treatments-xml/data/03/FF/87/03FF878E8C0910346784FB198041FBC6.xml
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<document id="668358869F986473965438AFF1E81D68" ID-DOI="10.26107/RBZ-2021-0013" ID-ISSN="2345-7600" ID-Zenodo-Dep="5352036" ID-ZooBank="C6DD0960-51EE-40C6-B948-BFAEF4E9F304" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630207527469" checkinUser="diego" docAuthor="Komai, Tomoyuki &amp; Rahayu, Dwi Listyo" docDate="2021" docId="03FF878E8C0910346784FB198041FBC6" docLanguage="en" docName="RBZ-2021-0013.pdf" docOrigin="Raffles Bulletin of Zoology 69" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Catapagurus exilidigitus Komai &amp; Rahayu 2021, new species" docType="treatment" docUuid="C49C2D29-4768-4620-AD49-7358E6830A0D" docUuidSource="ZooBank" docVersion="3" lastPageNumber="162" masterDocId="FFC6FFF68C0810326771FF848470FFF3" masterDocTitle="Three new species of the pagurid hermit crab genus Catapagurus A. Milne-Edwards, 1880 (Decapoda: Anomura: Paguroidea) from the Bohol Sea, the Philippines" masterLastPageNumber="174" masterPageNumber="156" pageNumber="157" updateTime="1699185298364" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FF0E92A80C1085E6C0C485C5575A595D">Three new species of the pagurid hermit crab genus Catapagurus A. Milne-Edwards, 1880 (Decapoda: Anomura: Paguroidea) from the Bohol Sea, the Philippines</mods:title>
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<mods:name id="3F7C3CA755F5D3236B8D79E9C3090A99" type="personal">
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<mods:namePart id="E766C7597A73E610B0253A67986C806A">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="FF94E67F246ADF5FE8F8A0E793B92441">Rahayu, Dwi Listyo</mods:namePart>
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<mods:title id="7E9E841C633CB41B2A7B39C62170269C">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="DF00D0BAE42FEA0F3E0EBD8E5441B890">2021</mods:date>
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<mods:number id="4E4E3ED50395DEF9FF12510F5989D363">2021-05-17</mods:number>
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<treatment id="03FF878E8C0910346784FB198041FBC6" LSID="urn:lsid:zoobank.org:act:C49C2D29-4768-4620-AD49-7358E6830A0D" httpUri="http://treatment.plazi.org/id/03FF878E8C0910346784FB198041FBC6" lastPageId="6" lastPageNumber="162" pageId="1" pageNumber="157">
<subSubSection id="C34C65138C0910336784FB1986EAFB46" box="[245,666,1181,1205]" pageId="1" pageNumber="157" type="nomenclature">
<paragraph id="8BE936988C0910336784FB1986EAFB46" blockId="1.[245,666,1181,1237]" box="[245,666,1181,1205]" pageId="1" pageNumber="157">
<heading id="D0A181F48C0910336784FB1986EAFB46" bold="true" box="[245,666,1181,1205]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="157" reason="2">
<emphasis id="B922EA8A8C0910336784FB1986EAFB46" bold="true" box="[245,666,1181,1205]" pageId="1" pageNumber="157">
<taxonomicName id="4C564D1B8C0910336784FB198676FB46" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[245,518,1181,1205]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="157" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0910336784FB198676FB46" bold="true" box="[245,518,1181,1205]" italics="true" pageId="1" pageNumber="157">Catapagurus exilidigitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21157F18C0910336565FB1986EAFB46" box="[532,666,1181,1205]" pageId="1" pageNumber="157" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C34C65138C09103366FDFB398672FB26" box="[396,514,1213,1237]" pageId="1" pageNumber="157" type="description">
<paragraph id="8BE936988C09103366FDFB398672FB26" blockId="1.[245,666,1181,1237]" box="[396,514,1213,1237]" pageId="1" pageNumber="157">
(
<figureCitation id="136D2A1D8C09103366E2FB39858AFB26" box="[403,506,1213,1237]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[113,148,1627,1649]" captionStartId-1="3.[136,171,1987,2009]" captionStartId-2="4.[113,148,1410,1432]" captionStartId-3="5.[136,171,1984,2006]" captionTargetBox-0="[116,1450,188,1609]" captionTargetBox-1="[136,1474,266,1972]" captionTargetBox-2="[111,1452,190,1394]" captionTargetBox-3="[129,1475,267,1968]" captionTargetId-0="figure-11@2.[92,1469,169,1620]" captionTargetId-1="figure-8@3.[115,1484,261,1989]" captionTargetId-2="figure-11@4.[99,1460,166,1405]" captionTargetId-3="figure-8@5.[118,1494,212,2011]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." captionText-1="Fig. 2. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of right cheliped, dorsal view; B, right cheliped, mesial view; C, same, lateral view; D, carpus of right cheliped, dorsal view." captionText-2="Fig. 3. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of left cheliped, dorsal view; B, left cheliped, mesial view; C, same, lateral view; D, carpus of left cheliped, dorsal view." captionText-3="Fig. 4. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, right pereopod 2, lateral view; B, same, dactylus and distal part of propodus, mesial view; C, left pereopod 3, lateral view; D, same, dactylus and distal part of propodus, mesial view." figureDoi-0="http://doi.org/10.5281/zenodo.5352038" figureDoi-1="http://doi.org/10.5281/zenodo.5352040" figureDoi-2="http://doi.org/10.5281/zenodo.5352042" figureDoi-3="http://doi.org/10.5281/zenodo.5352044" httpUri-0="https://zenodo.org/record/5352038/files/figure.png" httpUri-1="https://zenodo.org/record/5352040/files/figure.png" httpUri-2="https://zenodo.org/record/5352042/files/figure.png" httpUri-3="https://zenodo.org/record/5352044/files/figure.png" pageId="1" pageNumber="157">Figs. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C65138C09103367F9FB798532FA26" pageId="1" pageNumber="157" type="materials_examined">
