B997149B18A362F79959F65C4D">2021</mods:date>
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<mods:number id="041E6EED1E506A99F246138BE3C2EAAA">2021-05-31</mods:number>
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<treatment id="03F07D0C0439FFF7F0957E5AFB9404D2" ID-DOI="http://doi.org/10.5281/zenodo.5915222" ID-GBIF-Taxon="192720199" ID-Zenodo-Dep="5915222" LSID="urn:lsid:plazi:treatment:03F07D0C0439FFF7F0957E5AFB9404D2" httpUri="http://treatment.plazi.org/id/03F07D0C0439FFF7F0957E5AFB9404D2" lastPageId="2" lastPageNumber="144" pageId="1" pageNumber="143">
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<subSubSection id="C3439F910439FFF4F0957E5AFDE201CA" box="[151,639,1663,1689]" pageId="1" pageNumber="143" type="nomenclature">
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<paragraph id="8BE6CC1A0439FFF4F0957E5AFDE201CA" blockId="1.[151,639,1663,1689]" box="[151,639,1663,1689]" pageId="1" pageNumber="143">
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<heading id="D0AE7B760439FFF4F0957E5AFDE201CA" bold="true" box="[151,639,1663,1689]" fontSize="11" level="1" pageId="1" pageNumber="143" reason="1">
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<emphasis id="B92D10080439FFF4F0957E5AFDE201CA" bold="true" box="[151,639,1663,1689]" pageId="1" pageNumber="143">
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Subfamily
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<taxonomicName id="4C59B7990439FFF4F11D7E5AFDE201CA" authority="Kruger, 1913" authorityName="Kruger" authorityYear="1913" box="[287,639,1663,1689]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="143" phylum="Arthropoda" rank="subFamily" subFamily="Protosmylinae">
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Protosmylinae
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<bibRefCitation id="EFC8B1EB0439FFF4F1D97E5AFDE201CA" author="Kruger, L." box="[475,639,1663,1689]" pageId="1" pageNumber="143" pagination="3 - 123" refId="ref7259" refString="Kruger, L. (1913) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913">Krüger, 1913</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3439F910439FFF7F0957EE2FB910531" lastPageId="2" lastPageNumber="144" pageId="1" pageNumber="143" type="type_taxon">
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<paragraph id="8BE6CC1A0439FFF4F0957EE2FC3201B2" blockId="1.[151,1436,1735,2014]" box="[151,943,1735,1762]" pageId="1" pageNumber="143">
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<emphasis id="B92D10080439FFF4F0957EE2FE8001B2" bold="true" box="[151,285,1735,1761]" pageId="1" pageNumber="143">
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<typeStatus id="54E272B80439FFF4F0957EE2FF4F01B2" box="[151,210,1735,1761]" pageId="1" pageNumber="143">Type</typeStatus>
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genus
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</emphasis>
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.
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<taxonomicName id="4C59B7990439FFF4F1297EE2FDCC01B2" ID-CoL="6XYL" authority="Kruger, 1913" authorityName="Kruger" authorityYear="1913" box="[299,593,1735,1762]" class="Insecta" family="Osmylidae" genus="Protosmylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="143" phylum="Arthropoda" rank="genus">
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<emphasis id="B92D10080439FFF4F1297EE2FE2B01B2" box="[299,438,1735,1761]" italics="true" pageId="1" pageNumber="143">Protosmylus</emphasis>
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<bibRefCitation id="EFC8B1EB0439FFF4F1BF7EEDFDCC01B2" author="Kruger, L." box="[445,593,1735,1762]" pageId="1" pageNumber="143" pagination="3 - 123" refId="ref7259" refString="Kruger, L. (1913) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913">Krüger, 1913</bibRefCitation>
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</taxonomicName>
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(=
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<taxonomicName id="4C59B7990439FFF4F2737EE2FC3F01B2" ID-CoL="63GPY" authority="Cockerell, 1908" authorityName="Cockerell" authorityYear="1908" box="[625,930,1735,1761]" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="143" phylum="Arthropoda" rank="genus">
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<emphasis id="B92D10080439FFF4F2737EE2FD7901B2" box="[625,740,1735,1761]" italics="true" pageId="1" pageNumber="143">Osmylidia</emphasis>
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<bibRefCitation id="EFC8B1EB0439FFF4F2EE7EE2FC3F01B2" author="Cockerell, T. D. A." box="[748,930,1735,1761]" pageId="1" pageNumber="143" pagination="341 - 342" refId="ref6760" refString="Cockerell, T. D. A. (1908) Fossil Osmylidae (Neuroptera) in America. The Canadian Entomologist, 40, 341 - 342. https: // doi. org / 10.4039 / Ent 40341 - 10" type="journal article" year="1908">Cockerell, 1908</bibRefCitation>
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</taxonomicName>
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).
