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<document id="67FAF814709CD9D7D007DFF0EB1B7720" ID-DOI="10.11646/zootaxa.5383.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10350971" ID-ZooBank="1D2AD0FF-CE98-462C-A238-CC3DBF2906C9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702283818271" checkinUser="plazi" docAuthor="Lin, Chen, Zhang, Chunmin &amp; Yang, Ding" docDate="2023" docId="03F03213FFA6196F9AD6FF0264D7457D" docLanguage="en" docName="zootaxa.5383.2.3.pdf" docOrigin="Zootaxa 5383 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.2.3/52446" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus huanglianus Lin &amp; Zhang &amp; Yang 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="161" masterDocId="FFC94A6BFFAE19679A41FF95672F4308" masterDocTitle="The genus Systenus Meigen, 1803 (Diptera: Dolichopodidae) from Yunnan, China, with descriptions of six new species" masterLastPageNumber="166" masterPageNumber="153" pageNumber="161" updateTime="1702284527099" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="699CDEB31FAE5F680789F8676F040F58">The genus Systenus Meigen, 1803 (Diptera: Dolichopodidae) from Yunnan, China, with descriptions of six new species</mods:title>
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<mods:namePart id="AEEA95FE7D95A431AAC76559D6061682">Lin, Chen</mods:namePart>
<mods:affiliation id="6866CB38F21B4AD1624D60BF95207FD9">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100193, China &amp; Institute of Life Science and Technology, Inner Mongolia Normal University, Huhhot 010022, China</mods:affiliation>
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<mods:namePart id="291C4800650EFECDA75C7EA79B0D3EEF">Zhang, Chunmin</mods:namePart>
<mods:affiliation id="BE14BBBF008B274E79210533F84BF5FB">Tongliao Agricultural Technology Extension Center, Tongliao 028000, China</mods:affiliation>
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<mods:namePart id="228CE0BBF665AEDD80B20F61FE9B4DD8">Yang, Ding</mods:namePart>
<mods:affiliation id="9E36987FF5C9CA5C5BF78BA17B46279E">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100193, China</mods:affiliation>
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<subSubSection id="C343D08EFFA6196F9AD6FF02651843B9" box="[151,567,151,178]" pageId="8" pageNumber="161" type="nomenclature">
<paragraph id="8BE68305FFA6196F9AD6FF02651843B9" blockId="8.[151,567,151,214]" box="[151,567,151,178]" pageId="8" pageNumber="161">
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<emphasis id="B92D5F17FFA6196F9AD6FF02651843B9" bold="true" box="[151,567,151,178]" pageId="8" pageNumber="161">
<emphasis id="B92D5F17FFA6196F9AD6FF0266FE43BA" bold="true" box="[151,465,151,178]" italics="true" pageId="8" pageNumber="161">
5.
<taxonomicName id="4C59F886FFA6196F9A86FF0266FE43BA" authority="Lin &amp; Zhang &amp; Yang, 2023" authorityName="Lin &amp; Zhang &amp; Yang" authorityYear="2023" box="[199,465,151,178]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="161" phylum="Arthropoda" rank="species" species="huanglianus" status="sp. nov.">Systenus huanglianus</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A21EE26CFFA6196F9B98FF02651843B9" box="[473,567,151,177]" pageId="8" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C343D08EFFA6196F9AD6FF2E666C43DE" box="[151,323,187,214]" pageId="8" pageNumber="161" type="description">
<paragraph id="8BE68305FFA6196F9AD6FF2E666C43DE" blockId="8.[151,567,151,214]" box="[151,323,187,214]" pageId="8" pageNumber="161">
(
<figureCitation id="13629F80FFA6196F9ADEFF2E67C743DE" box="[159,232,187,214]" captionStart="FIGURES 16. 1" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[203,1384,189,1907]" captionTargetId="figure-21@2.[193,1394,179,1917]" captionTargetPageId="2" captionText="FIGURES 16. 1. Systenus apicidigitatus sp. nov.; 2. Systenus basiflavus sp. nov.; 3. Systenus digitatus sp. nov.; 4. Systenus flavilateralis sp. nov.; 5. Systenus huanglianus sp. nov.; 6. Systenus latimaculatus sp. nov. All male. Scale bar =1.0mm." figureDoi="http://doi.org/10.5281/zenodo.10350973" httpUri="https://zenodo.org/record/10350973/files/figure.png" pageId="8" pageNumber="161">Figs 5</figureCitation>
,