<paragraph id="8BE936988C09103367F9FB798604FA86" blockId="1.[136,775,1277,1397]" pageId="1" pageNumber="157">
<emphasis id="B922EA8A8C09103367F9FB79851AFAE6" bold="true" box="[136,362,1277,1301]" pageId="1" pageNumber="157">Material examined.</emphasis>
<materialsCitation id="3B3E3CC58C0910336602FB798604FA86" collectingDate="2004-07-06" collectionCode="T" county="Sea" elevation="119" latitude="9.501667" location="Cervera" longLatPrecision="129" longitude="123.84" municipality="Pamilacan Island" pageId="1" pageNumber="157" specimenCode="T39" specimenCount="1" stateProvince="Bohol" typeStatus="holotype">
<typeStatus id="54ED883A8C0910336602FB7985AAFAE6" box="[371,474,1277,1301]" pageId="1" pageNumber="157" type="holotype">Holotype</typeStatus>
: male (sl
<quantity id="4CAE9B7D8C091033653BFB7986D2FAE6" box="[586,674,1277,1301]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="157" unit="mm" value="1.7">1.7 mm</quantity>
), NMCR 50801, PANGLAO 2004, stn
<specimenCode id="DBF09EE38C0910336696FA998666FAC6" box="[487,534,1309,1333]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="157">T39</specimenCode>
,
<collectingRegion id="4992F87A8C0910336557FA99861AFAC6" box="[550,618,1309,1333]" country="Philippines" name="Bohol" pageId="1" pageNumber="157">Bohol</collectingRegion>
<collectingCounty id="62884E148C0910336507FA9986D1FAC6" box="[630,673,1309,1333]" pageId="1" pageNumber="157">Sea</collectingCounty>
, west of
<collectingMunicipality id="6B8DACE28C09103367F9FAB9853EFAA6" box="[136,334,1341,1365]" pageId="1" pageNumber="157">Pamilacan Island</collectingMunicipality>
,
<location id="8E8960438C091033662CFAB985C5FAA6" LSID="urn:lsid:plazi:treatment:03FF878E8C0910346784FB198041FBC6:8E8960438C091033662CFAB985C5FAA6" box="[349,437,1341,1365]" county="Sea" latitude="9.501667" longLatPrecision="129" longitude="123.84" municipality="Pamilacan Island" name="Cervera" pageId="1" pageNumber="157" stateProvince="Bohol">Cervera</location>
shoal,
<geoCoordinate id="EE62505F8C091033657CFAB88603FAA6" box="[525,627,1340,1365]" degrees="9" direction="north" minutes="30.1" orientation="latitude" pageId="1" pageNumber="157" precision="92" value="9.501667">9°30.1N</geoCoordinate>
,
<geoCoordinate id="EE62505F8C09103365F2FAB88771FAA6" box="[643,769,1340,1365]" degrees="123" direction="east" minutes="50.4" orientation="longitude" pageId="1" pageNumber="157" precision="92" value="123.84">123°50.4E</geoCoordinate>
,
<quantity id="4CAE9B7D8C09103367F8FAD98576FA86" box="[137,262,1373,1397]" metricMagnitude="2" metricUnit="m" metricValue="1.19" metricValueMax="1.38" metricValueMin="1.0" pageId="1" pageNumber="157" unit="m" value="119.0" valueMax="138.0" valueMin="100.0">
<elevation id="007BD1AB8C09103367F8FAD98576FA86" box="[137,262,1373,1397]" metricMagnitude="2" metricUnit="m" metricValue="1.19" metricValueMax="1.38" metricValueMin="1.0" pageId="1" pageNumber="157" unit="m" value="119.0" valueMax="138.0" valueMin="100.0">100138 m</elevation>
</quantity>
, muddy sand, trawl,
<date id="FFE810588C091033669DFAD98600FA86" box="[492,624,1373,1397]" pageId="1" pageNumber="157" value="2004-07-06">
<collectingDate id="EFACE9B08C091033669DFAD98600FA86" box="[492,624,1373,1397]" pageId="1" pageNumber="157" value="2004-07-06">6 July 2004</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE936988C09103367F9FA1A8532FA26" blockId="1.[136,774,1437,1493]" pageId="1" pageNumber="157">
<materialsCitation id="3B3E3CC58C09103367F9FA1A854EFA26" collectingDate="2004-07-06" collectionCode="ZRC" county="Sea" elevation="119" latitude="9.501667" location="Cervera" longLatPrecision="129" longitude="123.84" municipality="Pamilacan Island" pageId="1" pageNumber="157" specimenCode="ZRC 2021.0413" specimenCount="2" specimenCount-male="2" stateProvince="Bohol" typeStatus="paratype">
<typeStatus id="54ED883A8C09103367F9FA1A8484FA45" box="[136,244,1438,1462]" pageId="1" pageNumber="157" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D50FD118C091033678EFA198522FA46" box="[255,338,1437,1461]" pageId="1" pageNumber="157" type="male">2 males</specimenCount>
(sl 1.5,
<quantity id="4CAE9B7D8C09103366DAFA198671FA46" box="[427,513,1437,1461]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="157" unit="mm" value="1.8">1.8 mm</quantity>
),
<specimenCode id="DBF09EE38C0910336565FA1986B4FA46" box="[532,708,1437,1461]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="157">ZRC 2021.0413</specimenCode>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C65138C09103467F9FA7985A6FAA6" lastPageId="6" lastPageNumber="162" pageId="1" pageNumber="157" type="description">
<paragraph id="8BE936988C09103367F9FA7985CDF886" blockId="1.[134,776,1533,1909]" pageId="1" pageNumber="157">
<emphasis id="B922EA8A8C09103367F9FA79857FF9E6" bold="true" box="[136,271,1533,1557]" pageId="1" pageNumber="157">Description.</emphasis>
Shield (
<figureCitation id="136D2A1D8C0910336612FA7985C3F9E6" box="[355,435,1533,1557]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="1" pageNumber="157">Fig. 1A</figureCitation>
) approximately as long as broad; anterior margin between rostral lobe and lateral projections concave; anterolateral margins sloping; posterior margin roundly truncate; dorsal surface with longitudinal rows of tufts of short setae posterior to lateral projections; carapace lateral lobes very narrow, well calcified. Rostral lobe rounded, produced as far as lateral projections. Lateral projections broadly triangular, each with terminal, submarginal spinule. Posterior carapace 0.6 length of shield; posterolateral plates moderately wide anteriorly, drawn out into relatively thin bands reaching to posterior margins, posterior median plate short, only weakly calcified.