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</paragraph>
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<paragraph id="8BE6CC1A0439FFF4F0C57ECEFDD6007A" blockId="1.[151,1436,1735,2014]" pageId="1" pageNumber="143">
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<emphasis id="B92D10080439FFF4F0C57ECEFDE50056" bold="true" box="[199,632,1771,1797]" pageId="1" pageNumber="143">Emended diagnosis (based on wings)</emphasis>
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. Wings may be distinguished from those of other subfamilies of
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<taxonomicName id="4C59B7990439FFF4F5427ECEFF23007A" ID-CoL="DRB" authorityName="Leach" authorityYear="1815" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="143" phylum="Arthropoda" rank="family">Osmylidae</taxonomicName>
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by a combination of the following.
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</paragraph>
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<paragraph id="8BE6CC1A0439FFF7F0C57F16FF7D064E" blockId="1.[151,1436,1735,2014]" lastBlockId="2.[151,1437,151,897]" lastPageId="2" lastPageNumber="144" pageId="1" pageNumber="143">
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Forewing: Subcostal veinlets almost always simple (very rarely some forked) [
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<taxonomicName id="4C59B7990439FFF4F45E7F16FB40001E" authorityName="Leach" authorityYear="1815" box="[1116,1245,1843,1869]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="143" phylum="Arthropoda" rank="subFamily" subFamily="Osmylinae">Osmylinae</taxonomicName>
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: mostly forked]; trichosors along almost entire wing margin [
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<taxonomicName id="4C59B7990439FFF4F28D7F72FC890022" authorityName="Navas" authorityYear="1912" box="[655,788,1879,1905]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="143" phylum="Arthropoda" rank="subFamily" subFamily="Gumillinae">Gumillinae</taxonomicName>
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: restricted to apical region]; one subcostal crossvein, positioned basally [Porisminae: numerous along entire subcostal space]; all or nearly all crossveins in radial space forming three to four gradate series (except some
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<taxonomicName id="4C59B7990439FFF4F2DA7F85FCB600EA" ID-CoL="5HVQ" authorityName="Navas" authorityYear="1911" box="[728,811,1952,1977]" class="Insecta" family="Osmylidae" genus="Lysmus" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="143" phylum="Arthropoda" rank="genus">
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<emphasis id="B92D10080439FFF4F2DA7F85FCB600EA" box="[728,811,1952,1977]" italics="true" pageId="1" pageNumber="143">Lysmus</emphasis>
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</taxonomicName>
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) [all or nearly all crossveins (
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<taxonomicName id="4C59B7990439FFF4F4857FBAFA9700EA" authorityName="Navas" authorityYear="1912" box="[1159,1290,1951,1977]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="143" phylum="Arthropoda" rank="subFamily" subFamily="Gumillinae">Gumillinae</taxonomicName>
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, Porisminae,
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<taxonomicName id="4C59B7990439FFF4F0957FE6FED7008D" authorityName="Esben-Petersen" authorityYear="1917" box="[151,330,1987,2014]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="143" phylum="Arthropoda" rank="subFamily" subFamily="Eidoporisminae">Eidoporisminae</taxonomicName>
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) or proximal crossveins (other subfamilies) irregularly arranged]; M forked at level or only slightly proximad termination of A1 [
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<taxonomicName id="4C59B799043AFFF7F1E378B2FDFD07E2" authorityName="Leach" authorityYear="1815" box="[481,608,151,177]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Osmylinae">Osmylinae</taxonomicName>
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,
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<taxonomicName id="4C59B799043AFFF7F26E78B2FC9D07E2" authorityName="Carpenter" authorityYear="1943" box="[620,768,151,177]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Kempyninae">Kempyninae</taxonomicName>
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: much distad;