<figureCitation id="13629F80FFA6196F9AB5FF2E661443DE" box="[244,315,187,214]" captionStart="FIGURES 1516" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[286,1301,191,1909]" captionTargetId="figure-18@9.[276,1310,181,1918]" captionTargetPageId="9" captionText="FIGURES 1516. Systenus huanglianus sp. nov. (male). 15. Antenna; 16. Male genitalia, lateral view. Scale bar = 0.1mm. FIGURES 1718. Systenus latimaculatus sp. nov. (male). 17. Antenna; 18. Male genitalia, lateral view. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10350981" httpUri="https://zenodo.org/record/10350981/files/figure.png" pageId="8" pageNumber="161">1516</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA6196F9AD6FE9666674282" pageId="8" pageNumber="161" type="diagnosis">
<paragraph id="8BE68305FFA6196F9AD6FE9666674282" blockId="8.[151,1437,259,1654]" pageId="8" pageNumber="161">
<emphasis id="B92D5F17FFA6196F9AD6FE9666254215" bold="true" box="[151,266,259,285]" pageId="8" pageNumber="161">Diagnosis</emphasis>
. Postpedicel rather long, 4.0 times longer than wide; arista 0.88 times as long as postpedicel. All coxae yellow, but mid coxa brownish yellow at base. R
<subScript id="17DD8140FFA6196F988AFEAC65CA424F" attach="left" box="[715,741,313,327]" fontSize="6" pageId="8" pageNumber="161">4+5</subScript>
and M
<subScript id="17DD8140FFA6196F9978FEAF646E4240" attach="left" box="[825,833,314,328]" fontSize="6" pageId="8" pageNumber="161">1</subScript>
weakly convergent apically. Surstylus basally rather thick, apical long thin and finger-like. Male cercus strongly bent, basally slightly thick, apically with long thick ventral process.
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA6196F9A86FE06668D465D" pageId="8" pageNumber="161" type="description">
<paragraph id="8BE68305FFA6196F9A86FE06637F42A6" blockId="8.[151,1437,259,1654]" box="[199,1104,403,430]" pageId="8" pageNumber="161">
<emphasis id="B92D5F17FFA6196F9A86FE06667642A5" bold="true" box="[199,345,403,429]" pageId="8" pageNumber="161">Description.</emphasis>
Male (
<figureCitation id="13629F80FFA6196F9BE9FE0666C242A6" box="[424,493,403,430]" captionStart="FIGURES 16. 1" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[203,1384,189,1907]" captionTargetId="figure-21@2.[193,1394,179,1917]" captionTargetPageId="2" captionText="FIGURES 16. 1. Systenus apicidigitatus sp. nov.; 2. Systenus basiflavus sp. nov.; 3. Systenus digitatus sp. nov.; 4. Systenus flavilateralis sp. nov.; 5. Systenus huanglianus sp. nov.; 6. Systenus latimaculatus sp. nov. All male. Scale bar =1.0mm." figureDoi="http://doi.org/10.5281/zenodo.10350973" httpUri="https://zenodo.org/record/10350973/files/figure.png" pageId="8" pageNumber="161">Fig. 5</figureCitation>
). Body length 3.0
<quantity id="4CA12EE0FFA6196F9885FE06640542A6" box="[708,810,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" pageId="8" pageNumber="161" unit="mm" value="3.25">3.25 mm</quantity>
, wing length 2.53.0 mm.
</paragraph>
<paragraph id="8BE68305FFA6196F9A86FE226301418D" blockId="8.[151,1437,259,1654]" pageId="8" pageNumber="161">
Head metallic green with pale gray pollinosity. Frons metallic green with white gray pollinosity; face dark metallic green with dense silvery white pollinosity, narrower than width of postpedicel. Hairs and bristles on head black. 2 oc, 2 vt, 2 pvt. Antenna black; pedicel with blackish apical bristles; postpedicel rather long, 4.0 times longer than wide, basally rather thick, apically finger-like, and with dense dark brown pubescence; arista apical, 0.9 times as long as postpedicel, with very short dark brown pubescence (
<figureCitation id="13629F80FFA6196F991AFDD26482416A" box="[859,941,583,610]" captionStart="FIGURES 1516" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[286,1301,191,1909]" captionTargetId="figure-18@9.[276,1310,181,1918]" captionTargetPageId="9" captionText="FIGURES 1516. Systenus huanglianus sp. nov. (male). 15. Antenna; 16. Male genitalia, lateral view. Scale bar = 0.1mm. FIGURES 1718. Systenus latimaculatus sp. nov. (male). 17. Antenna; 18. Male genitalia, lateral view. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10350981" httpUri="https://zenodo.org/record/10350981/files/figure.png" pageId="8" pageNumber="161">Fig. 15</figureCitation>
). Proboscis short, brown with brown or dark brown hairs; palpus small, brown, with black hairs and 1 long black apical bristle.
</paragraph>
<paragraph id="8BE68305FFA6196F9A86FD1A6618401D" blockId="8.[151,1437,259,1654]" pageId="8" pageNumber="161">Thorax metallic green with pale gray pollinosity, but pleura blackish except pteropleuron, hypopleuron black and anterior portion of sternopleuron. Hairs and bristles on thorax black; 6 dc, 1213 irregularly paired acr; 2 strong npl, 1 strong sa, 2 strong pa; sc strong and long, lsc thin and short. Propleuron with 1 brownish yellow bristle on lower portion.</paragraph>
<paragraph id="8BE68305FFA6196F9A86FCB566F9471A" blockId="8.[151,1437,259,1654]" pageId="8" pageNumber="161">Legs mainly yellow to dark yellow; all coxae yellow, but mid coxa brownish yellow at base; mid and hind tibiae dark yellow at tip; all tarsi dark yellow to brown onwards from tarsomere 1. Hairs and bristles on legs mainly black; hairs on all coxae yellow, bristles brownish; hind coxa with 1 brown outer bristle at middle. Mid trochanter with 3 black bristles; hind trochanter with 1 black outer bristle at middle. Mid tibia with 1 ad, 2 pd and 2 apical bristles; hind tibia with 3 pd and 2 apical bristles. Mid and hind tarsomeres 14 each with 4 short apical bristles. Relative length of tibiae and five tarsomeres of legs LI 3.8: 1.7: 1.2: 0.8: 0.4: 0.6; LII 5.7: 2.8: 1.4: 1.2: 0.6: 0.5; LIII 6.1: 1.1: 3.0: 1.3: 0.7: 0.5.</paragraph>
<paragraph id="8BE68305FFA6196F9A86FB8E64AE4751" blockId="8.[151,1437,259,1654]" pageId="8" pageNumber="161">
Wing hyaline, somewhat brownish; veins dark brown, R
<subScript id="17DD8140FFA6196F9978FBB8647C4733" attach="left" box="[825,851,1069,1083]" fontSize="6" pageId="8" pageNumber="161">4+5</subScript>
and M
<subScript id="17DD8140FFA6196F99E0FBBB64864734" attach="left" box="[929,937,1070,1084]" fontSize="6" pageId="8" pageNumber="161">1</subScript>
weakly convergent apically; CuAx ratio 0.5. Squama yellow, with fan of pale white bristles. Halter pale yellow.
</paragraph>
<paragraph id="8BE68305FFA6196F9A86FBF667F447A9" blockId="8.[151,1437,259,1654]" pageId="8" pageNumber="161">Abdomen dark metallic green with pale gray pollinosity, venter dark yellow. Hairs and bristles on abdomen black.</paragraph>
<paragraph id="8BE68305FFA6196F9A86FB3E661C4639" blockId="8.[151,1437,259,1654]" pageId="8" pageNumber="161">
Male genitalia (
<figureCitation id="13629F80FFA6196F9B39FB3E66E247CD" box="[376,461,1195,1221]" captionStart="FIGURES 1516" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[286,1301,191,1909]" captionTargetId="figure-18@9.[276,1310,181,1918]" captionTargetPageId="9" captionText="FIGURES 1516. Systenus huanglianus sp. nov. (male). 15. Antenna; 16. Male genitalia, lateral view. Scale bar = 0.1mm. FIGURES 1718. Systenus latimaculatus sp. nov. (male). 17. Antenna; 18. Male genitalia, lateral view. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10350981" httpUri="https://zenodo.org/record/10350981/files/figure.png" pageId="8" pageNumber="161">Fig. 16</figureCitation>
): epandrium distinctly longer than wide; epandrial lobe long, uniformly thick, apically with two minute denticles; surstylus basally rather thick, apically long finger-like. Cercus strongly bent, basally somewhat thick, apically with long thick finger-like ventral process. Hypandrium rather short, subconical. Phallus long and thin.