</paragraph>
<paragraph id="8BE936988C09103367F9F81985E8F7C6" blockId="1.[135,776,1949,2101]" pageId="1" pageNumber="157">
Ocular peduncles (
<figureCitation id="136D2A1D8C0910336624F81985D9F846" box="[341,425,1949,1973]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="1" pageNumber="157">Fig. 1A</figureCitation>
) moderately short (0.8 times as long as shield), stout, widened distally; corneal width 0.6 of peduncular length. Ocular acicles narrow, spike-like, slender, reaching 0.4 length of ocular peduncles, terminating acutely; widely separated basally.
</paragraph>
<paragraph id="8BE936988C0910336432FF398037FE46" blockId="1.[834,1475,189,437]" pageId="1" pageNumber="157">
Antennular peduncle (
<figureCitation id="136D2A1D8C091033635AFF3980E9FF26" box="[1067,1177,189,213]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="1" pageNumber="157">Fig. 1A, B</figureCitation>
) overreaching distal corneal margin by full length of ultimate article. Ultimate article 0.9 times as long as shield, 5.3 times as long as distal height, with tuft of 7 long setae at dorsodistal lateral margin and with row of some short setae on dorsal surface. Penultimate article almost glabrous. Basal article with slightly produced, acute ventrodistal margin; statocyst lobe weakly inflated in proximal half of article.
</paragraph>
<paragraph id="8BE936988C0910336432FE59815DFD06" blockId="1.[834,1476,477,757]" pageId="1" pageNumber="157">
Antennal peduncle (
<figureCitation id="136D2A1D8C0910336356FE5980F0FE06" box="[1063,1152,477,501]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="1" pageNumber="157">Fig. 1A</figureCitation>
) overreaching distal corneal margin by 0.20.3 length of article 5. Articles 5 and 4 with few setae. Article 3 with few setae on unarmed ventromesial distal angle. Article 2 with produced, spiniform dorsolateral distal angle reaching midlength of article 4; dorsomesial distal angle with small spine. Article 1 with small spine at dorsolateral distal angle. Antennal acicle reaching midlength of article 5 and overreaching corneal base, terminating in spine. Flagella broken off and not preserved.
</paragraph>
<paragraph id="8BE936988C0910336432FC9987DEFC66" blockId="1.[835,1476,797,917]" pageId="1" pageNumber="157">
Maxilliped 3 endopod moderately stout; merus with small dorsodistal spine; ischium with crista dentata consisting of 6 small teeth; accessory tooth small, subdistal in position (
<figureCitation id="136D2A1D8C091033643AFCF987D1FC66" box="[843,929,893,917]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="1" pageNumber="157">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="8BE936988C0910336432FC3980EFF846" blockId="1.[834,1475,957,1973]" pageId="1" pageNumber="157">
Right cheliped (
<figureCitation id="136D2A1D8C0910336484FC39801CFC26" box="[1013,1132,957,981]" captionStart="Fig" captionStartId="3.[136,171,1987,2009]" captionTargetBox="[136,1474,266,1972]" captionTargetId="figure-8@3.[115,1484,261,1989]" captionTargetPageId="3" captionText="Fig. 2. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of right cheliped, dorsal view; B, right cheliped, mesial view; C, same, lateral view; D, carpus of right cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352040" httpUri="https://zenodo.org/record/5352040/files/figure.png" pageId="1" pageNumber="157">Fig. 2AD</figureCitation>
) long, moderately stout. Chela (
<figureCitation id="136D2A1D8C091033643DFC5987D6FC06" box="[844,934,989,1013]" captionStart="Fig" captionStartId="3.[136,171,1987,2009]" captionTargetBox="[136,1474,266,1972]" captionTargetId="figure-8@3.[115,1484,261,1989]" captionTargetPageId="3" captionText="Fig. 2. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of right cheliped, dorsal view; B, right cheliped, mesial view; C, same, lateral view; D, carpus of right cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352040" httpUri="https://zenodo.org/record/5352040/files/figure.png" pageId="1" pageNumber="157">Fig. 2A</figureCitation>
) elongate subovate, 2.6 times as long as wide. Dactylus 0.7 length of palm; weakly convex dorsal surface almost smooth and with tufts of setae dorsomesially; dorsomesial margin not delimited, rounded; mesial face with scattered granules; ventral surface with some tufts of long setae; occlusal margin with 2 obtuse calcareous teeth, slightly overlapped by fixed finger. Palm subequal in length to carpus, 1.6 times as long as wide; dorsomesial and dorsolateral margins not delimited, convex dorsal surface almost smooth except for granulate dorsolateral part; lateral and mesial faces granulate; ventral surface gently convex, with scattered tufts of moderately long setae; fixed finger with 2 rows of short setae on dorsal surface, occlusal margin with low, broad calcareous tooth proximal to midlength, unarmed in distal half. Carpus (
<figureCitation id="136D2A1D8C09103363E8FA19809DFA46" box="[1177,1261,1437,1461]" captionStart="Fig" captionStartId="3.[136,171,1987,2009]" captionTargetBox="[136,1474,266,1972]" captionTargetId="figure-8@3.[115,1484,261,1989]" captionTargetPageId="3" captionText="Fig. 2. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of right cheliped, dorsal view; B, right cheliped, mesial view; C, same, lateral view; D, carpus of right cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352040" httpUri="https://zenodo.org/record/5352040/files/figure.png" pageId="1" pageNumber="157">Fig. 2D</figureCitation>
) subequal in length to merus, widened distally, 2.2 times as long as distal width; dorsomesial and dorsolateral margins each delimited with row of tiny spines, strongest distally, and with few bristlelike setae proximally, dorsal surface almost smooth with few granules; lateral surface with scattered granules; mesial face almost smooth except for some granules adjacent to dorsal margin, ventromesial distal angle with tiny spine; ventral surface gently convex, smooth. Merus subtriangular in cross section; dorsodistal margin without spine, but with few bristle-like setae; dorsal surface with widely spaced stiff setae; lateral surface with sparse granules, gently convex ventrolateral margin without conspicuous spines; mesial surface almost glabrous except for few short stiff setae, weakly convex ventromesial margin with small distal spine; ventral surface with sparse granules. Ischium unarmed. Coxa unarmed on ventromesial angle.