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<taxonomicName id="4C59B799043AFFF7F3A178B2FBD607E2" authorityName="Kruger" authorityYear="1913" box="[931,1099,151,177]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Stenosmylinae">Stenosmylinae</taxonomicName>
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, Porisminae,
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<taxonomicName id="4C59B799043AFFF7F4E178B2FA0407E2" authorityName="Esben-Petersen" authorityYear="1917" box="[1251,1433,151,177]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Eidoporisminae">Eidoporisminae</taxonomicName>
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: much proximad]; between CuA, CuP ≤ 5 crossveins (up to
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<quantity id="4CA161FF043AFFF7F337789EFCFC0786" box="[821,865,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="2" pageNumber="144" unit="in" value="8.0">8 in</quantity>
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||
some
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<taxonomicName id="4C59B799043AFFF7F3A9789EFC630787" ID-CoL="5HVQ" authorityName="Navas" authorityYear="1911" box="[939,1022,187,212]" class="Insecta" family="Osmylidae" genus="Lysmus" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
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<emphasis id="B92D1008043AFFF7F3A9789EFC630787" box="[939,1022,187,212]" italics="true" pageId="2" pageNumber="144">Lysmus</emphasis>
|
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</taxonomicName>
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) [
|
||
<taxonomicName id="4C59B799043AFFF7F414789EFB230786" authorityName="Kruger" authorityYear="1913" box="[1046,1214,187,213]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
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||
: about 10 or more]; CuA pectinate, with 2–6 branches; CuP with 3–7 pectinate branches (up to
|
||
<quantity id="4CA161FF043AFFF7F40578FAFBA807AA" box="[1031,1077,223,249]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="2" pageNumber="144" unit="in" value="9.0">9 in</quantity>
|
||
some
|
||
<taxonomicName id="4C59B799043AFFF7F48678FAFB4A07AB" ID-CoL="5HVQ" authorityName="Navas" authorityYear="1911" box="[1156,1239,223,248]" class="Insecta" family="Osmylidae" genus="Lysmus" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F48678FAFB4A07AB" box="[1156,1239,223,248]" italics="true" pageId="2" pageNumber="144">Lysmus</emphasis>
|
||
</taxonomicName>
|
||
) [
|
||
<taxonomicName id="4C59B799043AFFF7F4F378FAFA0407AA" authorityName="Kruger" authorityYear="1913" box="[1265,1433,223,249]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
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: 8–15].
|
||
</paragraph>
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||
<paragraph id="8BE6CC1A043AFFF7F0C57902FDB906DA" blockId="2.[151,1437,151,897]" pageId="2" pageNumber="144">
|
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Hind wing: MP lacks anteriorly directed basal spur-like process [
|
||
<taxonomicName id="4C59B799043AFFF7F39E7902FBD90612" authorityName="Kruger" authorityYear="1913" box="[924,1092,295,321]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
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: present]; CuP relatively short, shallowly (not pectinately) branched or simple [
|
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<taxonomicName id="4C59B799043AFFF7F2BB796EFCD30636" authorityName="Carpenter" authorityYear="1943" box="[697,846,331,357]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Kempyninae">Kempyninae</taxonomicName>
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,
|
||
<taxonomicName id="4C59B799043AFFF7F359796EFC4A0636" authorityName="Leach" authorityYear="1815" box="[859,983,331,357]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Osmylinae">Osmylinae</taxonomicName>
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,
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<taxonomicName id="4C59B799043AFFF7F3E6796EFB110636" authorityName="Kruger" authorityYear="1913" box="[996,1164,331,357]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Stenosmylinae">Stenosmylinae</taxonomicName>
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, Porisminae,
|
||
<taxonomicName id="4C59B799043AFFF7F524796EFF7D06DA" authorityName="Esben-Petersen" authorityYear="1917" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Eidoporisminae">Eidoporisminae</taxonomicName>
|
||
: long, pectinately branched].
|
||
</paragraph>
|
||
<paragraph id="8BE6CC1A043AFFF7F0C579B7FB910531" blockId="2.[151,1437,151,897]" pageId="2" pageNumber="144">
|
||
<emphasis id="B92D1008043AFFF7F0C579B7FE1306FF" bold="true" box="[199,398,402,428]" pageId="2" pageNumber="144">Genera included</emphasis>
|
||
. Ten genera: †
|
||
<taxonomicName id="4C59B799043AFFF7F23779B6FCC906FE" ID-CoL="7JBZ" authority="Martynova, 1958" authorityName="Martynova" authorityYear="1958" box="[565,852,402,429]" class="Insecta" family="Osmylidae" genus="Sogjuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F23779B6FD1606FE" box="[565,651,403,429]" italics="true" pageId="2" pageNumber="144">Sogjuta</emphasis>
|
||
<bibRefCitation id="EFC8B1EB043AFFF7F29079B6FCC906FE" author="Martynova, O. M." box="[658,852,402,429]" pageId="2" pageNumber="144" refId="ref7960" refString="Martynova, O. M. (1958) New insects from the Permian and Mesozoic deposits of the USSR. In: Sharov, A. G. (Ed.), Materialy k Osnovam Paleontologii [Materials to " Fundamentals of Paleontology "]. Vol. 2. Academy of Sciences of the USSR, Moscow, pp. 69 - 94. [in Russian]" type="book" year="1958">Martynova, 1958</bibRefCitation>
|
||
</taxonomicName>
|
||
(Early to Late Jurassic,
|
||
<collectingCountry id="F34E8C8A043AFFF7F46579B6FB7506FE" box="[1127,1256,403,429]" name="Kyrgyzstan" pageId="2" pageNumber="144">Kyrgyzstan</collectingCountry>
|
||
and
|
||
<collectingCountry id="F34E8C8A043AFFF7F51D79B6FA0D06FD" box="[1311,1424,403,430]" name="Mongolia" pageId="2" pageNumber="144">Mongolia</collectingCountry>
|
||
),
|
||
<emphasis id="B92D1008043AFFF7F0957993FF6F0682" bold="true" box="[151,242,438,465]" pageId="2" pageNumber="144">sit. nov.</emphasis>
|
||
; †
|
||
<taxonomicName id="4C59B799043AFFF7F10D7992FD3C0682" ID-CoL="56Z5" authority="Wang et al., 2010" authorityName="Wang" authorityYear="2010" box="[271,673,439,465]" class="Insecta" family="Osmylidae" genus="Juraheterosmylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F10D7992FE480682" box="[271,469,439,465]" italics="true" pageId="2" pageNumber="144">Juraheterosmylus</emphasis>
|
||
<bibRefCitation id="EFC8B1EB043AFFF7F1DE7992FD3C0682" author="Wang, Y. J. & Liu, Z. Q. & Ren, D. & Shih, C. K." box="[476,673,439,465]" pageId="2" pageNumber="144" pagination="45 - 53" refId="ref8804" refString="Wang, Y. J., Liu, Z. Q., Ren, D. & Shih, C. K. (2010) A new genus of Protosmylinae from the Middle Jurassic of China (Neuroptera: Osmylidae). Zootaxa, 2480 (1), 45 - 53. https: // doi. org / 10.11646 / zootaxa. 2480.1.4" type="journal article" year="2010">
|
||
Wang
|
||
<emphasis id="B92D1008043AFFF7F2217992FDC90682" box="[547,596,439,465]" italics="true" pageId="2" pageNumber="144">et al</emphasis>
|
||
., 2010
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(late Middle / early Late Jurassic,
|
||
<collectingCountry id="F34E8C8A043AFFF7F4277992FBF00681" box="[1061,1133,439,466]" name="China" pageId="2" pageNumber="144">China</collectingCountry>
|
||
); †
|
||
<taxonomicName id="4C59B799043AFFF7F48F7992FED306A6" ID-CoL="5729" authority="Makarkin & Archibald, 2005" authorityName="Makarkin & Archibald" authorityYear="2005" class="Insecta" family="Osmylidae" genus="Jurosmylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F48F7992FA910682" box="[1165,1292,439,465]" italics="true" pageId="2" pageNumber="144">Jurosmylus</emphasis>
|
||
<bibRefCitation id="EFC8B1EB043AFFF7F5117992FED306A6" author="Makarkin, V. N. & Archibald, S. B." pageId="2" pageNumber="144" pagination="15 - 23" refId="ref7699" refString="Makarkin, V. N. & Archibald, S. B. (2005) Substitute names for three genera of fossil Neuroptera, with taxonomic notes. Zootaxa, 1054 (1), 15 - 23. https: // doi. org / 10.11646 / zootaxa. 1054.1.2" type="journal article" year="2005">Makarkin & Archibald, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
(Late Jurassic,
|
||
<collectingCountry id="F34E8C8A043AFFF7F1FD79FEFD1A06A5" box="[511,647,475,502]" name="Kazakhstan" pageId="2" pageNumber="144">Kazakhstan</collectingCountry>
|
||
); †
|
||
<taxonomicName id="4C59B799043AFFF7F2AA79FEFB8D06A6" ID-CoL="6XYK" authority="Jepson et al. 2009" authorityName="Jepson" authorityYear="2009" box="[680,1040,475,501]" class="Insecta" family="Osmylidae" genus="Protosmylina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F2AA79FEFCA306A6" box="[680,830,475,501]" italics="true" pageId="2" pageNumber="144">Protosmylina</emphasis>
|
||
<bibRefCitation id="EFC8B1EB043AFFF7F34779FEFB8D06A6" author="Jepson, J. E. & Makarkin, V. N. & Jarzembowski, E. A." box="[837,1040,475,501]" pageId="2" pageNumber="144" pagination="1325 - 1338" refId="ref7031" refString="Jepson, J. E., Makarkin, V. N. & Jarzembowski, E. A. (2009) New lacewings (Insecta: Neuroptera) from the Lower Cretaceous Wealden Supergroup of southern England. Cretaceous Research, 30, 1325 - 1338. https: // doi. org / 10.1016 / j. cretres. 2009.07.012" type="journal article" year="2009">
|
||
Jepson
|
||
<emphasis id="B92D1008043AFFF7F39B79FEFC5706A6" box="[921,970,475,501]" italics="true" pageId="2" pageNumber="144">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(Early Cretaceous,
|
||
<collectingCountry id="F34E8C8A043AFFF7F4F279FEFACF06A5" box="[1264,1362,475,502]" name="United Kingdom" pageId="2" pageNumber="144">England</collectingCountry>
|
||
); †
|
||
<taxonomicName id="4C59B799043AFFF7F57679FEFF76054A" ID-CoL="63GPY" authorityName="Cockerell" authorityYear="1908" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F57679FEFF76054A" italics="true" pageId="2" pageNumber="144">Osmylidia</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C59B799043AFFF7F10C79DAFDEB054A" ID-CoL="6XYL" authority=", Priabonian, Baltic" authorityName="Priabonian, Baltic" box="[270,630,511,537]" class="Insecta" family="Osmylidae" genus="Protosmylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F10C79DAFE04054A" box="[270,409,511,537]" italics="true" pageId="2" pageNumber="144">Protosmylus</emphasis>
|
||
, Priabonian, Baltic
|
||
</taxonomicName>
|
||
amber,
|
||
<emphasis id="B92D1008043AFFF7F2D779DAFCA2054A" bold="true" box="[725,831,511,537]" pageId="2" pageNumber="144">syn. nov.</emphasis>
|
||
), and †
|
||
<taxonomicName id="4C59B799043AFFF7F39B79DAFA94054A" ID-CoL="73KP" authority="Makarkin, 2017" authorityName="Makarkin" authorityYear="2017" box="[921,1289,511,537]" class="Insecta" family="Osmylidae" genus="Pseudosmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F39B79DAFBD7054A" box="[921,1098,511,537]" italics="true" pageId="2" pageNumber="144">Pseudosmylidia</emphasis>
|
||
<bibRefCitation id="EFC8B1EB043AFFF7F45679DAFA94054A" author="Makarkin, V. N." box="[1108,1289,511,537]" pageId="2" pageNumber="144" pagination="581 - 587" refId="ref7649" refString="Makarkin, V. N. (2017) A remarkable new genus of Protosmylinae (Neuroptera: Osmylidae) from late Eocene Florissant, Colorado. Zootaxa, 4268 (4), 581 - 587. https: // doi. org / 10.11646 / zootaxa. 4268.4.9" type="journal article" year="2017">Makarkin, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
(Priabonian, North America and Europe);
|
||
<taxonomicName id="4C59B799043AFFF7F1E97A06FCB4056E" ID-CoL="4WRY" authority="Kruger, 1913" authorityName="Kruger" authorityYear="1913" box="[491,809,547,573]" class="Insecta" family="Osmylidae" genus="Heterosmylus" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F1E97A06FD19056E" box="[491,644,547,573]" italics="true" pageId="2" pageNumber="144">Heterosmylus</emphasis>
|
||
<bibRefCitation id="EFC8B1EB043AFFF7F28C7A06FCB4056E" author="Kruger, L." box="[654,809,547,573]" pageId="2" pageNumber="144" pagination="3 - 123" refId="ref7259" refString="Kruger, L. (1913) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913">Krüger, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C59B799043AFFF7F3357A06FBF2056E" ID-CoL="4RB5" authority="Kruger, 1913" authorityName="Kruger" authorityYear="1913" box="[823,1135,547,573]" class="Insecta" family="Osmylidae" genus="Gryposmylus" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F3357A06FC56056E" box="[823,971,547,573]" italics="true" pageId="2" pageNumber="144">Gryposmylus</emphasis>
|
||
<bibRefCitation id="EFC8B1EB043AFFF7F3D77A06FBF2056E" author="Kruger, L." box="[981,1135,547,573]" pageId="2" pageNumber="144" pagination="3 - 123" refId="ref7259" refString="Kruger, L. (1913) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913">Krüger, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C59B799043AFFF7F4B37A06FA01056E" ID-CoL="5HVQ" authority="Navas, 1911" authorityName="Navas" authorityYear="1911" box="[1201,1436,547,573]" class="Insecta" family="Osmylidae" genus="Lysmus" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F4B37A06FA99056F" box="[1201,1284,547,572]" italics="true" pageId="2" pageNumber="144">Lysmus</emphasis>
|
||
<bibRefCitation id="EFC8B1EB043AFFF7F50C7A06FA01056E" author="Navas, L." box="[1294,1436,547,573]" pageId="2" pageNumber="144" pagination="111 - 117" refId="ref8162" refString="Navas, L. (1911) Nevropteres nouveaux de l'extreme Orient. Russkoe Entomologicheskoe Obozrenie [Revue Russe d'Entomologie], 11, 111 - 117." type="journal article" year="1911">Navás, 1911</bibRefCitation>
|
||
</taxonomicName>
|
||
(extant, all southeastern Asia);
|
||
<taxonomicName id="4C59B799043AFFF7F1F17A62FCA60532" ID-CoL="63LBD" authority="Kruger, 1913" authorityName="Kruger" authorityYear="1913" box="[499,827,583,609]" class="Insecta" family="Osmylidae" genus="Paryphosmylus" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F1F17A62FD3D0532" box="[499,672,583,609]" italics="true" pageId="2" pageNumber="144">Paryphosmylus</emphasis>
|
||
<bibRefCitation id="EFC8B1EB043AFFF7F2A47A62FCA60532" author="Kruger, L." box="[678,827,583,609]" pageId="2" pageNumber="144" pagination="3 - 123" refId="ref7259" refString="Kruger, L. (1913) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913">Krüger, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
(extant,
|
||
<collectingCountry id="F34E8C8A043AFFF7F39F7A62FB9D0531" box="[925,1024,583,610]" name="Ecuador" pageId="2" pageNumber="144">Ecuador</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3439F91043AFFF7F0C57A4EFB9404D2" pageId="2" pageNumber="144" type="discussion">
|
||
<paragraph id="8BE6CC1A043AFFF7F0C57A4EFCBC0446" blockId="2.[151,1437,151,897]" pageId="2" pageNumber="144">
|
||
<emphasis id="B92D1008043AFFF7F0C57A4EFEA605D6" bold="true" box="[199,315,619,645]" pageId="2" pageNumber="144">Remarks.</emphasis>
|
||
The venation of some species of
|
||
<taxonomicName id="4C59B799043AFFF7F2BD7A4EFC8F05D7" authorityName="Navas" authorityYear="1911" box="[703,786,619,644]" class="Insecta" family="Osmylidae" genus="Lysmus" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F2BD7A4EFC8F05D7" box="[703,786,619,644]" italics="true" pageId="2" pageNumber="144">Lysmus</emphasis>
|
||
</taxonomicName>
|
||
is very similar to that of the
|
||
<taxonomicName id="4C59B799043AFFF7F4677A4EFA9505D6" authorityName="Kruger" authorityYear="1913" box="[1125,1288,619,645]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
|
||
,
|
||
<emphasis id="B92D1008043AFFF7F5177A4EFAA005D7" box="[1301,1341,619,644]" italics="true" pageId="2" pageNumber="144">e.g.</emphasis>
|
||
,
|
||
<taxonomicName id="4C59B799043AFFF7F54F7A4EFE0505FA" authority="Navas, 1910" authorityName="Navas" authorityYear="1910" class="Insecta" family="Osmylidae" genus="Lysmus" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="species" species="harmandinus">
|
||
<emphasis id="B92D1008043AFFF7F54F7A4EFE9905FA" italics="true" pageId="2" pageNumber="144">L. harmandinus</emphasis>
|
||
Navás, 1910
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFC8B1EB043AFFF7F1D67AAAFD1905FA" author="Makarkin, V. N." box="[468,644,654,681]" pageId="2" pageNumber="144" pagination="35 - 47" refId="ref7562" refString="Makarkin, V. N. (1985) Review of the family Osmylidae (Neuroptera) of the fauna of the USSR. In: Lehr, P. A. & Storozhenko, S. Yu. (Chief Eds.), Taxonomy and ecology of arthropods of the Far East of the USSR. 129. Abstract. Far Eastern Scientific Centre of the Academy of Sciences of the USSR, Vladivostok, pp. 35 - 47. [in Russian]" type="book chapter" year="1985">Makarkin 1985</bibRefCitation>
|
||
:
|
||
<figureCitation id="1362D09F043AFFF7F28D7AAAFD4E05FA" box="[655,723,655,681]" captionStart="FIGURE 3" captionStartId="5.[151,250,1867,1891]" captionTargetBox="[151,1436,633,1835]" captionTargetId="figure-177@5.[151,1436,633,1843]" captionTargetPageId="5" captionText="FIGURE 3. Osmylidia taliae sp. nov., holotype UCM 67996 from the Green River Formation. A, photograph of the part; B, drawing of forewing venation; C, drawing of hind wing venation. Posterior branch of MA in red. Scale bar = 5 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4883189" httpUri="https://zenodo.org/record/4883189/files/figure.png" pageId="2" pageNumber="144">Fig. 3</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFC8B1EB043AFFF7F2E27AAAFC4E05FA" author="Winterton, S. L. & Martins, C. C. & Makarkin, V. N. & Ardila-Camacho, A. & Wang, Y. J." box="[736,979,655,681]" pageId="2" pageNumber="144" pagination="1 - 99" refId="ref8940" refString="Winterton, S. L., Martins, C. C., Makarkin, V. N., Ardila-Camacho, A. & Wang, Y. J. (2019) Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae & Saucrosmylidae): a review of living and fossil genera. Zootaxa, 4581 (1), 1 - 99. https: // doi. org / 10.11646 / zootaxa. 4581.1.1" type="journal article" year="2019">