</paragraph>
<paragraph id="8BE68305FFA6196F9A86FAAE668D465D" blockId="8.[151,1437,259,1654]" box="[199,418,1339,1365]" pageId="8" pageNumber="161">
<emphasis id="B92D5F17FFA6196F9A86FAAE660A465D" bold="true" box="[199,293,1339,1365]" pageId="8" pageNumber="161">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
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<emphasis id="B92D5F17FFA6196F9A86FACA665E4671" bold="true" box="[199,369,1375,1401]" pageId="8" pageNumber="161">Type material.</emphasis>
<materialsCitation id="3B318958FFA6196F9B32FACA63794695" collectingDate="2019-04-19" collectingDateMax="2019-05-19" collectingDateMin="2019-04-19" collectionCode="CAU" collectorName="Malaise &amp; Liang Wang &amp; Xin Li" country="China" county="Lvchun" elevation="1779" latitude="23.057" location="Shuikuxiafang" longLatPrecision="1" longitude="102.777145" municipality="Yakou" pageId="8" pageNumber="161" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B92D5F17FFA6196F9B32FACA65364671" bold="true" box="[371,537,1375,1401]" pageId="8" pageNumber="161">
<typeStatus id="54E23DA7FFA6196F9B32FACA65364671" box="[371,537,1375,1401]" pageId="8" pageNumber="161" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
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,
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:
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,
<collectingCounty id="6287FB89FFA6196F98A7FACA64114671" box="[742,830,1375,1401]" pageId="8" pageNumber="161">Lvchun</collectingCounty>
,
<collectingMunicipality id="6B82197FFFA6196F9905FACA64A04671" box="[836,911,1375,1401]" pageId="8" pageNumber="161">Yakou</collectingMunicipality>
,
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[
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,
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], collected by
<collectorName id="26ACE6D3FFA6196F9BD3FA1666C44695" box="[402,491,1411,1437]" pageId="8" pageNumber="161">Malaise</collectorName>
trap,
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</date>
,
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and
<collectorName id="26ACE6D3FFA6196F99F5FA1664D14695" box="[948,1022,1411,1437]" pageId="8" pageNumber="161">Xin Li</collectorName>
(
<collectionCode id="ED481BC0FFA6196F9E4CFA1663614695" box="[1037,1102,1411,1437]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="8" pageNumber="161">CAU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B318958FFA6196F9E20FA16655846C9" collectingDate="2019-03-19" collectingDateMax="2019-04-19" collectingDateMin="2019-03-19" collectionCode="CAU" country="China" pageId="8" pageNumber="161" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B92D5F17FFA6196F9E20FA1662294695" bold="true" box="[1121,1286,1411,1437]" pageId="8" pageNumber="161">
<typeStatus id="54E23DA7FFA6196F9E20FA1663D34695" box="[1121,1276,1411,1437]" pageId="8" pageNumber="161" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
<specimenCount id="9D5F488CFFA6196F9F4CFA1662164696" box="[1293,1337,1411,1438]" count="1" pageId="8" pageNumber="161" type="male">1 ♂</specimenCount>
, same as
<typeStatus id="54E23DA7FFA6196F9AD6FA3267D646C9" box="[151,249,1447,1473]" pageId="8" pageNumber="161" type="holotype">holotype</typeStatus>
except,
<date id="FFE7A5C5FFA6196F9B16FA32653046C9" box="[343,543,1447,1473]" pageId="8" pageNumber="161" value="2019-03-19" valueMax="2019-04-19" valueMin="2019-03-19">
<collectingDate id="EFA35C2DFFA6196F9B16FA32653046C9" box="[343,543,1447,1473]" pageId="8" pageNumber="161" value="2019-03-19" valueMax="2019-04-19" valueMin="2019-03-19">2019.III.19IV.19</collectingDate>
</date>
(
<collectionCode id="ED481BC0FFA6196F986FFA32654046C9" box="[558,623,1447,1473]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="8" pageNumber="161">CAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA6196F9A86FA5E650E46ED" box="[199,545,1483,1509]" pageId="8" pageNumber="161" type="distribution">
<paragraph id="8BE68305FFA6196F9A86FA5E650E46ED" blockId="8.[151,1437,259,1654]" box="[199,545,1483,1509]" pageId="8" pageNumber="161">
<emphasis id="B92D5F17FFA6196F9A86FA5E664E46ED" bold="true" box="[199,353,1483,1509]" pageId="8" pageNumber="161">Distribution.</emphasis>
<collectingCountry id="F34EC395FFA6196F9B26FA5E668546ED" box="[359,426,1483,1509]" name="China" pageId="8" pageNumber="161">China</collectingCountry>
(
<collectingRegion id="499D4DE7FFA6196F9BF8FA5E653A46ED" box="[441,533,1483,1509]" country="China" name="Yunnan" pageId="8" pageNumber="161">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA6196F9A86FA7A62574559" pageId="8" pageNumber="161" type="discussion">
<paragraph id="8BE68305FFA6196F9A86FA7A62574559" blockId="8.[151,1437,259,1654]" pageId="8" pageNumber="161">
<emphasis id="B92D5F17FFA6196F9A86FA7A66144501" bold="true" box="[199,315,1519,1545]" pageId="8" pageNumber="161">Remarks.</emphasis>
<taxonomicName id="4C59F886FFA6196F9B00FA7A651E4501" authority="Lin &amp; Zhang &amp; Yang, 2023" authorityName="Lin &amp; Zhang &amp; Yang" authorityYear="2023" box="[321,561,1519,1545]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="161" phylum="Arthropoda" rank="species" species="huanglianus" status="sp. nov.">
<emphasis id="B92D5F17FFA6196F9B00FA7A651E4501" box="[321,561,1519,1545]" italics="true" pageId="8" pageNumber="161">Systenus huanglianus</emphasis>
</taxonomicName>
<emphasis id="B92D5F17FFA6196F9879FA7A65BF4501" bold="true" box="[568,656,1519,1545]" pageId="8" pageNumber="161">
<taxonomicNameLabel id="A21EE26CFFA6196F9879FA7A65BF4501" box="[568,656,1519,1545]" pageId="8" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is very similar to
<taxonomicName id="4C59F886FFA6196F9917FA7A63244501" authority="Lin &amp; Zhang &amp; Yang, 2023" authorityName="Lin &amp; Zhang &amp; Yang" authorityYear="2023" box="[854,1035,1519,1545]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="161" phylum="Arthropoda" rank="species" species="apicidigitatus" status="sp. nov.">
<emphasis id="B92D5F17FFA6196F9917FA7A63244501" box="[854,1035,1519,1545]" italics="true" pageId="8" pageNumber="161">S. apicidigitatus</emphasis>
</taxonomicName>
<emphasis id="B92D5F17FFA6196F9E50FA7A63464501" bold="true" box="[1041,1129,1519,1545]" pageId="8" pageNumber="161">
<taxonomicNameLabel id="A21EE26CFFA6196F9E50FA7A63464501" box="[1041,1129,1519,1545]" pageId="8" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but may be separated from the latter by the following features: antenna wholly dark brown; R
<subScript id="17DD8140FFA6196F992DF9B064A9453B" attach="left" box="[876,902,1573,1587]" fontSize="6" pageId="8" pageNumber="161">4+5</subScript>
and M
<subScript id="17DD8140FFA6196F9993F9B364F5453C" attach="left" box="[978,986,1574,1588]" fontSize="6" pageId="8" pageNumber="161">1</subScript>
weakly convergent apically; postpedicel rather long, 4.0 times longer than wide; arista 0.88 times as long as postpedicel; cercus basally somewhat thick.
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA6196F9A86F9CE64D7457D" box="[199,1016,1627,1654]" pageId="8" pageNumber="161" type="etymology">
<paragraph id="8BE68305FFA6196F9A86F9CE64D7457D" blockId="8.[151,1437,259,1654]" box="[199,1016,1627,1654]" pageId="8" pageNumber="161">
<emphasis id="B92D5F17FFA6196F9A86F9CE6661457D" bold="true" box="[199,334,1627,1653]" pageId="8" pageNumber="161">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="54E23DA7FFA6196F98F7F9C965C9457E" box="[694,742,1628,1654]" pageId="8" pageNumber="161">type</typeStatus>
locality Huanglianshan.
</paragraph>
</subSubSection>
</treatment>
</document>