</paragraph>
<paragraph id="8BE936988C0910366432F85986A9F9D7" blockId="1.[834,1475,2013,2101]" lastBlockId="4.[113,752,1516,1572]" lastPageId="4" lastPageNumber="160" pageId="1" pageNumber="157">
Left cheliped (
<figureCitation id="136D2A1D8C09103364A8F859803AF806" box="[985,1098,2013,2037]" captionStart="Fig" captionStartId="4.[113,148,1410,1432]" captionTargetBox="[111,1452,190,1394]" captionTargetId="figure-11@4.[99,1460,166,1405]" captionTargetPageId="4" captionText="Fig. 3. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of left cheliped, dorsal view; B, left cheliped, mesial view; C, same, lateral view; D, carpus of left cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352042" httpUri="https://zenodo.org/record/5352042/files/figure.png" pageId="1" pageNumber="157">Fig. 3AD</figureCitation>
) slender, shorter than right cheliped; dactyl and fixed finger slightly curved ventrally. Chela (
<figureCitation id="136D2A1D8C09103362E4F8798714F7C6" captionStart="Fig" captionStartId="4.[113,148,1410,1432]" captionTargetBox="[111,1452,190,1394]" captionTargetId="figure-11@4.[99,1460,166,1405]" captionTargetPageId="4" captionText="Fig. 3. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of left cheliped, dorsal view; B, left cheliped, mesial view; C, same, lateral view; D, carpus of left cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352042" httpUri="https://zenodo.org/record/5352042/files/figure.png" pageId="1" pageNumber="157">Fig. 3A</figureCitation>
) 5.0 times as long as wide. Dactylus 1.5 times as long as palm, terminating in small, curved corneous claw; surfaces almost smooth, with sparse setae; dorsomesial margin not delimited; occlusal margin with row of minute corneous teeth almost over entire length. Palm 0.5 length of carpus, 2.0 times as long as wide; dorsomesial and dorsolateral margins not delimited, transversely convex dorsal surface with sparse granules; ventral surface slightly convex, smooth; fixed finger smooth on surfaces, terminating in small, curved corneous claw, with sparse short setae; occlusal margin with row of minute corneous teeth. Carpus (
<figureCitation id="136D2A1D8C0A10306690F79B864BF7C4" box="[481,571,2079,2103]" captionStart="Fig" captionStartId="4.[113,148,1410,1432]" captionTargetBox="[111,1452,190,1394]" captionTargetId="figure-11@4.[99,1460,166,1405]" captionTargetPageId="4" captionText="Fig. 3. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of left cheliped, dorsal view; B, left cheliped, mesial view; C, same, lateral view; D, carpus of left cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352042" httpUri="https://zenodo.org/record/5352042/files/figure.png" pageId="2" pageNumber="158">Fig. 3D</figureCitation>
) slightly longer than merus; dorsomesial and dorsolateral margins each with row of tiny spines, strongest distally, dorsal surface almost smooth; lateral surface with sparse granules; mesial surface nearly smooth except for granulate ventral part, granules extending onto slightly convex ventral surface. Merus dorsodistal margin unarmed; dorsal surface with several setae; lateral surface with sparse granules, particularly on ventral part, gently convex ventral margin unarmed; mesial surface almost smooth, with row of stiff setae adjacent to ventromesial margin, gently convex ventromesial margin with 3 tiny spines on distal margin; ventral surface granulates. Ischium unarmed. Coxa unarmed on ventromesial angle.
</paragraph>
<caption id="DF2966108C0A10306700F9DF871BF93B" ID-DOI="http://doi.org/10.5281/zenodo.5352038" ID-Zenodo-Dep="5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="2" pageNumber="158" startId="2.[113,148,1627,1649]" targetBox="[116,1450,188,1609]" targetPageId="2">
<paragraph id="8BE936988C0A10306700F9DF871BF93B" blockId="2.[113,1451,1627,1736]" pageId="2" pageNumber="158">
Fig. 1.
<taxonomicName id="4C564D1B8C0A103067C8F9D885D9F983" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[185,425,1627,1649]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="158" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0A103067C8F9D885D9F983" box="[185,425,1627,1649]" italics="true" pageId="2" pageNumber="158">Catapagurus exilidigitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21157F18C0A103066C6F9D8865FF982" box="[439,559,1627,1649]" pageId="2" pageNumber="158" rank="species">new species</taxonomicNameLabel>
, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view.
</paragraph>
</caption>
<caption id="DF2966108C0B103167F9F84787A7F805" ID-DOI="http://doi.org/10.5281/zenodo.5352040" ID-Zenodo-Dep="5352040" httpUri="https://zenodo.org/record/5352040/files/figure.png" pageId="3" pageNumber="159" startId="3.[136,171,1987,2009]" targetBox="[136,1474,266,1972]" targetPageId="3">
<paragraph id="8BE936988C0B103167F9F84787A7F805" blockId="3.[136,1474,1987,2038]" pageId="3" pageNumber="159">
Fig. 2.
<taxonomicName id="4C564D1B8C0B103167BFF84785CDF82B" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[206,445,1987,2008]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0B103167BFF84785CDF82B" box="[206,445,1987,2008]" italics="true" pageId="3" pageNumber="159">Catapagurus exilidigitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21157F18C0B103166B8F8478630F82A" box="[457,576,1987,2009]" pageId="3" pageNumber="159" rank="species">new species</taxonomicNameLabel>
, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of right cheliped, dorsal view; B, right cheliped, mesial view; C, same, lateral view; D, carpus of right cheliped, dorsal view.
</paragraph>
</caption>
<caption id="DF2966108C0C10366700FA0687C1FA46" ID-DOI="http://doi.org/10.5281/zenodo.5352042" ID-Zenodo-Dep="5352042" httpUri="https://zenodo.org/record/5352042/files/figure.png" pageId="4" pageNumber="160" startId="4.[113,148,1410,1432]" targetBox="[111,1452,190,1394]" targetPageId="4">
<paragraph id="8BE936988C0C10366700FA0687C1FA46" blockId="4.[113,1451,1410,1461]" pageId="4" pageNumber="160">
Fig. 3.
<taxonomicName id="4C564D1B8C0C103667C8FA0685D9FA64" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[185,425,1410,1431]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="160" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0C103667C8FA0685D9FA64" box="[185,425,1410,1431]" italics="true" pageId="4" pageNumber="160">Catapagurus exilidigitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21157F18C0C103666C6FA07865FFA6B" box="[439,559,1410,1432]" pageId="4" pageNumber="160" rank="species">new species</taxonomicNameLabel>
, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of left cheliped, dorsal view; B, left cheliped, mesial view; C, same, lateral view; D, carpus of left cheliped, dorsal view.
</paragraph>
</caption>
<paragraph id="8BE936988C0C10366700F9C88080F977" blockId="4.[111,753,1612,2116]" lastBlockId="4.[813,1452,1516,1668]" pageId="4" pageNumber="160">
Pereopods 2 (
<figureCitation id="136D2A1D8C0C10366660F9C885E0F997" box="[273,400,1612,1636]" captionStart="Fig" captionStartId="5.[136,171,1984,2006]" captionTargetBox="[129,1475,267,1968]" captionTargetId="figure-8@5.[118,1494,212,2011]" captionTargetPageId="5" captionText="Fig. 4. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, right pereopod 2, lateral view; B, same, dactylus and distal part of propodus, mesial view; C, left pereopod 3, lateral view; D, same, dactylus and distal part of propodus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5352044" httpUri="https://zenodo.org/record/5352044/files/figure.png" pageId="4" pageNumber="160">Fig. 4A, B</figureCitation>
) slender, overreaching tips of outstretched chelipeds. Dactyli (
<figureCitation id="136D2A1D8C0C103666B4F9E88666F977" box="[453,534,1644,1668]" captionStart="Fig" captionStartId="5.[136,171,1984,2006]" captionTargetBox="[129,1475,267,1968]" captionTargetId="figure-8@5.[118,1494,212,2011]" captionTargetPageId="5" captionText="Fig. 4. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, right pereopod 2, lateral view; B, same, dactylus and distal part of propodus, mesial view; C, left pereopod 3, lateral view; D, same, dactylus and distal part of propodus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5352044" httpUri="https://zenodo.org/record/5352044/files/figure.png" pageId="4" pageNumber="160">Fig. 4B</figureCitation>
) very slender (about 23 times as long as wide), 1.4 times as long as propodi; in dorsal view, straight; in lateral view, slightly curved ventrally; dorsal margins each with row of moderately spaced bristlelike or spiniform setae, becoming longer distally; mesial faces each with row of 9 moderately spaced, spiniform or bristle-like setae extending near ventral margin to midline toward distal; lateral surfaces and ventral margins unarmed. Propodi slightly more than twice length of carpi; dorsal and ventral margins granulate, former with few stiff setae subdistally; lateral and mesial surfaces smooth; ventrodistal margin with 1 spiniform seta. Carpi short, about half-length of meri; dorsal surfaces each with row of spinules, strongest distally; lateral surface with few granules, mesial and ventral surfaces almost smooth. Meri each with 1 small dorsodistal spine; dorsal surface with few low transverse ridges and moderately spaced bristle-like setae, but without spines; lateral and mesial surfaces smooth except for granulate ventral parts; ventral surfaces granulate, ventrolateral distal angle with minute spine. Ischia unarmed.
</paragraph>
<paragraph id="8BE936988C0C1034645CF928863BFE86" blockId="4.[811,1453,1708,2116]" lastBlockId="6.[112,754,189,373]" lastPageId="6" lastPageNumber="162" pageId="4" pageNumber="160">
Pereopods 3 (
<figureCitation id="136D2A1D8C0C103664B0F9288047F937" box="[961,1079,1708,1732]" captionStart="Fig" captionStartId="5.[136,171,1984,2006]" captionTargetBox="[129,1475,267,1968]" captionTargetId="figure-8@5.[118,1494,212,2011]" captionTargetPageId="5" captionText="Fig. 4. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, right pereopod 2, lateral view; B, same, dactylus and distal part of propodus, mesial view; C, left pereopod 3, lateral view; D, same, dactylus and distal part of propodus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5352044" httpUri="https://zenodo.org/record/5352044/files/figure.png" pageId="4" pageNumber="160">Fig. 4C, D</figureCitation>
) generally similar to pereopods 2. Dactyli (
<figureCitation id="136D2A1D8C0C103664E1F9488798F917" box="[912,1000,1740,1764]" captionStart="Fig" captionStartId="5.[136,171,1984,2006]" captionTargetBox="[129,1475,267,1968]" captionTargetId="figure-8@5.[118,1494,212,2011]" captionTargetPageId="5" captionText="Fig. 4. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, right pereopod 2, lateral view; B, same, dactylus and distal part of propodus, mesial view; C, left pereopod 3, lateral view; D, same, dactylus and distal part of propodus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5352044" httpUri="https://zenodo.org/record/5352044/files/figure.png" pageId="4" pageNumber="160">Fig. 4D</figureCitation>
) 26 times as long as wide, 1.5 times as long as propodi; dorsal margins each with row of moderately spaced bristle-like or spiniform setae, becoming longer distally; mesial faces each with row of about 10 moderately spaced spiniform setae extending near ventral margin to midline toward distal; lateral surfaces and ventral margins unarmed. Propodi about twice length of carpi; dorsal margins faintly granulate, ventral margins nearly smooth; lateral and mesial surfaces also smooth; ventrodistal margin with 1 spiniform seta. Carpi short, about half-length of meri; dorsal surfaces each with row of spinules, strongest distally; lateral, mesial and ventral surfaces almost smooth. Meri unarmed on dorsodistal margin, but with bristle-like seta; dorsal surface with 2 low transverse ridges and bristle-like setae subdistally, remainder with irregular rows of bristlelike or stiff setae; lateral and mesial surfaces smooth; ventral surfaces granulate, ventrolateral distal angle unarmed. Ischia longer than those of pereopods 2, unarmed.
</paragraph>
<caption id="DF2966108C0D103767F9F84481DAF800" ID-DOI="http://doi.org/10.5281/zenodo.5352044" ID-Zenodo-Dep="5352044" httpUri="https://zenodo.org/record/5352044/files/figure.png" pageId="5" pageNumber="161" startId="5.[136,171,1984,2006]" targetBox="[129,1475,267,1968]" targetPageId="5">
<paragraph id="8BE936988C0D103767F9F84481DAF800" blockId="5.[135,1474,1984,2035]" pageId="5" pageNumber="161">
Fig. 4.
<taxonomicName id="4C564D1B8C0D103767B8F84485C6F826" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[201,438,1984,2005]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="161" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0D103767B8F84485C6F826" box="[201,438,1984,2005]" italics="true" pageId="5" pageNumber="161">Catapagurus exilidigitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21157F18C0D103766B3F8458646F825" box="[450,566,1984,2006]" pageId="5" pageNumber="161" rank="species">new species</taxonomicNameLabel>
, holotype, male (sl 1.7 mm), NMCR 50801. A, right pereopod 2, lateral view; B, same, dactylus and distal part of propodus, mesial view; C, left pereopod 3, lateral view; D, same, dactylus and distal part of propodus, mesial view.
</paragraph>
</caption>
<paragraph id="8BE936988C0E10346700FE1984AAFDC6" blockId="6.[112,753,413,565]" pageId="6" pageNumber="162">
Pereopods 4 (
<figureCitation id="136D2A1D8C0E10346671FE198522FE46" box="[256,338,413,437]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="6" pageNumber="162">Fig. 1D</figureCitation>
) weakly semichelate; dactyli each with well developed preungual process far extending beyond tip of corneous claw; propodal rasp consisting of single row of corneous scales on distal 0.8 of ventral margin. Pereopods 5 chelate.
</paragraph>
<paragraph id="8BE936988C0E10346700FDD986FCFD06" blockId="6.[113,752,605,758]" pageId="6" pageNumber="162">
Males with long right sexual tube (right pereopod 5 coxa) curved dorsally over pleon and reaching pleonal midline, terminus simple, with few minute setae (
<figureCitation id="136D2A1D8C0E10346520FD1986D9FD46" box="[593,681,669,693]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="6" pageNumber="162">Fig. 1F</figureCitation>
). Left pereopod 5 coxa with small, papilla-like protrusion from gonopore, circumscribed by short setae (
<figureCitation id="136D2A1D8C0E1034655DFD59860FFD06" box="[556,639,733,757]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="6" pageNumber="162">Fig. 1F</figureCitation>
).
</paragraph>
<paragraph id="8BE936988C0E10346700FC998632FC46" blockId="6.[113,753,797,949]" pageId="6" pageNumber="162">
Thoracic sternite 6 (pereopods 3) (
<figureCitation id="136D2A1D8C0E10346690FC998643FCC6" box="[481,563,797,821]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="6" pageNumber="162">Fig. 1G</figureCitation>
) with moderately wide, subsemicircular anterior lobe, much shorter than large posterior lobe; posterior lobe distinctly longer than anterior lobe. Sternite 8 (pereopods 5) (
<figureCitation id="136D2A1D8C0E103466BCFCF98650FC66" box="[461,544,893,917]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="6" pageNumber="162">Fig. 1E</figureCitation>
) consisting of two small rounded lobes moderately separated.
</paragraph>
<paragraph id="8BE936988C0E10346700FC5985FFFBC6" blockId="6.[112,752,989,1077]" pageId="6" pageNumber="162">Pleon in males with small, uniramous unpaired left pleopods 3 and 4, no pleopod 5; females unknown. Uropodal protopods not protruding posteriorly.</paragraph>
<paragraph id="8BE936988C0E10346700FBD984B4FAE6" blockId="6.[113,752,1117,1301]" pageId="6" pageNumber="162">
Telson (
<figureCitation id="136D2A1D8C0E103467A5FBD98541FB86" box="[212,305,1117,1141]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="6" pageNumber="162">Fig. 1H</figureCitation>
) with lateral indentations suggesting separation of anterior and posterior portions; triangular posterior lobes separated by moderately deep U-shaped median cleft, each terminating in subacute tip; oblique terminal margins unarmed, with few minute setae on right margin.
</paragraph>
<paragraph id="8BE936988C0E10346700FAB985A6FAA6" blockId="6.[113,470,1341,1365]" box="[113,470,1341,1365]" pageId="6" pageNumber="162">
<emphasis id="B922EA8A8C0E10346700FAB9853DFAA6" bold="true" box="[113,333,1341,1365]" pageId="6" pageNumber="162">Colouration in life.</emphasis>
<collectingRegion id="4992F87A8C0E10346627FABA850EFAA6" box="[342,382,1342,1365]" country="Maldives" name="Uthuru" pageId="6" pageNumber="162">Not</collectingRegion>
known.
</paragraph>
</subSubSection>
<subSubSection id="C34C65138C0E10346700FAF981DCFC46" pageId="6" pageNumber="162" type="distribution">
<paragraph id="8BE936988C0E10346700FAF98625FA46" blockId="6.[113,752,1405,1461]" pageId="6" pageNumber="162">
<emphasis id="B922EA8A8C0E10346700FAF98576FA66" bold="true" box="[113,262,1405,1429]" pageId="6" pageNumber="162">Distribution.</emphasis>
Presently known only from the
<collectingRegion id="4992F87A8C0E10346507FAF986C8FA66" box="[630,696,1405,1429]" country="Philippines" name="Bohol" pageId="6" pageNumber="162">Bohol</collectingRegion>
Sea, central
<collectingCountry id="F34176088C0E103467B2FA19854EFA46" box="[195,318,1437,1461]" name="Philippines" pageId="6" pageNumber="162">Philippines</collectingCountry>
, at depths of
<quantity id="4CAE9B7D8C0E103466A2FA198621FA46" box="[467,593,1437,1461]" metricMagnitude="2" metricUnit="m" metricValue="1.19" metricValueMax="1.38" metricValueMin="1.0" pageId="6" pageNumber="162" unit="m" value="119.0" valueMax="138.0" valueMin="100.0">
100
<specimenCount id="9D50FD118C0E10346578FA198621FA46" box="[521,593,1437,1461]" pageId="6" pageNumber="162" type="generic">138 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="8BE936988C0E1034645CFF3981DCFC46" blockId="6.[811,1454,189,949]" pageId="6" pageNumber="162">
Takeda, 2006;
<bibRefCitation id="EFC74B698C0E103464BCFF39817EFF26" author="McLaughlin PA &amp; Rahayu DL" box="[973,1294,189,213]" pageId="6" pageNumber="162" pagination="899 - 916" refId="ref10249" refString="McLaughlin PA &amp; Rahayu DL (2008) Pteropagurus and Catapagurus (Decapoda, Anomura, Paguridae): resource sharing or &quot; any port in a storm &quot;? Zoosystema, 30 (4): 899 - 916." type="journal article" year="2008">McLaughlin &amp; Rahayu, 2008</bibRefCitation>
).
<taxonomicName id="4C564D1B8C0E10346250FF3A87D4FF06" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0E10346250FF3A87D4FF06" italics="true" pageId="6" pageNumber="162">Catapagurus exilidigitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21157F18C0E103464C0FF5A8044FF06" box="[945,1076,221,245]" pageId="6" pageNumber="162" rank="species">new species</taxonomicNameLabel>
, is primarily distinguished from
<taxonomicName id="4C564D1B8C0E103462E2FF5A87B4FEE6" authorityName="McLaughlin" authorityYear="1997" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="tanimbarensis">
<emphasis id="B922EA8A8C0E103462E2FF5A87B4FEE6" italics="true" pageId="6" pageNumber="162">C. tanimbarensis</emphasis>
</taxonomicName>
by the telson with a wide, U-shaped median cleft (
<figureCitation id="136D2A1D8C0E10346418FE9987C9FEC6" box="[873,953,285,309]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="6" pageNumber="162">Fig. 1H</figureCitation>
) (versus telson with a narrow, slit-like V-shaped median cleft;
<bibRefCitation id="EFC74B698C0E103464BCFEB980D5FEA6" author="McLaughlin PA" box="[973,1189,317,341]" pageId="6" pageNumber="162" pagination="433 - 572" refId="ref9882" refString="McLaughlin PA (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier A &amp; Bouchet P (eds.) Resultats des Campagnes MUSORSTOM, Volume 16. Memoires du Museum national d'Histoire naturelle, Paris, 172: 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
: fig. 18i, k, m). Other differentiating characters are: (1) the ultimate article of the antennular peduncle is relatively longer in
<taxonomicName id="4C564D1B8C0E10346390FEFA8102FE66" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[1249,1394,381,405]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0E10346390FEFA8102FE66" box="[1249,1394,381,405]" italics="true" pageId="6" pageNumber="162">C. exilidigitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21157F18C0E1034620EFEFA870BFE46" pageId="6" pageNumber="162" rank="species">new species</taxonomicNameLabel>
, than in
<taxonomicName id="4C564D1B8C0E10346491FE1A80E8FE46" authorityName="McLaughlin" authorityYear="1997" box="[992,1176,413,437]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="tanimbarensis">
<emphasis id="B922EA8A8C0E10346491FE1A80E8FE46" box="[992,1176,413,437]" italics="true" pageId="6" pageNumber="162">C. tanimbarensis</emphasis>
</taxonomicName>
(0.9 times as long as the shield versus about 0.7 times) (
<figureCitation id="136D2A1D8C0E1034631CFE3980A9FE26" box="[1133,1241,445,469]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="6" pageNumber="162">Fig. 1A, B</figureCitation>
versus
<bibRefCitation id="EFC74B698C0E10346257FE398714FE06" author="McLaughlin PA" pageId="6" pageNumber="162" pagination="433 - 572" refId="ref9882" refString="McLaughlin PA (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier A &amp; Bouchet P (eds.) Resultats des Campagnes MUSORSTOM, Volume 16. Memoires du Museum national d'Histoire naturelle, Paris, 172: 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
: fig. 18a, b); the meri of chelipeds are unarmed on the dorsodistal margins in
<taxonomicName id="4C564D1B8C0E10346354FE7A80CAFDE6" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[1061,1210,509,533]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0E10346354FE7A80CAFDE6" box="[1061,1210,509,533]" italics="true" pageId="6" pageNumber="162">C. exilidigitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21157F18C0E103463B8FE7A813AFDE6" box="[1225,1354,509,533]" pageId="6" pageNumber="162" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="136D2A1D8C0E1034622BFE7981D6FDE6" box="[1370,1446,509,533]" captionStart="Fig" captionStartId="3.[136,171,1987,2009]" captionTargetBox="[136,1474,266,1972]" captionTargetId="figure-8@3.[115,1484,261,1989]" captionTargetPageId="3" captionText="Fig. 2. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of right cheliped, dorsal view; B, right cheliped, mesial view; C, same, lateral view; D, carpus of right cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352040" httpUri="https://zenodo.org/record/5352040/files/figure.png" pageId="6" pageNumber="162">Figs. 2</figureCitation>
,
<figureCitation id="136D2A1D8C0E1034645FFD99874DFDC6" box="[814,829,541,565]" captionStart="Fig" captionStartId="4.[113,148,1410,1432]" captionTargetBox="[111,1452,190,1394]" captionTargetId="figure-11@4.[99,1460,166,1405]" captionTargetPageId="4" captionText="Fig. 3. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, chela of left cheliped, dorsal view; B, left cheliped, mesial view; C, same, lateral view; D, carpus of left cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352042" httpUri="https://zenodo.org/record/5352042/files/figure.png" pageId="6" pageNumber="162">3</figureCitation>
), whereas armed with one or more dorsodistal spine(s) in
<taxonomicName id="4C564D1B8C0E1034643BFDBA8072FDA6" authorityName="McLaughlin" authorityYear="1997" box="[842,1026,573,597]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="tanimbarensis">
<emphasis id="B922EA8A8C0E1034643BFDBA8072FDA6" box="[842,1026,573,597]" italics="true" pageId="6" pageNumber="162">C. tanimbarensis</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFC74B698C0E10346349FDB9814CFDA6" author="McLaughlin PA" box="[1080,1340,573,597]" pageId="6" pageNumber="162" pagination="433 - 572" refId="ref9882" refString="McLaughlin PA (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier A &amp; Bouchet P (eds.) Resultats des Campagnes MUSORSTOM, Volume 16. Memoires du Museum national d'Histoire naturelle, Paris, 172: 433 - 572." type="journal article" year="1997">McLaughlin, 1997: 499</bibRefCitation>
; fig. 18c); the ambulatory dactyli are more elongate and slender in
<taxonomicName id="4C564D1B8C0E103462E5FDDA87D4FD66" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0E103462E5FDDA87D4FD66" italics="true" pageId="6" pageNumber="162">C. exilidigitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21157F18C0E103464C3FDFA8046FD66" box="[946,1078,637,661]" pageId="6" pageNumber="162" rank="species">new species</taxonomicNameLabel>
, than in
<taxonomicName id="4C564D1B8C0E103463E5FDFA813CFD66" authorityName="McLaughlin" authorityYear="1997" box="[1172,1356,637,661]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="tanimbarensis">
<emphasis id="B922EA8A8C0E103463E5FDFA813CFD66" box="[1172,1356,637,661]" italics="true" pageId="6" pageNumber="162">C. tanimbarensis</emphasis>
</taxonomicName>
(1.41.5 times as long as the propodus; more than 20 times as long as wide in
<taxonomicName id="4C564D1B8C0E103464DAFD3A8036FD26" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[939,1094,701,725]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0E103464DAFD3A8036FD26" box="[939,1094,701,725]" italics="true" pageId="6" pageNumber="162">C. exilidigitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21157F18C0E10346326FD3A80AFFD26" box="[1111,1247,701,725]" pageId="6" pageNumber="162" rank="species">new species</taxonomicNameLabel>
, cf.
<figureCitation id="136D2A1D8C0E10346262FD398127FD26" box="[1299,1367,701,725]" captionStart="Fig" captionStartId="5.[136,171,1984,2006]" captionTargetBox="[129,1475,267,1968]" captionTargetId="figure-8@5.[118,1494,212,2011]" captionTargetPageId="5" captionText="Fig. 4. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, right pereopod 2, lateral view; B, same, dactylus and distal part of propodus, mesial view; C, left pereopod 3, lateral view; D, same, dactylus and distal part of propodus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5352044" httpUri="https://zenodo.org/record/5352044/files/figure.png" pageId="6" pageNumber="162">Fig. 4</figureCitation>
; versus 1.01.25 times as long as the propodus and less than 15 times as long as wide in
<taxonomicName id="4C564D1B8C0E10346487FD7A80DDFCE6" authorityName="McLaughlin" authorityYear="1997" box="[1014,1197,765,789]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="tanimbarensis">
<emphasis id="B922EA8A8C0E10346487FD7A80DDFCE6" box="[1014,1197,765,789]" italics="true" pageId="6" pageNumber="162">C. tanimbarensis</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFC74B698C0E103463ACFD7981D8FCE6" author="McLaughlin PA" box="[1245,1448,765,789]" pageId="6" pageNumber="162" pagination="433 - 572" refId="ref9882" refString="McLaughlin PA (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier A &amp; Bouchet P (eds.) Resultats des Campagnes MUSORSTOM, Volume 16. Memoires du Museum national d'Histoire naturelle, Paris, 172: 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
: fig. 18df;
<bibRefCitation id="EFC74B698C0E103464DBFC99803DFCC6" author="Asakura A" box="[938,1101,797,821]" pageId="6" pageNumber="162" pagination="823 - 891" refId="ref9098" refString="Asakura A (2001) A revision of the hermit crab genera Catapagurus A. Milne-Edwards and Hemipagurus Smith from the Indo- West Pacific (Crustacea: Decapoda: Anomura: Paguridae). Invertebrate Taxonomy, 15: 823 - 891." type="journal article" year="2001">Asakura, 2001</bibRefCitation>
: fig. 37F); the pereopod 2 meri are only armed with a dorsodistal spine in
<taxonomicName id="4C564D1B8C0E1034627DFCBA81D4FCA6" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[1292,1444,829,853]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0E1034627DFCBA81D4FCA6" box="[1292,1444,829,853]" italics="true" pageId="6" pageNumber="162">C. exilidigitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21157F18C0E1034645CFCDA87DEFC86" box="[813,942,861,885]" pageId="6" pageNumber="162" rank="species">new species</taxonomicNameLabel>
(cf.
<figureCitation id="136D2A1D8C0E10346492FCD9804AFC86" box="[995,1082,861,885]" captionStart="Fig" captionStartId="5.[136,171,1984,2006]" captionTargetBox="[129,1475,267,1968]" captionTargetId="figure-8@5.[118,1494,212,2011]" captionTargetPageId="5" captionText="Fig. 4. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, right pereopod 2, lateral view; B, same, dactylus and distal part of propodus, mesial view; C, left pereopod 3, lateral view; D, same, dactylus and distal part of propodus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5352044" httpUri="https://zenodo.org/record/5352044/files/figure.png" pageId="6" pageNumber="162">Fig. 4A</figureCitation>
), rather than armed with subdistal spines on the dorsal surface in addition to the dorsodistal spine in
<taxonomicName id="4C564D1B8C0E103464FBFC1A8031FC46" authorityName="McLaughlin" authorityYear="1997" box="[906,1089,925,949]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="tanimbarensis">
<emphasis id="B922EA8A8C0E103464FBFC1A8031FC46" box="[906,1089,925,949]" italics="true" pageId="6" pageNumber="162">C. tanimbarensis</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFC74B698C0E10346302FC19814EFC46" author="McLaughlin PA" box="[1139,1342,925,949]" pageId="6" pageNumber="162" pagination="433 - 572" refId="ref9882" refString="McLaughlin PA (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier A &amp; Bouchet P (eds.) Resultats des Campagnes MUSORSTOM, Volume 16. Memoires du Museum national d'Histoire naturelle, Paris, 172: 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
: fig. 18d).
</paragraph>
</subSubSection>
<subSubSection id="C34C65138C0E1034645CFC598041FBC6" pageId="6" pageNumber="162" type="etymology">
<paragraph id="8BE936988C0E1034645CFC598041FBC6" blockId="6.[811,1451,989,1077]" pageId="6" pageNumber="162">
<emphasis id="B922EA8A8C0E1034645CFC5987DFFC06" bold="true" box="[813,943,989,1013]" pageId="6" pageNumber="162">Etymology.</emphasis>
The Latin
<taxonomicName id="4C564D1B8C0E1034635CFC5980D4FC06" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[1069,1188,989,1013]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="exilidigitus">
<emphasis id="B922EA8A8C0E1034635CFC5980D4FC06" box="[1069,1188,989,1013]" italics="true" pageId="6" pageNumber="162">exilidigitus</emphasis>
</taxonomicName>
(
<emphasis id="B922EA8A8C0E103463C4FC59809DFC06" box="[1205,1261,989,1013]" italics="true" pageId="6" pageNumber="162">exilis</emphasis>
= thin;
<emphasis id="B922EA8A8C0E1034623BFC5981E3FC06" box="[1354,1427,989,1013]" italics="true" pageId="6" pageNumber="162">digitus</emphasis>
= digit, toe) refers to the very slender dactyli of the ambulatory legs of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>