|
||
Winterton
|
||
<emphasis id="B92D1008043AFFF7F35A7AAAFC1405FA" box="[856,905,655,681]" italics="true" pageId="2" pageNumber="144">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
: Fig. 34). Moreover, this genus had long been considered a member of the
|
||
<taxonomicName id="4C59B799043AFFF7F21C7A96FD5F059E" authorityName="Kruger" authorityYear="1913" box="[542,706,691,717]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C59B799043AFFF7F2CC7A96FCBC059F" authorityName="Navas" authorityYear="1911" box="[718,801,691,716]" class="Insecta" family="Osmylidae" genus="Lysmus" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F2CC7A96FCBC059F" box="[718,801,691,716]" italics="true" pageId="2" pageNumber="144">Lysmus</emphasis>
|
||
</taxonomicName>
|
||
, however, lacks an important synapomorphy of
|
||
<taxonomicName id="4C59B799043AFFF7F54F7A96FF6905A2" authorityName="Kruger" authorityYear="1913" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
|
||
, the presence of an anteriorly directed basal spur-like process in the hind wing MP. Also, the male genitalia of
|
||
<taxonomicName id="4C59B799043AFFF7F0B47ADEFE940447" authorityName="Navas" authorityYear="1911" box="[182,265,763,788]" class="Insecta" family="Osmylidae" genus="Lysmus" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F0B47ADEFE940447" box="[182,265,763,788]" italics="true" pageId="2" pageNumber="144">Lysmus</emphasis>
|
||
</taxonomicName>
|
||
are typical for
|
||
<taxonomicName id="4C59B799043AFFF7F1B67ADEFDCB0446" authorityName="Kruger" authorityYear="1913" box="[436,598,763,789]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Protosmylinae">Protosmylinae</taxonomicName>
|
||
(see Wang 2010).
|
||
</paragraph>
|
||
<paragraph id="8BE6CC1A043AFFF7F0C57B3AFB9404D2" blockId="2.[151,1437,151,897]" pageId="2" pageNumber="144">
|
||
According to Article 40.1 of ICZN, a family-group name is not replaced with a new family-group name when the name of its
|
||
<typeStatus id="54E272B8043AFFF7F1437B66FEEC040E" box="[321,369,835,861]" pageId="2" pageNumber="144">type</typeStatus>
|
||
genus is considered to be a junior synonym of another genus. Hence,
|
||
<taxonomicName id="4C59B799043AFFF7F4797B66FA80040E" authorityName="Kruger" authorityYear="1913" box="[1147,1309,835,861]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Protosmylinae">Protosmylinae</taxonomicName>
|
||
retains this name following designation of
|
||
<taxonomicName id="4C59B799043AFFF7F1F77B42FD1D04D2" box="[501,640,871,897]" class="Insecta" family="Osmylidae" genus="Protosmylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F1F77B42FD1D04D2" box="[501,640,871,897]" italics="true" pageId="2" pageNumber="144">Protosmylus</emphasis>
|
||
</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName id="4C59B799043AFFF7F38D7B42FB9F04D2" authorityName="Cockerell" authorityYear="1908" box="[911,1026,871,897]" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D1008043AFFF7F38D7B42FB9F04D2" box="[911,1026,871,897]" italics="true" pageId="2" pageNumber="144">Osmylidia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |