642 lines
129 KiB
XML
642 lines
129 KiB
XML
<document id="E75D950DE805F3202D2597542A57F415" ID-CLB-Dataset="29073" ID-DOI="10.11646/zootaxa.4457.1.8" ID-GBIF-Dataset="a40b8c40-0523-455c-8b9b-6804ffa1f188" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457655" ID-ZooBank="23929DBE-D0F0-4A06-AFA5-DC43630E656A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539263841310" checkinUser="plazi" docAuthor="Cavallari, Daniel C. & Simone, Luiz R. L." docDate="2018" docId="03FF878EC80BFFFFF29ED5E47C3839EE" docLanguage="en" docName="zootaxa.4457.1.8.pdf" docOrigin="Zootaxa 4457 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calliostoma melliferum Cavallari & Simone 2018, sp. nov." docType="treatment" docUuid="1D076080-D8FB-4F0C-89E1-A85074FAE8B8" docUuidSource="ZooBank" docVersion="8" lastPageNumber="163" masterDocId="FFC6FFF6C80AFFF8F209D02379703E1B" masterDocTitle="A new species of Calliostoma (Vetigastropoda: Calliostomatidae) from Canopus Bank, off northeastern Brazil" masterLastPageNumber="166" masterPageNumber="156" pageNumber="157" updateTime="1698576889864" updateUser="ExternalLinkService">
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<mods:mods id="F91AB5F5C4769867085D1D0E5076926E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="ACA654C0DCF36F613F729DD8A980DC23">
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<mods:title id="88FF77B5D032E29343EBDBF7DFA4048C">A new species of Calliostoma (Vetigastropoda: Calliostomatidae) from Canopus Bank, off northeastern Brazil</mods:title>
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<mods:name id="7FE23856D374C62584C9752D486BD058" type="personal">
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<mods:namePart id="CE8C741E96762AB9A04E8A1470B74C0A">Cavallari, Daniel C.</mods:namePart>
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<mods:name id="AC7478AA4A457F94FCF618BCE81CEBD4" type="personal">
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<mods:roleTerm id="9993E3F5746781067F68C304D53718D1">Author</mods:roleTerm>
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<mods:namePart id="AC43D3BD7BCE6CC60C3DDBB905A7C65D">Simone, Luiz R. L.</mods:namePart>
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<mods:typeOfResource id="15B35535714C00BFC6A3E46F61659FAB">text</mods:typeOfResource>
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<mods:title id="8F369EE82C38079F65B67A05D87CD3BF">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="BD521E30217557F34F37EECDBAE6A152">
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<mods:date id="5970FE9D0CACD913EBA647CC6F214867">2018</mods:date>
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<mods:number id="CB4DC9F0F99D832F39E6AAB85D74F023">2018-08-07</mods:number>
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<mods:number id="9D3A1F6678EA9F74BD4C4CBCBBA67E22">4457</mods:number>
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<mods:classification id="9443409AA6609FEFA2546E18A956A5BB">journal article</mods:classification>
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<mods:identifier id="F47E6F2AA54A2B58207D4D1CF459C24E" type="DOI">10.11646/zootaxa.4457.1.8</mods:identifier>
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<treatment id="03FF878EC80BFFFFF29ED5E47C3839EE" ID-DOI="http://doi.org/10.5281/zenodo.5976901" ID-GBIF-Taxon="148691934" ID-Zenodo-Dep="5976901" LSID="urn:lsid:zoobank.org:act:1D076080-D8FB-4F0C-89E1-A85074FAE8B8" httpUri="http://treatment.plazi.org/id/03FF878EC80BFFFFF29ED5E47C3839EE" lastPageId="7" lastPageNumber="163" pageId="1" pageNumber="157">
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<subSubSection id="C34C6513C80BFFF9F29ED5E478503818" pageId="1" pageNumber="157" type="nomenclature">
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<paragraph id="8BE93698C80BFFF9F29ED5E47B6A3BF9" blockId="1.[151,538,1479,1539]" box="[151,538,1479,1506]" pageId="1" pageNumber="157">
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<heading id="D0A181F4C80BFFF9F29ED5E47B6A3BF9" bold="true" box="[151,538,1479,1506]" fontSize="11" level="1" pageId="1" pageNumber="157" reason="1">
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<emphasis id="B922EA8AC80BFFF9F29ED5E47B6A3BF9" bold="true" box="[151,538,1479,1506]" pageId="1" pageNumber="157">
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<taxonomicName id="4C564D1BC80BFFF9F29ED5E478C43BFA" authority="Cavallari & Simone, 2018" authorityName="Cavallari & Simone" authorityYear="2018" box="[151,436,1479,1505]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="157" phylum="Mollusca" rank="species" species="melliferum" status="sp. nov.">
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<emphasis id="B922EA8AC80BFFF9F29ED5E478C43BFA" bold="true" box="[151,436,1479,1505]" italics="true" pageId="1" pageNumber="157">Calliostoma melliferum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21157F1C80BFFF9F3B5D5EB7B6A3BF9" box="[444,538,1480,1506]" pageId="1" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE93698C80BFFF9F29ED5C978503818" blockId="1.[151,538,1479,1539]" box="[151,288,1514,1539]" pageId="1" pageNumber="157">
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(
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<figureCitation id="136D2A1DC80BFFF9F296D5C978673818" box="[159,279,1514,1539]" captionStart-0="FIGURES 1–11" captionStart-1="FIGURES 12–20" captionStart-2="FIGURES 21–29" captionStart-3="FIGURES 30–33" captionStartId-0="3.[151,264,1865,1887]" captionStartId-1="4.[151,264,1860,1882]" captionStartId-2="5.[151,264,1865,1887]" captionStartId-3="6.[151,264,642,664]" captionTargetBox-0="[152,1435,193,1844]" captionTargetBox-1="[189,1426,284,1786]" captionTargetBox-2="[152,1435,193,1844]" captionTargetBox-3="[152,1435,193,621]" captionTargetId-0="figure@3.[152,1435,193,1844]" captionTargetId-1="figure@4.[151,1437,193,1838]" captionTargetId-2="figure@5.[152,1435,193,1844]" captionTargetId-3="figure@6.[152,1435,193,621]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." captionText-1="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." captionText-2="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." captionText-3="FIGURES 30–33. Calliostoma melliferum micro-computed tomography of Paratype MZSP 70308; 30. 3D reconstruction of shell, apertural view (H = 4.5 mm, D = 4.1 mm; voxel siZe = 8.8 µm); 31. same, longitudinal section showing the columellar axis, slightly lateral view; 32. same, longitudinal section showing the columellar axis, apertural view; 33. same, umbilical view." httpUri-0="https://zenodo.org/record/1457657/files/figure.png" httpUri-1="https://zenodo.org/record/1457659/files/figure.png" httpUri-2="https://zenodo.org/record/1457661/files/figure.png" httpUri-3="https://zenodo.org/record/1457663/files/figure.png" pageId="1" pageNumber="157">Figs. 1–33</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34C6513C80BFFF9F29ED60F78FB395B" pageId="1" pageNumber="157" type="materials_examined">
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<paragraph id="8BE93698C80BFFF9F29ED60F7AF138E3" blockId="1.[151,1437,1580,2037]" pageId="1" pageNumber="157">
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<emphasis id="B922EA8AC80BFFF9F29ED60F7837385E" bold="true" box="[151,327,1580,1605]" pageId="1" pageNumber="157">Type material.</emphasis>
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Holotype: MZSP 53718. Paratypes: MZSP 53923, 1 spm (
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<date id="FFE81058C80BFFF9F1E7D60F7D29385E" box="[1006,1113,1580,1605]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-08">viii/2005</date>
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); MZSP 53693, 5 spm (
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<date id="FFE81058C80BFFF9F764D60F79A43873" bridgedPair="/" pageId="1" pageNumber="157" value="2005-08">viii/ 2005</date>
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); MZSP 53711, 2 shells (
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<date id="FFE81058C80BFFF9F3F5D66C7B183873" box="[508,616,1615,1640]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-08">viii/2005</date>
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); MZSP 55522, 1 spm (
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<date id="FFE81058C80BFFF9F188D66C7A9C3873" box="[897,1004,1615,1640]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-08">viii/2005</date>
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), all from type locality (same depth);
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<materialsCitation id="3B3E3CC5C80BFFF9F29ED6567A063896" ID-GBIF-Occurrence="1917076103" box="[151,886,1652,1677]" collectingDate="2005-08" collectionCode="MZSP" elevation="250" pageId="1" pageNumber="157" specimenCode="MZSP 53939, 2" specimenCount="1">
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same locality data,
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<elevation id="007BD1ABC80BFFF9F379D657789D3897" box="[368,493,1652,1677]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="1" pageNumber="157" unit="m" value="250.0" valueMax="260.0" valueMin="240.0">
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<quantity id="4CAE9B7DC80BFFF9F379D657789D3897" box="[368,493,1652,1677]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="1" pageNumber="157" unit="m" value="250.0" valueMax="260.0" valueMin="240.0">240-260 m</quantity>
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</elevation>
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,
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<specimenCode id="DBF09EE3C80BFFF9F3F0D6577BE23896" box="[505,658,1652,1677]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="157">MZSP 53939</specimenCode>
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,
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<specimenCode id="DBF09EE3C80BFFF9F097D6577BDC3897" box="[670,684,1652,1676]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="157">2</specimenCode>
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shells (
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<collectingDate id="EFACE9B0C80BFFF9F10BD6577A1E3896" box="[770,878,1652,1677]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-08">
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<date id="FFE81058C80BFFF9F10BD6577A1E3896" box="[770,878,1652,1677]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-08">viii/2005</date>
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</collectingDate>
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)
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</materialsCitation>
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; MZSP 66589, 1 shell (
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<date id="FFE81058C80BFFF9F689D6577D9B3896" box="[1152,1259,1652,1677]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-08">viii/2005</date>
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); MZSP 67310, 6 shells (
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<date id="FFE81058C80BFFF9F2F4D6B4782938AB" box="[253,345,1687,1712]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-11">xi/2005</date>
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); MZSP 70308, 70 shells (
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<date id="FFE81058C80BFFF9F080D6B47B9538AB" box="[649,741,1687,1712]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-11">xi/2005</date>
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); MZSP 71960, 1 shell (
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<date id="FFE81058C80BFFF9F1F2D6B47D1638AB" box="[1019,1126,1687,1712]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-08">viii/2005</date>
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); MZSP 71961, 1 shell (
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<date id="FFE81058C80BFFF9F774D6B479A438CE" bridgedPair="/" pageId="1" pageNumber="157" value="2005-11">xi/ 2005</date>
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); MZSP 90268, 150 shells (
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<date id="FFE81058C80BFFF9F01CD69F7B0138CE" box="[533,625,1724,1749]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-11">xi/2005</date>
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); MZSP 94218, 27 shells (
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<date id="FFE81058C80BFFF9F1ACD69F7D7138CE" box="[933,1025,1724,1749]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-11">xi/2005</date>
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); MZSP 94254, 13 shells (
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<date id="FFE81058C80BFFF9F73DD69F7CE038CE" box="[1332,1424,1724,1749]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-11">xi/2005</date>
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); MZSP 94837, 1 shell (
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<date id="FFE81058C80BFFF9F39DD6FC788038E3" box="[404,496,1759,1784]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-11">xi/2005</date>
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); MZSP
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<date id="FFE81058C80BFFF9F05AD6C37BDA38E3" box="[595,682,1760,1784]" pageId="1" pageNumber="157" value="14-10-04">100414</date>
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, 6 shells (
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<date id="FFE81058C80BFFF9F110D6FC7A0538E3" box="[793,885,1759,1784]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-11">xi/2005</date>
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).
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</paragraph>
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<paragraph id="8BE93698C80BFFF9F2CED72778FB395B" blockId="1.[151,1437,1580,2037]" pageId="1" pageNumber="157">
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||
<materialsCitation id="3B3E3CC5C80BFFF9F2CED72778FB395B" ID-GBIF-Occurrence="1917076108" collectingDate="2005-08" collectorName="Coltro & Costa" country="Brazil" elevation="60" latitude="-2.2402778" location="Canopus Bank" longLatPrecision="21" longitude="-38.380554" municipality="Fortaleza" pageId="1" pageNumber="157" specimenCount="1" stateProvince="Ceara" typeStatus="holotype">
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||
<emphasis id="B922EA8AC80BFFF9F2CED72778163906" bold="true" box="[199,358,1796,1821]" pageId="1" pageNumber="157">
|
||
<typeStatus id="54ED883AC80BFFF9F2CED72778723906" box="[199,258,1796,1821]" pageId="1" pageNumber="157">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F3417608C80BFFF9F364D72778C43906" box="[365,436,1796,1821]" name="Brazil" pageId="1" pageNumber="157">Brazil</collectingCountry>
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,
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<collectingRegion id="4992F87AC80BFFF9F3C9D7277B743907" box="[448,516,1796,1820]" country="Brazil" name="Ceara" pageId="1" pageNumber="157">Ceará</collectingRegion>
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||
,
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||
<quantity id="4CAE9B7DC80BFFF9F019D7277B153906" box="[528,613,1796,1821]" metricMagnitude="5" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="157" unit="km" value="190.0">190 km</quantity>
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||
off
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<collectingMunicipality id="6B8DACE2C80BFFF9F09CD7277A703906" box="[661,768,1796,1821]" pageId="1" pageNumber="157">Fortaleza</collectingMunicipality>
|
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,
|
||
<location id="8E896043C80BFFF9F102D7277AC33906" LSID="urn:lsid:plazi:treatment:03FF878EC80BFFFFF29ED5E47C3839EE:8E896043C80BFFF9F102D7277AC33906" box="[779,947,1796,1821]" country="Brazil" latitude="-2.2402778" longLatPrecision="21" longitude="-38.380554" municipality="Fortaleza" name="Canopus Bank" pageId="1" pageNumber="157" stateProvince="Ceara">Canopus Bank</location>
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||
,
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,
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<geoCoordinate id="EE62505FC80BFFF9F65BD7277D913906" box="[1106,1249,1796,1821]" direction="west" orientation="longitude" pageId="1" pageNumber="157" precision="15" value="-38.380554">38°22’50”W</geoCoordinate>
|
||
,
|
||
<elevation id="007BD1ABC80BFFF9F6E4D7277C573907" box="[1261,1319,1796,1821]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="157" unit="m" value="60.0">
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<quantity id="4CAE9B7DC80BFFF9F6E4D7277C573907" box="[1261,1319,1796,1821]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="157" unit="m" value="60.0">60 m</quantity>
|
||
</elevation>
|
||
(
|
||
<collectorName id="26A3534EC80BFFF9F73FD7277C0F3906" box="[1334,1407,1796,1821]" pageId="1" pageNumber="157">Coltro</collectorName>
|
||
&
|
||
<collectorName id="26A3534EC80BFFF9F29ED70B79A7395B" box="[151,215,1832,1856]" pageId="1" pageNumber="157">Costa</collectorName>
|
||
leg.,
|
||
<collectingDate id="EFACE9B0C80BFFF9F31CD704780F395B" box="[277,383,1831,1856]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-08">
|
||
<date id="FFE81058C80BFFF9F31CD704780F395B" box="[277,383,1831,1856]" bridgedPair="/" pageId="1" pageNumber="157" value="2005-08">viii/2005</date>
|
||
</collectingDate>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C6513C80BFFF9F2CED76F7DA039EF" pageId="1" pageNumber="157" type="diagnosis">
|
||
<paragraph id="8BE93698C80BFFF9F2CED76F7DA039EF" blockId="1.[151,1437,1580,2037]" pageId="1" pageNumber="157">
|
||
<emphasis id="B922EA8AC80BFFF9F2CED76F784E397E" bold="true" box="[199,318,1868,1893]" pageId="1" pageNumber="157">Diagnosis.</emphasis>
|
||
Shell with slightly convex whorls and spire slightly taller than wide; background color ivory, with honey yellow spiral cords and apex; suture marked by distinct thin, smooth suprasutural cord; base inflated, smooth; umbilicus narrow, deep. Surface of snout smooth, with small lateral expansions; pair of anterior odontophore cartilages fused; anterior and posterior odontophore cartilages partially fused; horizontal muscle divided into m6 and m6a; spiral caecum in stomach absent; mantle border bearing long papillae aligned in a single row.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C6513C808FFFFF2CED0B47D4538E7" lastPageId="7" lastPageNumber="163" pageId="2" pageNumber="158" type="description">
|
||
<paragraph id="8BE93698C808FFFAF2CED0B47DB73D9B" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="158">
|
||
<emphasis id="B922EA8AC808FFFAF2CED0B478293EAB" bold="true" box="[199,345,151,176]" pageId="2" pageNumber="158">Description.</emphasis>
|
||
<emphasis id="B922EA8AC808FFFAF368D0BB78E93EAB" box="[353,409,152,176]" italics="true" pageId="2" pageNumber="158">Shell</emphasis>
|
||
(
|
||
<figureCitation id="136D2A1DC808FFFAF3A0D0B47B663EAB" box="[425,534,151,176]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Figs. 1–9</figureCitation>
|
||
,
|
||
<figureCitation id="136D2A1DC808FFFAF02DD0BB7B1C3EAB" box="[548,620,152,176]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="2" pageNumber="158">21–23</figureCitation>
|
||
,
|
||
<figureCitation id="136D2A1DC808FFFAF070D0BB7BB43EAB" box="[633,708,152,176]" captionStart="FIGURES 30–33" captionStartId="6.[151,264,642,664]" captionTargetBox="[152,1435,193,621]" captionTargetId="figure@6.[152,1435,193,621]" captionTargetPageId="6" captionText="FIGURES 30–33. Calliostoma melliferum micro-computed tomography of Paratype MZSP 70308; 30. 3D reconstruction of shell, apertural view (H = 4.5 mm, D = 4.1 mm; voxel siZe = 8.8 µm); 31. same, longitudinal section showing the columellar axis, slightly lateral view; 32. same, longitudinal section showing the columellar axis, apertural view; 33. same, umbilical view." httpUri="https://zenodo.org/record/1457663/files/figure.png" pageId="2" pageNumber="158">30–33</figureCitation>
|
||
). Shell small (H =
|
||
<quantity id="4CAE9B7DC808FFFAF194D0BB7D7E3EAB" box="[925,1038,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="2" pageNumber="158" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
|
||
), trochoid, slightly taller than wide (H/D ~1.2), of up to 7.25 slightly convex whorls; ground color ivory white, with honey yellow apex (
|
||
<figureCitation id="136D2A1DC808FFFAF717D09F7C133ECF" box="[1310,1379,188,213]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Fig. 2</figureCitation>
|
||
) and spiral cords (
|
||
<figureCitation id="136D2A1DC808FFFAF320D0FC781C3EE3" box="[297,364,223,248]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Fig. 3</figureCitation>
|
||
), occasionally with nacreous gleam underneath, visible through transparency (
|
||
<figureCitation id="136D2A1DC808FFFAF6E9D0FC7C3D3EE3" box="[1248,1357,223,248]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Figs. 5–8</figureCitation>
|
||
). Spire tall (height ~1/2 of shell height), prominent, high and broad; spire angle ~60°. Protoconch (
|
||
<figureCitation id="136D2A1DC808FFFAF693D1277D993F07" box="[1178,1257,260,285]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Figs. 9</figureCitation>
|
||
,
|
||
<figureCitation id="136D2A1DC808FFFAF6FCD1277C623F07" box="[1269,1298,260,284]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="2" pageNumber="158">23</figureCitation>
|
||
) prominent, of ~1 whorl (width = 0.32–0.35 mm), glossy, honey yellow, apical fold tip with a darker-orange shade; surface sculptured with intersected delicate lines forming a honeycomb-like pattern of hexagons; transition marked by a thick terminal varix. Teleoconch I short (~0.1 whorl after protoconch), sculptured with closely spaced axial striae and 4–5 sinuous spiral ribs with variable width; growth scar nearly indistinct. Teleoconch II with up to ~6 convex whorls; first 3.5 whorls internally calcified in adult specimens (
|
||
<figureCitation id="136D2A1DC808FFFAF164D1947A883FCB" box="[877,1016,439,464]" captionStart="FIGURES 30–33" captionStartId="6.[151,264,642,664]" captionTargetBox="[152,1435,193,621]" captionTargetId="figure@6.[152,1435,193,621]" captionTargetPageId="6" captionText="FIGURES 30–33. Calliostoma melliferum micro-computed tomography of Paratype MZSP 70308; 30. 3D reconstruction of shell, apertural view (H = 4.5 mm, D = 4.1 mm; voxel siZe = 8.8 µm); 31. same, longitudinal section showing the columellar axis, slightly lateral view; 32. same, longitudinal section showing the columellar axis, apertural view; 33. same, umbilical view." httpUri="https://zenodo.org/record/1457663/files/figure.png" pageId="2" pageNumber="158">Figs. 31–32</figureCitation>
|
||
); whorls mainly sculptured with 4–6 spiral cords of similar width (
|
||
<figureCitation id="136D2A1DC808FFFAF3E5D1FF7B2A3FEF" box="[492,602,476,501]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Figs. 2, 7</figureCitation>
|
||
) that multiply by intercalation, separated by wider interspaces (~
|
||
<date id="FFE81058C808FFFAF74DD1FF7C123FEF" box="[1348,1378,476,500]" pageId="2" pageNumber="158">2x</date>
|
||
cord width); primary cords presenting numerous small, closely-packed, angular nodules, secondary cords smooth (
|
||
<figureCitation id="136D2A1DC808FFFAF76DD1DC79C53C27" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="2" pageNumber="158">Figs. 21</figureCitation>
|
||
,
|
||
<figureCitation id="136D2A1DC808FFFAF2CBD20779923C27" box="[194,226,548,572]" captionStart="FIGURES 30–33" captionStartId="6.[151,264,642,664]" captionTargetBox="[152,1435,193,621]" captionTargetId="figure@6.[152,1435,193,621]" captionTargetPageId="6" captionText="FIGURES 30–33. Calliostoma melliferum micro-computed tomography of Paratype MZSP 70308; 30. 3D reconstruction of shell, apertural view (H = 4.5 mm, D = 4.1 mm; voxel siZe = 8.8 µm); 31. same, longitudinal section showing the columellar axis, slightly lateral view; 32. same, longitudinal section showing the columellar axis, apertural view; 33. same, umbilical view." httpUri="https://zenodo.org/record/1457663/files/figure.png" pageId="2" pageNumber="158">30</figureCitation>
|
||
); ontogeny of cords as follows (
|
||
<figureCitation id="136D2A1DC808FFFAF05AD2077BD93C27" box="[595,681,548,573]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="2" pageNumber="158">Fig. 22</figureCitation>
|
||
): P1 commencing at 0.75 teleoconch whorl; P2 commencing right after protoconch; P3 commencing at 1.3 teleoconch whorl; P4 commencing just before P3 on same whorl; S1 commencing at 4.5 whorls; S2 commencing at 4.3 whorls. Suture delicate but distinct, marked by a thin, smooth suprasutural cord (
|
||
<figureCitation id="136D2A1DC808FFFAF361D2AC78B53CB3" box="[360,453,655,680]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="2" pageNumber="158">Figs. 21</figureCitation>
|
||
,
|
||
<figureCitation id="136D2A1DC808FFFAF3D8D2B37B6A3CB3" box="[465,538,656,680]" captionStart="FIGURES 30–33" captionStartId="6.[151,264,642,664]" captionTargetBox="[152,1435,193,621]" captionTargetId="figure@6.[152,1435,193,621]" captionTargetPageId="6" captionText="FIGURES 30–33. Calliostoma melliferum micro-computed tomography of Paratype MZSP 70308; 30. 3D reconstruction of shell, apertural view (H = 4.5 mm, D = 4.1 mm; voxel siZe = 8.8 µm); 31. same, longitudinal section showing the columellar axis, slightly lateral view; 32. same, longitudinal section showing the columellar axis, apertural view; 33. same, umbilical view." httpUri="https://zenodo.org/record/1457663/files/figure.png" pageId="2" pageNumber="158">30, 33</figureCitation>
|
||
). Aperture rounded to subquadrate (
|
||
<figureCitation id="136D2A1DC808FFFAF1B9D2AC7D463CB3" box="[944,1078,655,680]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Figs. 1, 5, 8</figureCitation>
|
||
), prosocline (
|
||
<figureCitation id="136D2A1DC808FFFAF6D9D2AC7C653CB3" box="[1232,1301,655,680]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Fig. 6</figureCitation>
|
||
), height ~1/ 3 of shell height, ivory white, nacreous, somewhat dislocated laterally in older specimens (
|
||
<figureCitation id="136D2A1DC808FFFAF683D2977DBE3CD6" box="[1162,1230,692,717]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Fig. 5</figureCitation>
|
||
); inner lip slightly flared, forming a narrow callus (
|
||
<figureCitation id="136D2A1DC808FFFAF017D2F47BE43CEB" box="[542,660,727,752]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Figs. 1, 5</figureCitation>
|
||
). Base distinctly convex, mostly smooth (
|
||
<figureCitation id="136D2A1DC808FFFAF685D2F47D933CEB" box="[1164,1251,727,752]" captionStart="FIGURES 30–33" captionStartId="6.[151,264,642,664]" captionTargetBox="[152,1435,193,621]" captionTargetId="figure@6.[152,1435,193,621]" captionTargetPageId="6" captionText="FIGURES 30–33. Calliostoma melliferum micro-computed tomography of Paratype MZSP 70308; 30. 3D reconstruction of shell, apertural view (H = 4.5 mm, D = 4.1 mm; voxel siZe = 8.8 µm); 31. same, longitudinal section showing the columellar axis, slightly lateral view; 32. same, longitudinal section showing the columellar axis, apertural view; 33. same, umbilical view." httpUri="https://zenodo.org/record/1457663/files/figure.png" pageId="2" pageNumber="158">Fig. 33</figureCitation>
|
||
) or very rarely sculptured with smooth, almost obsolete spiral cords (
|
||
<figureCitation id="136D2A1DC808FFFAF10AD2DF7A3A3D0F" box="[771,842,764,789]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Fig. 4</figureCitation>
|
||
), yet always presenting two smooth, similarly wide peripheral cords, the uppermost of which is visible as a suprasutural cord in the preceding whorls (
|
||
<figureCitation id="136D2A1DC808FFFAF73FD33C7CFE3D23" box="[1334,1422,799,824]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="2" pageNumber="158">Fig. 21</figureCitation>
|
||
); umbilicus narrow but deep, reaching first teleoconch whorls (
|
||
<figureCitation id="136D2A1DC808FFFAF143D3677AA33D47" box="[842,979,836,861]" captionStart="FIGURES 30–33" captionStartId="6.[151,264,642,664]" captionTargetBox="[152,1435,193,621]" captionTargetId="figure@6.[152,1435,193,621]" captionTargetPageId="6" captionText="FIGURES 30–33. Calliostoma melliferum micro-computed tomography of Paratype MZSP 70308; 30. 3D reconstruction of shell, apertural view (H = 4.5 mm, D = 4.1 mm; voxel siZe = 8.8 µm); 31. same, longitudinal section showing the columellar axis, slightly lateral view; 32. same, longitudinal section showing the columellar axis, apertural view; 33. same, umbilical view." httpUri="https://zenodo.org/record/1457663/files/figure.png" pageId="2" pageNumber="158">Figs. 31–32</figureCitation>
|
||
), bordered by 1–2 smooth, periumbilical spiral cords (
|
||
<figureCitation id="136D2A1DC808FFFAF321D34478073D9B" box="[296,375,871,896]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="136D2A1DC808FFFAF38AD34B78D23D9B" box="[387,418,872,896]" captionStart="FIGURES 30–33" captionStartId="6.[151,264,642,664]" captionTargetBox="[152,1435,193,621]" captionTargetId="figure@6.[152,1435,193,621]" captionTargetPageId="6" captionText="FIGURES 30–33. Calliostoma melliferum micro-computed tomography of Paratype MZSP 70308; 30. 3D reconstruction of shell, apertural view (H = 4.5 mm, D = 4.1 mm; voxel siZe = 8.8 µm); 31. same, longitudinal section showing the columellar axis, slightly lateral view; 32. same, longitudinal section showing the columellar axis, apertural view; 33. same, umbilical view." httpUri="https://zenodo.org/record/1457663/files/figure.png" pageId="2" pageNumber="158">33</figureCitation>
|
||
); interior sculptured with thin, closely spaced axial striae (
|
||
<figureCitation id="136D2A1DC808FFFAF63BD3447DCA3D9B" box="[1074,1210,871,896]" captionStart="FIGURES 30–33" captionStartId="6.[151,264,642,664]" captionTargetBox="[152,1435,193,621]" captionTargetId="figure@6.[152,1435,193,621]" captionTargetPageId="6" captionText="FIGURES 30–33. Calliostoma melliferum micro-computed tomography of Paratype MZSP 70308; 30. 3D reconstruction of shell, apertural view (H = 4.5 mm, D = 4.1 mm; voxel siZe = 8.8 µm); 31. same, longitudinal section showing the columellar axis, slightly lateral view; 32. same, longitudinal section showing the columellar axis, apertural view; 33. same, umbilical view." httpUri="https://zenodo.org/record/1457663/files/figure.png" pageId="2" pageNumber="158">Figs. 31, 33</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE93698C808FFFAF2CED3AF7B743DD3" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="158">
|
||
<emphasis id="B922EA8AC808FFFAF2CED3AF78343DBF" box="[199,324,908,932]" italics="true" pageId="2" pageNumber="158">Operculum</emphasis>
|
||
(
|
||
<figureCitation id="136D2A1DC808FFFAF35BD3AF78B93DBF" box="[338,457,908,933]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="2" pageNumber="158">Figs. 8, 10</figureCitation>
|
||
) golden yellow, corneous, circular, multispiral, with central nucleus; inner scar shallow, occupying ~1/2 of inner surface.
|
||
</paragraph>
|
||
<paragraph id="8BE93698C808FFFAF2CED3F77D09384B" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="158">
|
||
<emphasis id="B922EA8AC808FFFAF2CED3F778493DF7" box="[199,313,980,1004]" italics="true" pageId="2" pageNumber="158">Head-foot</emphasis>
|
||
(
|
||
<figureCitation id="136D2A1DC808FFFAF346D3F778903DF7" box="[335,480,980,1005]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="2" pageNumber="158">Figs. 12–14</figureCitation>
|
||
). Total body length slightly exceeding half of body whorl length in preserved specimens. Head bulged, located approximately in middle region of head-foot. Snout (
|
||
<emphasis id="B922EA8AC808FFFAF66ED3D47DF13A0B" bold="true" box="[1127,1153,1015,1040]" pageId="2" pageNumber="158">sn</emphasis>
|
||
) wide, slightly flattened, slightly longer than wide, ~1/4 of head-foot length; anterior end simple, slightly flap-like, lacking papillae; lateral edges of snout bearing small lateral projection on each side (
|
||
<figureCitation id="136D2A1DC808FFFAF156D41C7AC63A43" box="[863,950,1087,1112]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="2" pageNumber="158">Fig. 14</figureCitation>
|
||
). Mouth (
|
||
<emphasis id="B922EA8AC808FFFAF622D41C7D203A43" bold="true" box="[1067,1104,1087,1112]" pageId="2" pageNumber="158">mo</emphasis>
|
||
) longitudinal, small, located centrally on ventral surface of snout, at distal end of pseudoproboscis (
|
||
<figureCitation id="136D2A1DC808FFFAF1A0D4477A8D3A67" box="[937,1021,1124,1149]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="2" pageNumber="158">Fig. 14</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC808FFFAF60ED4477D563A66" bold="true" box="[1031,1062,1124,1149]" pageId="2" pageNumber="158">pb</emphasis>
|
||
), bearing a pair of closely-placed folds, running posteriorly up to ventral region of origin of snout, abruptly twisting leftward, dorso-posteriorly lying at left border between snout and foot. Cephalic tentacles (
|
||
<emphasis id="B922EA8AC808FFFAF12FD48F7A4C3ADE" bold="true" box="[806,828,1196,1221]" pageId="2" pageNumber="158">te</emphasis>
|
||
) about as long as snout. Ommatophores (
|
||
<emphasis id="B922EA8AC808FFFAF71DD48F7C493ADE" bold="true" box="[1300,1337,1196,1221]" pageId="2" pageNumber="158">om</emphasis>
|
||
) located on outer base of cephalic tentacles, length ~1/5 of tentacle length and almost same width of their base. Eyes large, dark, occupying most of distal region of ommatophores. Foot thick, occupying about half of total head-foot length, divided into mesopodium (
|
||
<emphasis id="B922EA8AC808FFFAF3D8D53478953B2B" bold="true" box="[465,485,1303,1328]" pageId="2" pageNumber="158">fs</emphasis>
|
||
) and epipodium (
|
||
<emphasis id="B922EA8AC808FFFAF0B1D5347BA43B2B" bold="true" box="[696,724,1303,1328]" pageId="2" pageNumber="158">ep</emphasis>
|
||
), surrounding lateral-dorsal region of mesopodium, far away from sole; anterior edge wider than remaining foot area; furrow of pedal gland very narrow, running along entire anterior edge of foot. Anterior third of epipodium relatively small and short, forming neck lobes (
|
||
<figureCitation id="136D2A1DC808FFFAF6D1D57C7C123B63" box="[1240,1378,1375,1400]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="2" pageNumber="158">Figs. 12–13</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC808FFFAF765D57C7CFB3B63" bold="true" box="[1388,1419,1375,1400]" pageId="2" pageNumber="158">nu</emphasis>
|
||
); anterior end covered by ommatophore; remaining 2/3 of epipodium relatively low and thick, bearing 4 pairs of long, slender tentacles (
|
||
<emphasis id="B922EA8AC808FFFAF3A7D58478B43BDB" bold="true" box="[430,452,1447,1472]" pageId="2" pageNumber="158">et</emphasis>
|
||
) inserted in its distal edge; each epipodial tentacle almost equidistantly distributed, similar-sized, distinctly separated from each other, apparently symmetrical on both sides. Pair of columellar muscles (
|
||
<figureCitation id="136D2A1DC808FFFAF30DD5CC782B3813" box="[260,347,1519,1544]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="2" pageNumber="158">Fig. 12</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC808FFFAF360D5CC78FD3813" bold="true" box="[361,397,1519,1544]" pageId="2" pageNumber="158">mc</emphasis>
|
||
) completely fused with each other, thick, length ~1/4 whorl, fused with each other along median line at considerable length. Haemocoel (
|
||
<figureCitation id="136D2A1DC808FFFAF0B8D6377A713837" box="[689,769,1556,1581]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="2" pageNumber="158">Fig. 14</figureCitation>
|
||
) spanning ~1/2 of head-foot length, ~twice as long as wide; lacking any visceral structures; buccal mass occupying ~1/2 of inner haemocoel volume.
|
||
</paragraph>
|
||
<paragraph id="8BE93698C808FFFAF2CED67F7CED39A2" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="158">
|
||
<emphasis id="B922EA8AC808FFFAF2CED67F7800386F" box="[199,368,1628,1652]" italics="true" pageId="2" pageNumber="158">Mantle organs</emphasis>
|
||
(
|
||
<figureCitation id="136D2A1DC808FFFAF38DD67F78AD386E" box="[388,477,1628,1653]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="2" pageNumber="158">Fig. 16</figureCitation>
|
||
). Mantle border (
|
||
<emphasis id="B922EA8AC808FFFAF0B8D67F7BA8386E" bold="true" box="[689,728,1628,1653]" pageId="2" pageNumber="158">mb</emphasis>
|
||
) relatively thick, whitish, bearing small and closely spaced papillae. Pallial cavity ~3/4 whorl in length. Osphradium (
|
||
<emphasis id="B922EA8AC808FFFAF15FD65C7A1F3883" bold="true" box="[854,879,1663,1688]" pageId="2" pageNumber="158">os</emphasis>
|
||
) node-like, relatively large, located at base of suspensory stalk of gill. Gill (
|
||
<emphasis id="B922EA8AC808FFFAF3E8D687788738A6" bold="true" box="[481,503,1700,1725]" pageId="2" pageNumber="158">gi</emphasis>
|
||
) located approximately at the centre of the cavity, gradually expanding towards left posteriorly; length ~90% of pallial cavity length; projected anteriorly by suspensory stalk (
|
||
<emphasis id="B922EA8AC808FFFAF683D6E47DDF38FB" bold="true" box="[1162,1199,1735,1760]" pageId="2" pageNumber="158">gm</emphasis>
|
||
) for ~1/5 of total gill length (
|
||
<emphasis id="B922EA8AC808FFFAF2E4D6CF7879391E" bold="true" box="[237,265,1772,1797]" pageId="2" pageNumber="158">ga</emphasis>
|
||
). Anterior end of gill pointed, projecting outside cavity (
|
||
<emphasis id="B922EA8AC808FFFAF185D6CF7AD8391E" bold="true" box="[908,936,1772,1797]" pageId="2" pageNumber="158">ga</emphasis>
|
||
). Both gill filaments relatively symmetrical, with larger filaments at middle gill level, gradually decreasing toward both sides. Ctenidial or efferent gill vessel (
|
||
<emphasis id="B922EA8AC808FFFAF2A9D71779CB3956" bold="true" box="[160,187,1844,1869]" pageId="2" pageNumber="158">cv</emphasis>
|
||
) about three times wider than afferent vessel, running along basal region of central axis of gill; directly connected to pericardium; gill separated from rectum for about 1/4 of pallial cavity width. No clear hypobranchial gland. Rectum (
|
||
<emphasis id="B922EA8AC808FFFAF345D75F7812398E" bold="true" box="[332,354,1916,1941]" pageId="2" pageNumber="158">rt</emphasis>
|
||
) narrow, slightly sinuous, ~1/2 of pallial cavity in length, running close along right edge. Anus narrow, long siphoned (~1/5 of rectum length), located halfway through length of cavity, posterior to mantle border.
|
||
</paragraph>
|
||
<caption id="DF296610C809FFFBF29ED76A7D6639C1" httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="3" pageNumber="159" startId="3.[151,264,1865,1887]" targetBox="[152,1435,193,1844]" targetPageId="3">
|
||
<paragraph id="8BE93698C809FFFBF29ED76A7D6639C1" blockId="3.[151,1436,1865,2010]" pageId="3" pageNumber="159">
|
||
<emphasis id="B922EA8AC809FFFBF29ED76A78353944" bold="true" box="[151,325,1865,1887]" pageId="3" pageNumber="159">FIGURES 1–11.</emphasis>
|
||
<taxonomicName id="4C564D1BC809FFFBF342D76A7B4B3944" authorityName="Cavallari & Simone" authorityYear="2018" box="[331,571,1865,1887]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="3" pageNumber="159" phylum="Mollusca" rank="species" species="melliferum">
|
||
<emphasis id="B922EA8AC809FFFBF342D76A7B4B3944" box="[331,571,1865,1887]" italics="true" pageId="3" pageNumber="159">Calliostoma melliferum</emphasis>
|
||
</taxonomicName>
|
||
shells, operculum and jaw;
|
||
<emphasis id="B922EA8AC809FFFBF15CD76A7A183944" bold="true" box="[853,872,1865,1887]" pageId="3" pageNumber="159">1.</emphasis>
|
||
Holotype MZSP 53718, apertural view (H = 7.42 mm);
|
||
<emphasis id="B922EA8AC809FFFBF29ED74479DA3966" bold="true" box="[151,170,1895,1917]" pageId="3" pageNumber="159">2.</emphasis>
|
||
same, apical view (D = 6.13 mm);
|
||
<emphasis id="B922EA8AC809FFFBF013D7447B5C3966" bold="true" box="[538,556,1895,1917]" pageId="3" pageNumber="159">3.</emphasis>
|
||
same, abapertural view;
|
||
<emphasis id="B922EA8AC809FFFBF139D7447A323966" bold="true" box="[816,834,1895,1917]" pageId="3" pageNumber="159">4.</emphasis>
|
||
same, umbilical view.
|
||
<emphasis id="B922EA8AC809FFFBF638D7447D333966" bold="true" box="[1073,1091,1895,1917]" pageId="3" pageNumber="159">5.</emphasis>
|
||
Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm).
|
||
<emphasis id="B922EA8AC809FFFBF001D7A57B453987" bold="true" box="[520,565,1926,1948]" pageId="3" pageNumber="159">6–8.</emphasis>
|
||
Paratype MZSP 55522 shell with soft parts;
|
||
<emphasis id="B922EA8AC809FFFBF1F4D7A57D603987" bold="true" box="[1021,1040,1926,1948]" pageId="3" pageNumber="159">6.</emphasis>
|
||
whole specimen, left lateral view (H = 5.0 mm);
|
||
<emphasis id="B922EA8AC809FFFBF2F7D786786039A0" bold="true" box="[254,272,1957,1979]" pageId="3" pageNumber="159">7.</emphasis>
|
||
same, apical view;
|
||
<emphasis id="B922EA8AC809FFFBF3E9D786788339A0" bold="true" box="[480,499,1957,1979]" pageId="3" pageNumber="159">8.</emphasis>
|
||
same, apertural view (D = 4.6 mm);
|
||
<emphasis id="B922EA8AC809FFFBF173D7867AFD39A0" bold="true" box="[890,909,1957,1979]" pageId="3" pageNumber="159">9.</emphasis>
|
||
Holotype, protoconch detail (scale = 0.2 mm).
|
||
<emphasis id="B922EA8AC809FFFBF775D7867CEC39A0" bold="true" box="[1404,1436,1957,1979]" pageId="3" pageNumber="159">10.</emphasis>
|
||
Paratype MZSP 55522 operculum;
|
||
<emphasis id="B922EA8AC809FFFBF3F4D7E77B6A39C1" bold="true" box="[509,538,1988,2010]" pageId="3" pageNumber="159">11.</emphasis>
|
||
same, jaw plates, ventral view (scales = 0.5 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF296610C80EFFFCF29ED7677A8139E8" httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="4" pageNumber="160" startId="4.[151,264,1860,1882]" targetBox="[189,1426,284,1786]" targetPageId="4">
|
||
<paragraph id="8BE93698C80EFFFCF29ED7677A8139E8" blockId="4.[151,1436,1860,2035]" pageId="4" pageNumber="160">
|
||
<emphasis id="B922EA8AC80EFFFCF29ED76778253941" bold="true" box="[151,341,1860,1882]" pageId="4" pageNumber="160">FIGURES 12–20.</emphasis>
|
||
Anatomy of
|
||
<taxonomicName id="4C564D1BC80EFFFCF3D6D7677BA03941" authorityName="Cavallari & Simone" authorityYear="2018" box="[479,720,1860,1882]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="160" phylum="Mollusca" rank="species" species="melliferum">
|
||
<emphasis id="B922EA8AC80EFFFCF3D6D7677BA03941" box="[479,720,1860,1882]" italics="true" pageId="4" pageNumber="160">Calliostoma melliferum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B922EA8AC80EFFFCF0D6D7677B8F3941" bold="true" box="[735,767,1860,1882]" pageId="4" pageNumber="160">12.</emphasis>
|
||
head-foot, right view, anterior portion of mantle also shown;
|
||
<emphasis id="B922EA8AC80EFFFCF775D7677CEC3941" bold="true" box="[1404,1436,1860,1882]" pageId="4" pageNumber="160">13.</emphasis>
|
||
same, left view;
|
||
<emphasis id="B922EA8AC80EFFFCF349D74178103963" bold="true" box="[320,352,1890,1912]" pageId="4" pageNumber="160">14.</emphasis>
|
||
head and haemocoel, ventral view, foot and columellar muscle removed;
|
||
<emphasis id="B922EA8AC80EFFFCF65DD7417D043963" bold="true" box="[1108,1140,1890,1912]" pageId="4" pageNumber="160">15.</emphasis>
|
||
foregut and nerve ring, right view;
|
||
<emphasis id="B922EA8AC80EFFFCF2D0D7A27989398C" bold="true" box="[217,249,1921,1943]" pageId="4" pageNumber="160">16.</emphasis>
|
||
pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed;
|
||
<emphasis id="B922EA8AC80EFFFCF72AD7A27C32398C" bold="true" box="[1315,1346,1921,1943]" pageId="4" pageNumber="160">17.</emphasis>
|
||
foregut, ventral view, longitudinally opened, odontophore deflected to right;
|
||
<emphasis id="B922EA8AC80EFFFCF143D7BC7A1A39AE" bold="true" box="[842,874,1951,1973]" pageId="4" pageNumber="160">18.</emphasis>
|
||
nerve ring, ventral view;
|
||
<emphasis id="B922EA8AC80EFFFCF666D7BC7DFF39AE" bold="true" box="[1135,1167,1951,1973]" pageId="4" pageNumber="160">19.</emphasis>
|
||
odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected;
|
||
<emphasis id="B922EA8AC80EFFFCF1DDD79D7A8439CF" bold="true" box="[980,1012,1982,2004]" pageId="4" pageNumber="160">20.</emphasis>
|
||
same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF296610C80FFFFDF29ED76A7C0B39A0" httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="5" pageNumber="161" startId="5.[151,264,1865,1887]" targetBox="[152,1435,193,1844]" targetPageId="5">
|
||
<paragraph id="8BE93698C80FFFFDF29ED76A7C0B39A0" blockId="5.[151,1436,1865,1979]" pageId="5" pageNumber="161">
|
||
<emphasis id="B922EA8AC80FFFFDF29ED76A78243944" bold="true" box="[151,340,1865,1887]" pageId="5" pageNumber="161">FIGURES 21–29.</emphasis>
|
||
<taxonomicName id="4C564D1BC80FFFFDF353D76A7B3A3944" authorityName="Cavallari & Simone" authorityYear="2018" box="[346,586,1865,1887]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="5" pageNumber="161" phylum="Mollusca" rank="species" species="melliferum">
|
||
<emphasis id="B922EA8AC80FFFFDF353D76A7B3A3944" box="[346,586,1865,1887]" italics="true" pageId="5" pageNumber="161">Calliostoma melliferum</emphasis>
|
||
</taxonomicName>
|
||
in SEM.
|
||
<emphasis id="B922EA8AC80FFFFDF0A5D76A7BB53944" bold="true" box="[684,709,1865,1887]" pageId="5" pageNumber="161">21</emphasis>
|
||
. Paratype MZSP 90268, apertural view (D = 4.3 mm);
|
||
<emphasis id="B922EA8AC80FFFFDF6F1D76A7C683944" bold="true" box="[1272,1304,1865,1887]" pageId="5" pageNumber="161">22.</emphasis>
|
||
same, apical view (H = 4.8 mm);
|
||
<emphasis id="B922EA8AC80FFFFDF36FD74478F63966" bold="true" box="[358,390,1895,1917]" pageId="5" pageNumber="161">23.</emphasis>
|
||
same, protoconch detail (scale = 0.2 mm).
|
||
<emphasis id="B922EA8AC80FFFFDF132D7447AF03966" bold="true" box="[827,896,1895,1917]" pageId="5" pageNumber="161">24–29.</emphasis>
|
||
Radulae in SEM:
|
||
<emphasis id="B922EA8AC80FFFFDF632D7447D2B3966" bold="true" box="[1083,1115,1895,1917]" pageId="5" pageNumber="161">24.</emphasis>
|
||
wide view (scale = 50 µm);
|
||
<emphasis id="B922EA8AC80FFFFDF775D7447CEC3966" bold="true" box="[1404,1436,1895,1917]" pageId="5" pageNumber="161">25.</emphasis>
|
||
same, detail of right half (scale= 30 µm);
|
||
<emphasis id="B922EA8AC80FFFFDF04BD7A57B123987" bold="true" box="[578,610,1926,1948]" pageId="5" pageNumber="161">26.</emphasis>
|
||
same, detail of central region (scale = 10 µm);
|
||
<emphasis id="B922EA8AC80FFFFDF641D7A57D183987" bold="true" box="[1096,1128,1926,1948]" pageId="5" pageNumber="161">27.</emphasis>
|
||
same, detail of right marginal teeth (scale = 10 µm);
|
||
<emphasis id="B922EA8AC80FFFFDF375D78678EC39A0" bold="true" box="[380,412,1957,1979]" pageId="5" pageNumber="161">28.</emphasis>
|
||
another specimen, wide view, (scale = 50 µm);
|
||
<emphasis id="B922EA8AC80FFFFDF18BD7867AD239A0" bold="true" box="[898,930,1957,1979]" pageId="5" pageNumber="161">29.</emphasis>
|
||
same, detail of central region (scale = 20 µm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF296610C80CFFFEF29ED2A17CED3CCD" httpUri="https://zenodo.org/record/1457663/files/figure.png" pageId="6" pageNumber="162" startId="6.[151,264,642,664]" targetBox="[152,1435,193,621]" targetPageId="6">
|
||
<paragraph id="8BE93698C80CFFFEF29ED2A17CED3CCD" blockId="6.[151,1437,642,726]" pageId="6" pageNumber="162">
|
||
<emphasis id="B922EA8AC80CFFFEF29ED2A178263C83" bold="true" box="[151,342,642,664]" pageId="6" pageNumber="162">FIGURES 30–33.</emphasis>
|
||
<taxonomicName id="4C564D1BC80CFFFEF354D2A17B3F3C83" authorityName="Cavallari & Simone" authorityYear="2018" box="[349,591,642,664]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="6" pageNumber="162" phylum="Mollusca" rank="species" species="melliferum">
|
||
<emphasis id="B922EA8AC80CFFFEF354D2A17B3F3C83" box="[349,591,642,664]" italics="true" pageId="6" pageNumber="162">Calliostoma melliferum</emphasis>
|
||
</taxonomicName>
|
||
micro-computed tomography of Paratype MZSP 70308;
|
||
<emphasis id="B922EA8AC80CFFFEF696D2A17DCF3C83" bold="true" box="[1183,1215,642,664]" pageId="6" pageNumber="162">30.</emphasis>
|
||
3D reconstruction of shell, apertural view (H = 4.5 mm, D = 4.1 mm; voxel siZe = 8.8 µm);
|
||
<emphasis id="B922EA8AC80CFFFEF17BD2827AE23CAC" bold="true" box="[882,914,673,695]" pageId="6" pageNumber="162">31.</emphasis>
|
||
same, longitudinal section showing the columellar axis, slightly lateral view;
|
||
<emphasis id="B922EA8AC80CFFFEF3A8D2E378B13CCD" bold="true" box="[417,449,704,726]" pageId="6" pageNumber="162">32.</emphasis>
|
||
same, longitudinal section showing the columellar axis, apertural view;
|
||
<emphasis id="B922EA8AC80CFFFEF692D2E37DCA3CCD" bold="true" box="[1179,1210,704,726]" pageId="6" pageNumber="162">33.</emphasis>
|
||
same, umbilical view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE93698C80CFFFEF2CED3207DBD3DB0" blockId="6.[151,1437,770,2019]" pageId="6" pageNumber="162">
|
||
<emphasis id="B922EA8AC80CFFFEF2CED32078143D01" box="[199,356,771,795]" italics="true" pageId="6" pageNumber="162">Visceral mass</emphasis>
|
||
(
|
||
<figureCitation id="136D2A1DC80CFFFEF37CD32178BB3D00" box="[373,459,770,795]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="6" pageNumber="162">Fig. 16</figureCitation>
|
||
). Located about 2.5–3 whorls posterior to pallial cavity, leaving some of apical shell whorls empty. Gonad (
|
||
<emphasis id="B922EA8AC80CFFFEF3ACD30578B13D24" bold="true" box="[421,449,806,831]" pageId="6" pageNumber="162">go</emphasis>
|
||
) white, occupying ~2/3 of internal volume, located mainly in superior and external regions. Digestive gland (
|
||
<emphasis id="B922EA8AC80CFFFEF3C1D36978953D78" bold="true" box="[456,485,842,867]" pageId="6" pageNumber="162">dg</emphasis>
|
||
) beige, occupying regions close to columella; thicker around stomach. Stomach located close to posterior end of pallial cavity, occupying ~1/3 of local volume, ~0.15 whorl long, somewhat fusiform, slightly flattened. Reno-pericardial structures as anterior end, encroaching pallial roof.
|
||
</paragraph>
|
||
<paragraph id="8BE93698C80CFFFEF2CED3957D793A20" blockId="6.[151,1437,770,2019]" pageId="6" pageNumber="162">
|
||
<emphasis id="B922EA8AC80CFFFEF2CED3957B3E3DD5" box="[199,590,950,974]" italics="true" pageId="6" pageNumber="162">Circulatory and excretory systems</emphasis>
|
||
(
|
||
<figureCitation id="136D2A1DC80CFFFEF068D3957BC73DD4" box="[609,695,950,975]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="6" pageNumber="162">Fig. 16</figureCitation>
|
||
). Pericardium (
|
||
<emphasis id="B922EA8AC80CFFFEF163D3957AF63DD4" bold="true" box="[874,902,950,975]" pageId="6" pageNumber="162">pc</emphasis>
|
||
) not seen in detail due to preservation issues; relatively small, located between pallial cavity and visceral mass, just posterior to left kidney, close to median line, slightly dislocated to left. Left kidney (
|
||
<emphasis id="B922EA8AC80CFFFEF08DD3DD7BEB3A0C" bold="true" box="[644,667,1022,1047]" pageId="6" pageNumber="162">kl</emphasis>
|
||
) ~3-times longer than right kidney (
|
||
<emphasis id="B922EA8AC80CFFFEF662D3DD7DF73A0C" bold="true" box="[1131,1159,1022,1047]" pageId="6" pageNumber="162">kr</emphasis>
|
||
), white, solid, simple. Nephrostome (
|
||
<emphasis id="B922EA8AC80CFFFEF337D401782A3A20" bold="true" box="[318,346,1058,1083]" pageId="6" pageNumber="162">ne</emphasis>
|
||
) at anterior end of kidneys, preceded by a short hollow stalk.
|
||
</paragraph>
|
||
<paragraph id="8BE93698C80CFFFEF2CED4647DA639A4" blockId="6.[151,1437,770,2019]" pageId="6" pageNumber="162">
|
||
<emphasis id="B922EA8AC80CFFFEF2CED46478F33A45" box="[199,387,1095,1118]" italics="true" pageId="6" pageNumber="162">Digestive system</emphasis>
|
||
(
|
||
<figureCitation id="136D2A1DC80CFFFEF39BD465789E3A44" box="[402,494,1094,1119]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="6" pageNumber="162">Figs. 11</figureCitation>
|
||
,
|
||
<figureCitation id="136D2A1DC80CFFFEF3F3D4647BE93A44" box="[506,665,1095,1119]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="6" pageNumber="162">14–17, 19–20</figureCitation>
|
||
). Oral tube (
|
||
<figureCitation id="136D2A1DC80CFFFEF12DD4657A093A44" box="[804,889,1094,1119]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="6" pageNumber="162">Fig. 15</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80CFFFEF18DD4657AEB3A44" bold="true" box="[900,923,1094,1119]" pageId="6" pageNumber="162">ot</emphasis>
|
||
) with ~1/3 of odontophore length, wide, with relatively thick muscular walls. Jaw and peribuccal muscles (
|
||
<figureCitation id="136D2A1DC80CFFFEF16DD4497ACD3A98" box="[868,957,1130,1155]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="6" pageNumber="162">Fig. 15</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80CFFFEF1C2D4497A9C3A98" bold="true" box="[971,1004,1130,1155]" pageId="6" pageNumber="162">mj</emphasis>
|
||
) very thick, originating laterally and ventrally at inner surface of mouth, close to median line, running divergently towards posterior and lateral regions, inserting in ventral surface of odontophore; additionally, also inserting in narrow region of middle level on ventral side of anterior odontophore cartilages (
|
||
<figureCitation id="136D2A1DC80CFFFEF070D4F57BA43AF4" box="[633,724,1238,1263]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="6" pageNumber="162">Fig. 20</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80CFFFEF0EFD4F57A773AF4" bold="true" box="[742,775,1238,1263]" pageId="6" pageNumber="162">mj</emphasis>
|
||
). Odontophore (
|
||
<emphasis id="B922EA8AC80CFFFEF1D9D4F57A9D3AF4" bold="true" box="[976,1005,1238,1263]" pageId="6" pageNumber="162">od</emphasis>
|
||
) occupying ~1/4 haemocoel length. Odontophore muscles (
|
||
<figureCitation id="136D2A1DC80CFFFEF395D4D97B563B08" box="[412,550,1274,1299]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="6" pageNumber="162">Figs. 19–20</figureCitation>
|
||
):
|
||
<emphasis id="B922EA8AC80CFFFEF030D4D97B2E3B08" bold="true" box="[569,606,1274,1299]" pageId="6" pageNumber="162">m1</emphasis>
|
||
, series of small muscles connecting buccal mass to adjacent inner surface of snout and haemocoel;
|
||
<emphasis id="B922EA8AC80CFFFEF3B9D53D78AD3B2C" bold="true" box="[432,477,1310,1335]" pageId="6" pageNumber="162">m1l</emphasis>
|
||
, pair of small lateral protractors (
|
||
<figureCitation id="136D2A1DC80CFFFEF15CD53D7AD73B2C" box="[853,935,1310,1335]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="6" pageNumber="162">Fig. 15</figureCitation>
|
||
), originating on lateral middle wall of snout, running towards anterior region, inserting in lateral-posterior region of odontophore; pair of buccal mass retractors absent;
|
||
<emphasis id="B922EA8AC80CFFFEF2E6D54578653B64" bold="true" box="[239,277,1382,1407]" pageId="6" pageNumber="162">m4</emphasis>
|
||
, broad pair of dorsal tensor muscles of radula and subradular membrane, originating part in anterior cartilages, along their ventral surfaces at some distance from median line, and part in posterior cartilages, on their posterior and lateral surfaces, surrounding lateral and ventral surfaces of anterior cartilages, inserting along subradular membrane, in its dorsal region exposed inside buccal cavity, with a portion in radular ribbon in region preceding buccal cavity;
|
||
<emphasis id="B922EA8AC80CFFFEF3B9D5D578A53814" bold="true" box="[432,469,1526,1551]" pageId="6" pageNumber="162">m5</emphasis>
|
||
, pair of small auxiliary dorsal tensor muscles of radula, originating on ventral surface of posterior cartilages, running towards dorsal and medial regions along ~1/5 of odontophore length, inserting in radular ribbon in region preceding buccal cavity;
|
||
<emphasis id="B922EA8AC80CFFFEF0D2D61D7A70384C" bold="true" box="[731,768,1598,1623]" pageId="6" pageNumber="162">m6</emphasis>
|
||
, horizontal muscle, uniting both anterior cartilages along ~half their ventral edge (just posterior to fusion of cartilages), on their external surface;
|
||
<emphasis id="B922EA8AC80CFFFEF679D6417DD43860" bold="true" box="[1136,1188,1634,1659]" pageId="6" pageNumber="162">m6a</emphasis>
|
||
, secondary horizontal muscle, length ~1/10 odontophore’s length, located at middle level between posterior end of
|
||
<emphasis id="B922EA8AC80CFFFEF6E1D6A57C7E3884" bold="true" box="[1256,1294,1670,1695]" pageId="6" pageNumber="162">m6</emphasis>
|
||
and that of posterior cartilage;
|
||
<emphasis id="B922EA8AC80CFFFEF37BD68978E738D8" bold="true" box="[370,407,1706,1731]" pageId="6" pageNumber="162">m8</emphasis>
|
||
, pair of broad approximators of cartilages, originating in anterior cartilages in a small area on their lateral surface, posterior to insertion of
|
||
<emphasis id="B922EA8AC80CFFFEF0D7D6ED7B8F38FC" bold="true" box="[734,767,1742,1767]" pageId="6" pageNumber="162">mj</emphasis>
|
||
, running towards posterior region, decreasing gradually, inserting in middle region of anterior surface of posterior cartilages;
|
||
<emphasis id="B922EA8AC80CFFFEF1BDD6D17A973910" bold="true" box="[948,999,1778,1803]" pageId="6" pageNumber="162">m10</emphasis>
|
||
, small pair of odontophore protractor muscles (
|
||
<figureCitation id="136D2A1DC80CFFFEF309D73578223934" box="[256,338,1814,1839]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="6" pageNumber="162">Fig. 15</figureCitation>
|
||
), originating at middle level of ventral surface of oral tube, running towards posterior region along ~1/4 of odontophore length, inserting in anterior-ventral region of odontophore;
|
||
<emphasis id="B922EA8AC80CFFFEF64CD7197D063948" bold="true" box="[1093,1142,1850,1875]" pageId="6" pageNumber="162">m11</emphasis>
|
||
, pair of narrow and thin ventral tensor muscles of radula (
|
||
<figureCitation id="136D2A1DC80CFFFEF013D77D7B00396C" box="[538,624,1886,1911]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="6" pageNumber="162">Fig. 20</figureCitation>
|
||
), originating at middle region of ventral surface of posterior cartilage, separated from each other close to median line, running anteriorly covering
|
||
<emphasis id="B922EA8AC80CFFFEF1E4D7A17D643980" bold="true" box="[1005,1044,1922,1947]" pageId="6" pageNumber="162">m6</emphasis>
|
||
and
|
||
<emphasis id="B922EA8AC80CFFFEF643D7A17D1F3980" bold="true" box="[1098,1135,1922,1947]" pageId="6" pageNumber="162">m8</emphasis>
|
||
, inserting in distal edge of subradular membrane at some distance beyond it towards radular ribbon, along the inner surface.
|
||
</paragraph>
|
||
<paragraph id="8BE93698C80CFFFFF2CED7E87D533C03" blockId="6.[151,1437,770,2019]" lastBlockId="7.[151,1437,151,2038]" lastPageId="7" lastPageNumber="163" pageId="6" pageNumber="162">
|
||
<emphasis id="B922EA8AC80CFFFEF2CED7E8784E39F8" box="[199,318,1995,2019]" italics="true" pageId="6" pageNumber="162">Jaw plates</emphasis>
|
||
(
|
||
<figureCitation id="136D2A1DC80CFFFEF347D7E978DB39F8" box="[334,427,1994,2019]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="6" pageNumber="162">Figs. 11</figureCitation>
|
||
,
|
||
<figureCitation id="136D2A1DC80CFFFEF3BED7E878A739F8" box="[439,471,1995,2019]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="6" pageNumber="162">17</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80CFFFEF3EBD7E9788F39F8" bold="true" box="[482,511,1994,2019]" pageId="6" pageNumber="162">jw</emphasis>
|
||
). Thick, barely triangular; cutting edge anterior, occupying most of dorsal surface of buccal cavity; separated from each other by a narrow space; central projection of jaw (
|
||
<emphasis id="B922EA8AC80DFFFFF682D0B47DD23EAB" bold="true" box="[1163,1186,151,176]" pageId="7" pageNumber="163">ja</emphasis>
|
||
) well-developed. Pair of dorsal folds (
|
||
<figureCitation id="136D2A1DC80DFFFFF350D09F78C33ECF" box="[345,435,188,213]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Fig. 17</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80DFFFFF3CAD09F78AC3ECE" bold="true" box="[451,476,188,213]" pageId="7" pageNumber="163">df</emphasis>
|
||
) initiating just posterior to jaw plates, fused with each other, encircling an area equivalent to jaws, having a distal edge wider than their base. Salivary glands absent. Anterior esophagus relatively wide, inner surface bearing a pair of tall and narrow folds (
|
||
<figureCitation id="136D2A1DC80DFFFFF127D1277AF33F07" box="[814,899,260,285]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Fig. 17</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80DFFFFF187D1277AD43F06" bold="true" box="[910,932,260,285]" pageId="7" pageNumber="163">ef</emphasis>
|
||
) as a continuation of dorsal folds, producing wide dorsal furrow. Esophageal crop (
|
||
<figureCitation id="136D2A1DC80DFFFFF036D1047BB73F5B" box="[575,711,295,320]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Figs. 15, 17</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80DFFFFF0DBD1047B9C3F5B" bold="true" box="[722,748,295,320]" pageId="7" pageNumber="163">eg</emphasis>
|
||
) occupying central region of esophagus, spanning along ~1/3 of its length; pair of longitudinal folds as continuation from those of anterior esophagus (
|
||
<emphasis id="B922EA8AC80DFFFFF673D16F7DE03F7E" bold="true" box="[1146,1168,332,357]" pageId="7" pageNumber="163">ef</emphasis>
|
||
), remaining area totally covered by small papillae; each papilla short, separated from each other by interspaces equivalent to their respective size, tip rounded. Pair of esophageal folds (
|
||
<emphasis id="B922EA8AC80DFFFFF0FCD1B77A7B3FB6" bold="true" box="[757,779,404,429]" pageId="7" pageNumber="163">ef</emphasis>
|
||
) disappearing after crop; posterior esophagus with simple, smooth inner surface. Stomach (
|
||
<figureCitation id="136D2A1DC80DFFFFF004D1947B143FCB" box="[525,612,439,464]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Fig. 16</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80DFFFFF07AD1947BF73FCB" bold="true" box="[627,647,439,464]" pageId="7" pageNumber="163">st</emphasis>
|
||
) fusiform, location and size described above; inner surface smooth; single duct to digestive glans (
|
||
<emphasis id="B922EA8AC80DFFFFF3E4D1FF7B7C3FEE" bold="true" box="[493,524,476,501]" pageId="7" pageNumber="163">dd</emphasis>
|
||
) located close to intestinal origin; stomach tapering gradually both at esophageal insertion and intestinal origin. Rectum and anus described above (pallial organs).
|
||
</paragraph>
|
||
<paragraph id="8BE93698C80DFFFFF2CED2077B653D65" blockId="7.[151,1437,151,2038]" pageId="7" pageNumber="163">
|
||
<emphasis id="B922EA8AC80DFFFFF2CED2077B063C27" box="[199,630,548,572]" italics="true" pageId="7" pageNumber="163">Non-muscular odontophore structures</emphasis>
|
||
(
|
||
<figureCitation id="136D2A1DC80DFFFFF08ED2077A633C27" box="[647,787,548,573]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Figs. 19–20</figureCitation>
|
||
).
|
||
<emphasis id="B922EA8AC80DFFFFF121D2077A323C26" bold="true" box="[808,834,548,573]" pageId="7" pageNumber="163">oc</emphasis>
|
||
, pair of anterior odontophore cartilages, ~4-times as long as wide, flat, anterior and posterior ends rounded, about same length as odontophore; fused with its respective pair at the first ~1/4 of anterior region of ventral edge (
|
||
<figureCitation id="136D2A1DC80DFFFFF139D2487AF93C9F" box="[816,905,619,644]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Fig. 19</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80DFFFFF190D2487AC03C9F" bold="true" box="[921,944,619,644]" pageId="7" pageNumber="163">of</emphasis>
|
||
);
|
||
<emphasis id="B922EA8AC80DFFFFF1C4D2487A9A3C9F" bold="true" box="[973,1002,619,644]" pageId="7" pageNumber="163">po</emphasis>
|
||
, small pair of posterior odontophore cartilages, antero-posteriorly short, length ~1/10 of anterior cartilages length, ~2/3 their width; located in posterior region, slightly lateral side of anterior cartilages, its outer side partially fused with anterior cartilages (
|
||
<figureCitation id="136D2A1DC80DFFFFF707D2907C123CD0" box="[1294,1378,691,716]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Fig. 20</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80DFFFFF764D2907CFA3CD7" bold="true" box="[1389,1418,691,716]" pageId="7" pageNumber="163">po</emphasis>
|
||
);
|
||
<emphasis id="B922EA8AC80DFFFFF29ED2F579C23CF4" bold="true" box="[151,178,726,751]" pageId="7" pageNumber="163">br</emphasis>
|
||
, subradular membrane, covering most of exposed portion of odontophore in buccal cavity (
|
||
<figureCitation id="136D2A1DC80DFFFFF701D2F57C123CF4" box="[1288,1378,726,751]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Fig. 19</figureCitation>
|
||
);
|
||
<emphasis id="B922EA8AC80DFFFFF774D2F57CE43CF4" bold="true" box="[1405,1428,726,751]" pageId="7" pageNumber="163">sc</emphasis>
|
||
, subradular cartilage, maintaining radular ribbon and expanding in buccal cavity beyond radula, covering about half of exposed portion of subradular membrane (
|
||
<figureCitation id="136D2A1DC80DFFFFF0A2D33D7A733D2C" box="[683,771,798,823]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Fig. 19</figureCitation>
|
||
). Radular sac (
|
||
<figureCitation id="136D2A1DC80DFFFFF1BED33D7D393D2C" box="[951,1097,798,823]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Figs. 15, 17</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80DFFFFF651D33D7D1F3D2C" bold="true" box="[1112,1135,798,823]" pageId="7" pageNumber="163">rs</emphasis>
|
||
) wide, projecting beyond posterior side of odontophore for approximately the same length the odontophore itself. Radular nucleus (
|
||
<emphasis id="B922EA8AC80DFFFFF73DD3627C3F3D41" bold="true" box="[1332,1359,833,858]" pageId="7" pageNumber="163">rn</emphasis>
|
||
) bifid, divided in the middle by a furrow.
|
||
</paragraph>
|
||
<paragraph id="8BE93698C80DFFFFF2CED3AA78A43B54" blockId="7.[151,1437,151,2038]" pageId="7" pageNumber="163">
|
||
<emphasis id="B922EA8AC80DFFFFF2CED3AA78683DBA" box="[199,280,905,929]" italics="true" pageId="7" pageNumber="163">Radula</emphasis>
|
||
(
|
||
<figureCitation id="136D2A1DC80DFFFFF321D3AA78C33DB9" box="[296,435,905,930]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="7" pageNumber="163">Figs. 24–29</figureCitation>
|
||
).
|
||
<emphasis id="B922EA8AC80DFFFFF3CFD3AA7B313DB9" bold="true" box="[454,577,905,930]" pageId="7" pageNumber="163">Rachidian</emphasis>
|
||
occupying ~15% of radula width, ~twice as long as wide; base rectangular, with abrupt notch preceding cutting edge (
|
||
<figureCitation id="136D2A1DC80DFFFFF07ED38E7BBB3DDE" box="[631,715,941,966]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="7" pageNumber="163">Fig. 28</figureCitation>
|
||
); cutting edge ~half of tooth height, strongly curved, triangular, pointed tip angle ~25° (
|
||
<figureCitation id="136D2A1DC80DFFFFF3A6D3F37B313DF2" box="[431,577,976,1001]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="7" pageNumber="163">Figs. 26, 29</figureCitation>
|
||
); ~10 pairs of secondary cusps along lateral border of cutting edge; each secondary cusp small, narrow, ~4-times longer than tall, tip sharply pointed, separated from each other by space equivalent to their basal width.
|
||
<emphasis id="B922EA8AC80DFFFFF009D43B7BEC3A2A" bold="true" box="[512,668,1048,1073]" pageId="7" pageNumber="163">Lateral teeth</emphasis>
|
||
five pairs in number. Similar shape as rachidian, except for having ~1/3 of its width and being slightly curved medially; apical angle ~10° (
|
||
<figureCitation id="136D2A1DC80DFFFFF1E9D41F7DEC3A4F" box="[992,1180,1084,1109]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="7" pageNumber="163">Figs. 25, 26, 29</figureCitation>
|
||
).
|
||
<emphasis id="B922EA8AC80DFFFFF6BBD41F7C173A4E" bold="true" box="[1202,1383,1084,1109]" pageId="7" pageNumber="163">Marginal teeth</emphasis>
|
||
~25 pairs, medial teeth slightly broader (as wide as lateral teeth), gradually becoming narrower towards lateral ends of radula; innermost marginal tooth with 4 large, wide cusps in subapical region of inner edge and series of minute marginal cusps along entire outer edge (
|
||
<figureCitation id="136D2A1DC80DFFFFF053D4847B933ADB" box="[602,739,1191,1216]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="7" pageNumber="163">Figs. 25, 26</figureCitation>
|
||
), 25~ times as tall as wide; second innermost tooth with 8–10 strong, thin cusps in subapical region of inner edge (
|
||
<figureCitation id="136D2A1DC80DFFFFF0E9D4E87A413AF8" box="[736,817,1227,1252]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="7" pageNumber="163">Fig. 27</figureCitation>
|
||
) and series of minute marginal cusps along entire outer edge (
|
||
<figureCitation id="136D2A1DC80DFFFFF2D7D4CD785E3B1C" box="[222,302,1262,1287]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="7" pageNumber="163">Fig 27</figureCitation>
|
||
), ~25 times as tall as wide; remaining marginal teeth as narrower versions of preceding marginal tooth; outermost teeth bearing taller, filiform secondary cusps (
|
||
<figureCitation id="136D2A1DC80DFFFFF16AD5317A9F3B30" box="[867,1007,1298,1323]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="7" pageNumber="163">Figs. 27, 29</figureCitation>
|
||
). Row of radular teeth clearly arched, with no signs of asymmetry.
|
||
</paragraph>
|
||
<paragraph id="8BE93698C80DFFFFF2CED5797D3B3B69" blockId="7.[151,1437,151,2038]" box="[199,1099,1369,1394]" pageId="7" pageNumber="163">
|
||
<emphasis id="B922EA8AC80DFFFFF2CED579781D3B6A" box="[199,365,1370,1394]" italics="true" pageId="7" pageNumber="163">Genital system</emphasis>
|
||
. Gonad described above, no additional genital structure detected.
|
||
</paragraph>
|
||
<paragraph id="8BE93698C80DFFFFF2CED55E7D4538E7" blockId="7.[151,1437,151,2038]" pageId="7" pageNumber="163">
|
||
<emphasis id="B922EA8AC80DFFFFF2CED55E78A13B8E" box="[199,465,1405,1429]" italics="true" pageId="7" pageNumber="163">Central nervous system</emphasis>
|
||
(
|
||
<figureCitation id="136D2A1DC80DFFFFF3EBD55E7B1D3B8D" box="[482,621,1405,1430]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Figs. 15, 18</figureCitation>
|
||
). Nerve ring surrounding middle level of buccal mass (
|
||
<figureCitation id="136D2A1DC80DFFFFF6ECD55E7C4A3B8D" box="[1253,1338,1405,1430]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Fig. 15</figureCitation>
|
||
); pair of pedal ganglia located just posterior to ventral base of snout, encased inside pedal musculature (
|
||
<figureCitation id="136D2A1DC80DFFFFF70FD5827C103BA1" box="[1286,1376,1441,1466]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Fig. 14</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80DFFFFF766D5827CFB3BA1" bold="true" box="[1391,1419,1441,1466]" pageId="7" pageNumber="163">nr</emphasis>
|
||
). Cerebral ganglia (
|
||
<emphasis id="B922EA8AC80DFFFFF365D5E678F53BC5" bold="true" box="[364,389,1477,1502]" pageId="7" pageNumber="163">ce</emphasis>
|
||
) broad, rather flattened, located in lateral region of buccal mass (
|
||
<figureCitation id="136D2A1DC80DFFFFF693D5E67D833BC5" box="[1178,1267,1477,1502]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Fig. 15</figureCitation>
|
||
); size of each ganglion equivalent to ~1/15 of that of buccal mass. Cerebral commissure narrow and long (~½ buccal mass width). Pleural and pedal ganglia (
|
||
<emphasis id="B922EA8AC80DFFFFF012D62F7B48383E" bold="true" box="[539,568,1548,1573]" pageId="7" pageNumber="163">gp</emphasis>
|
||
) hard to distinguish from each other; pedal ganglia widely connected to each other close do median line; each pedal ganglion with about same size as cerebral ganglion, relatively cylindrical. Cerebro-pleural and cerebro-pedal connectives relatively long, left connectives (right in
|
||
<figureCitation id="136D2A1DC80DFFFFF685D6777DAF3877" box="[1164,1247,1620,1645]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Fig. 18</figureCitation>
|
||
) slightly shorter than right connectives. Statocysts (
|
||
<emphasis id="B922EA8AC80DFFFFF02DD6547B4D388B" bold="true" box="[548,573,1655,1680]" pageId="7" pageNumber="163">sy</emphasis>
|
||
) small (size ~1/12 of each pleuro-pedal ganglion size), located side by side and close to each other; bearing several internal statoconia. Pair of buccal ganglia well-developed and individualized (
|
||
<figureCitation id="136D2A1DC80DFFFFF340D69C78A838C3" box="[329,472,1727,1752]" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1860,1882]" captionTargetBox="[189,1426,284,1786]" captionTargetId="figure@4.[151,1437,193,1838]" captionTargetPageId="4" captionText="FIGURES 12–20. Anatomy of Calliostoma melliferum; 12. head-foot, right view, anterior portion of mantle also shown; 13. same, left view; 14. head and haemocoel, ventral view, foot and columellar muscle removed; 15. foregut and nerve ring, right view; 16. pallial cavity and visceral mass, ventral view, layer of digestive gland ventral to stomach removed; 17. foregut, ventral view, longitudinally opened, odontophore deflected to right; 18. nerve ring, ventral view; 19. odontophore, dorsal view, first layer of membranes and muscles removed, both cartilages slightly deflected; 20. same, ventral view, anterior region of m4 and br removed, right-anterior portion of m11 showing its topology (scales = 1 mm)." httpUri="https://zenodo.org/record/1457659/files/figure.png" pageId="7" pageNumber="163">Figs. 15, 17</figureCitation>
|
||
:
|
||
<emphasis id="B922EA8AC80DFFFFF3ECD69C7B7338C3" bold="true" box="[485,515,1727,1752]" pageId="7" pageNumber="163">bg</emphasis>
|
||
), located at intersection between odontophore and esophageal origin, with each ganglion relatively separated from each other, size ~1/10 of cerebral ganglion size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C6513C80DFFFFF2CED7257DFF3958" pageId="7" pageNumber="163" type="distribution">
|
||
<paragraph id="8BE93698C80DFFFFF2CED7257BB83904" blockId="7.[151,1437,151,2038]" box="[199,712,1798,1823]" pageId="7" pageNumber="163">
|
||
<emphasis id="B922EA8AC80DFFFFF2CED72578103904" bold="true" box="[199,352,1798,1823]" pageId="7" pageNumber="163">Distribution.</emphasis>
|
||
Known only from
|
||
<typeStatus id="54ED883AC80DFFFFF03ED7247B173904" box="[567,615,1799,1823]" pageId="7" pageNumber="163">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="8BE93698C80DFFFFF2CED7097DFF3958" blockId="7.[151,1437,151,2038]" box="[199,1167,1834,1859]" pageId="7" pageNumber="163">
|
||
<emphasis id="B922EA8AC80DFFFFF2CED709785A3958" bold="true" box="[199,298,1834,1859]" pageId="7" pageNumber="163">Habitat.</emphasis>
|
||
Coralline bottoms, from
|
||
<quantity id="4CAE9B7DC80DFFFFF04CD7097B0E3959" box="[581,638,1834,1859]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="163" unit="m" value="60.0">60 m</quantity>
|
||
(live specimens) to
|
||
<quantity id="4CAE9B7DC80DFFFFF168D7097AD83959" box="[865,936,1834,1859]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="163" unit="m" value="260.0">260 m</quantity>
|
||
(empty shells only).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C6513C80DFFFFF2CED76D7C203991" pageId="7" pageNumber="163" type="etymology">
|
||
<paragraph id="8BE93698C80DFFFFF2CED76D7C203991" blockId="7.[151,1437,151,2038]" pageId="7" pageNumber="163">
|
||
<emphasis id="B922EA8AC80DFFFFF2CED76D783E397C" bold="true" box="[199,334,1870,1895]" pageId="7" pageNumber="163">Etymology.</emphasis>
|
||
The specific name is from Latin
|
||
<emphasis id="B922EA8AC80DFFFFF0D6D76D7A77397D" box="[735,775,1870,1894]" italics="true" pageId="7" pageNumber="163">mel</emphasis>
|
||
(“honey”) + -
|
||
<emphasis id="B922EA8AC80DFFFFF1BBD76D7AA9397D" box="[946,985,1870,1894]" italics="true" pageId="7" pageNumber="163">ifer</emphasis>
|
||
(“-bearing”), an allusion to the honey yellow coloration of the shell’s apex and cords, and also to the honeycomb-like sculpture of the protoconch.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C6513C80DFFFFF2CED7B67C3839EE" pageId="7" pageNumber="163" type="description">
|
||
<paragraph id="8BE93698C80DFFFFF2CED7B67C3839EE" blockId="7.[151,1437,151,2038]" pageId="7" pageNumber="163">
|
||
<emphasis id="B922EA8AC80DFFFFF2CED7B6789A39B5" bold="true" box="[199,490,1941,1966]" pageId="7" pageNumber="163">Measurements (in mm):</emphasis>
|
||
<typeStatus id="54ED883AC80DFFFFF3FDD7B67B2F39B5" box="[500,607,1941,1966]" pageId="7" pageNumber="163" type="holotype">Holotype</typeStatus>
|
||
, 6.75 whorls, H = 7.42, D = 6.13;
|
||
<typeStatus id="54ED883AC80DFFFFF1FBD7B57D2439B5" box="[1010,1108,1942,1966]" pageId="7" pageNumber="163" type="paratype">Paratype</typeStatus>
|
||
MZSP 55522 #1 (
|
||
<figureCitation id="136D2A1DC80DFFFFF725D7B67CFD39B5" box="[1324,1421,1941,1966]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="7" pageNumber="163">Figs 6-8</figureCitation>
|
||
), 5.75 whorls, H = 5.00, D = 4.60;
|
||
<typeStatus id="54ED883AC80DFFFFF005D7997B1E39C9" box="[524,622,1978,2002]" pageId="7" pageNumber="163" type="paratype">Paratype</typeStatus>
|
||
MZSP 67310 (
|
||
<figureCitation id="136D2A1DC80DFFFFF112D79A7A2E39C9" box="[795,862,1977,2002]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@3.[152,1435,193,1844]" captionTargetPageId="3" captionText="FIGURES 1–11. Calliostoma melliferum shells, operculum and jaw; 1. Holotype MZSP 53718, apertural view (H = 7.42 mm); 2. same, apical view (D = 6.13 mm); 3. same, abapertural view; 4. same, umbilical view. 5. Paratype MZSP 67310, apertural view (H = 9.58 mm, D = 8.13 mm). 6–8. Paratype MZSP 55522 shell with soft parts; 6. whole specimen, left lateral view (H = 5.0 mm); 7. same, apical view; 8. same, apertural view (D = 4.6 mm); 9. Holotype, protoconch detail (scale = 0.2 mm). 10. Paratype MZSP 55522 operculum; 11. same, jaw plates, ventral view (scales = 0.5 mm)." httpUri="https://zenodo.org/record/1457657/files/figure.png" pageId="7" pageNumber="163">Fig. 5</figureCitation>
|
||
), 7.25 whorls, H = 9.58, D = 8.13;
|
||
<typeStatus id="54ED883AC80DFFFFF6E3D7997C3C39C9" box="[1258,1356,1978,2002]" pageId="7" pageNumber="163" type="paratype">Paratype</typeStatus>
|
||
MZSP 70308, 5 whorls, H = 4.50, D = 4.10;
|
||
<typeStatus id="54ED883AC80DFFFFF034D7FD7BEF39ED" box="[573,671,2014,2038]" pageId="7" pageNumber="163" type="paratype">Paratype</typeStatus>
|
||
MZSP 90268 (
|
||
<figureCitation id="136D2A1DC80DFFFFF142D7FE7AA439EE" box="[843,980,2013,2038]" captionStart="FIGURES 21–29" captionStartId="5.[151,264,1865,1887]" captionTargetBox="[152,1435,193,1844]" captionTargetId="figure@5.[152,1435,193,1844]" captionTargetPageId="5" captionText="FIGURES 21–29. Calliostoma melliferum in SEM. 21. Paratype MZSP 90268, apertural view (D = 4.3 mm); 22. same, apical view (H = 4.8 mm); 23. same, protoconch detail (scale = 0.2 mm). 24–29. Radulae in SEM: 24. wide view (scale = 50 µm); 25. same, detail of right half (scale= 30 µm); 26. same, detail of central region (scale = 10 µm); 27. same, detail of right marginal teeth (scale = 10 µm); 28. another specimen, wide view, (scale = 50 µm); 29. same, detail of central region (scale = 20 µm)." httpUri="https://zenodo.org/record/1457661/files/figure.png" pageId="7" pageNumber="163">Figs. 21–23</figureCitation>
|
||
), 5.5 whorls, H = 4.80, D = 4.30.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |