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<document id="8A3B5D975DFF9E30ED9354026455EEF9" ID-DOI="10.5281/zenodo.196399" ID-GBIF-Dataset="a8861489-e7c9-499e-be53-5925446a5aa5" ID-ISSN="1175-5326" ID-Zenodo-Dep="196399" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460243044829" checkinUser="plazi" docAuthor="Barnich, Ruth &amp; Fiege, Dieter" docDate="2010" docId="A04A87E7471BFF83FF5ACF8340AB31D7" docLanguage="en" docName="zt02525p018.pdf" docOrigin="Zootaxa 2525" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Harmothoe globifera G.O. Sars 1873" docType="treatment" docVersion="9" lastPageNumber="4" masterDocId="5C73FF9F471AFF80FFCDCB46416C341B" masterDocTitle="On the distinction of Harmothoe globifera (G. O. Sars, 1873) and some other easily confused polynoids in the NE Atlantic, with the description of a new species of Acanthicolepis Norman in McIntosh, 1900 (Polychaeta, Polynoidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="2" updateTime="1698241275722" updateUser="plazi">
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<mods:title id="412E5E8EA316280F2942FCC4A06E6F5D">On the distinction of Harmothoe globifera (G. O. Sars, 1873) and some other easily confused polynoids in the NE Atlantic, with the description of a new species of Acanthicolepis Norman in McIntosh, 1900 (Polychaeta, Polynoidae)</mods:title>
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<mods:date id="834C0B61AF2C9808D78F422418527E09">2010</mods:date>
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<taxonomicName id="EFE34D72471BFF81FF5ACF83431D30C4" ID-CoL="6LCFB" authority="G.O. Sars, 1873" authorityName="G.O. Sars" authorityYear="1873" box="[151,625,1221,1247]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE3471BFF81FF5ACF83431D30C4" bold="true" box="[151,625,1221,1247]" pageId="1" pageNumber="2">
<emphasis id="1A97EAE3471BFF81FF5ACF8340FA30C4" bold="true" box="[151,406,1221,1247]" italics="true" pageId="1" pageNumber="2">Harmothoe globifera</emphasis>
(
<bibRefCitation id="4C724B00471BFF81FE68CF83430530C4" author="Sars" box="[421,617,1221,1247]" pageId="1" pageNumber="18" refString="Sars, G. O. (1873) Bidrag til kundskaben om dyrelivet paa vore havbanker. Forhandlinger i Videnskabsselskabet i Kristiana, 73 - 119." type="book chapter" year="1873">G.O. Sars, 1873</bibRefCitation>
)
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(
<figureCitation id="B0D82A74471BFF81FF52CFAE418C3119" box="[159,224,1256,1282]" captionStart="FIGURE 1" captionStartId="4.[151,255,1880,1904]" captionTargetBox="[240,1348,202,1850]" captionTargetId="figure@4.[231,1353,194,1865]" captionTargetPageId="4" captionText="FIGURE 1. Harmothoe globifera (SMF 18887). (A) anterior end; (B) right anterior elytron from unknown segment, free in vial; (C) micro- and macrotubercles from same; (D) left elytragerous parapodium from segment 5, posterior view; (E) distal half of long notochaeta; (F) distal part of middle, unidentate neurochaeta; (G) tip of same; (H) tip of upper bidentate neurochaeta with fragile secondary tooth; (I) tip of upper bidentate neurochaeta with abraded secondary tooth; (J) tip of lower unidentate neurochaeta. (Scales: A, B = 2 mm; C = 250 µm; D = 500 µm; E, F = 100 µm; G – J = 50 µm)." httpUri="https://zenodo.org/record/196400/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
A–J)
</paragraph>
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<subSubSection id="60F9657A471BFF81FF5ACE7442E23292" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="285C36F1471BFF81FF5ACE7443453152" blockId="1.[151,1095,1330,1673]" box="[151,553,1330,1353]" pageId="1" pageNumber="2">
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<taxonomicName id="EFE34D72471BFF81FF5ACE7443493152" ID-CoL="484VB" authority="G.O. Sars, 1873: 95" authorityName="G.O. Sars" authorityPageNumber="95" authorityYear="1873" box="[151,549,1330,1353]" class="Polychaeta" family="Polynoidae" genus="Nychia" higherTaxonomySource="CoL" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE3471BFF81FF5ACE7440273152" box="[151,331,1330,1353]" italics="true" pageId="1" pageNumber="2">Nychia globifera</emphasis>
<treatmentCitation id="A94210E0471BFF81FE9CCE7443493152" author="Sars" box="[337,549,1330,1353]" page="95" pageId="1" pageNumber="2" year="1873">
<bibRefCitation id="4C724B00471BFF81FE9CCE7440913152" author="Sars" box="[337,509,1330,1353]" pageId="1" pageNumber="18" refString="Sars, G. O. (1873) Bidrag til kundskaben om dyrelivet paa vore havbanker. Forhandlinger i Videnskabsselskabet i Kristiana, 73 - 119." type="book chapter" year="1873">G.O. Sars, 1873</bibRefCitation>
: 95
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.
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<paragraph id="285C36F1471BFF81FF5ACE1443193192" blockId="1.[151,1095,1330,1673]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="08F311DF471BFF81FF5ACE1443193192" pageId="1" pageNumber="2">
<taxonomicName id="EFE34D72471BFF81FF5ACE14436A3172" ID-CoL="4LCD7" authority="Hansen (1882)" authorityName="Hansen" authorityYear="1882" box="[151,518,1362,1385]" class="Polychaeta" family="Polynoidae" genus="Polynoe" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE3471BFF81FF5ACE14403B3172" box="[151,343,1362,1385]" italics="true" pageId="1" pageNumber="2">Polynoe globifera</emphasis>
: 
<treatmentCitation id="A94210E0471BFF81FEA9CE14436A3172" author="Hansen" box="[356,518,1362,1385]" page="4" pageId="1" pageNumber="2" year="1882">
<bibRefCitation id="4C724B00471BFF81FEA9CE14436A3172" author="Hansen" box="[356,518,1362,1385]" pageId="1" pageNumber="17" refString="Hansen, G. A. (1882) Annelida. The Norwegian North-Atlantic Expedition 1876 - 1878. Zoology, 3 (2), 1 - 53." type="journal article" year="1882">Hansen (1882)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
: 4, 14, 17, 18, 23, pl. 2 figs. 1–9. 
<taxonomicName id="EFE34D72471BFF81FF5ACE3443253192" authority="Bidenkap (1894)" authorityName="Bidenkap" authorityYear="1894" box="[151,585,1394,1417]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE3471BFF81FF5ACE3440EF3192" box="[151,387,1394,1417]" italics="true" pageId="1" pageNumber="2">Lepidonotus globifera</emphasis>
: 
<treatmentCitation id="A94210E0471BFF81FE5CCE3443253192" author="Bidenkap" box="[401,585,1394,1417]" page="61" pageId="1" pageNumber="2" year="1894">
<bibRefCitation id="4C724B00471BFF81FE5CCE3443253192" author="Bidenkap" box="[401,585,1394,1417]" pageId="1" pageNumber="17" refString="Bidenkap, O. (1894) Systematisk oversigt over Norges Annulata Polychaeta. Christiania Videnskabs-Selskabs Forhandlinger, 10, 1 - 142." type="journal article" year="1894">Bidenkap (1894)</bibRefCitation>
</treatmentCitation>
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: 61.
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</paragraph>
<paragraph id="285C36F1471BFF81FF5ACED4427831B2" blockId="1.[151,1095,1330,1673]" box="[151,788,1426,1449]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="08F311DF471BFF81FF5ACED4427831B2" box="[151,788,1426,1449]" pageId="1" pageNumber="2">
<taxonomicName id="EFE34D72471BFF81FF5ACED4408731B2" ID-CoL="3CG3M" authority="Fauvel (1914)" authorityName="Fauvel" authorityYear="1914" box="[151,491,1426,1449]" class="Polychaeta" family="Polynoidae" genus="Eunoe" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE3471BFF81FF5ACED4402831B2" box="[151,324,1426,1449]" italics="true" pageId="1" pageNumber="2">Eunoe globifera</emphasis>
: 
<treatmentCitation id="A94210E0471BFF81FE9CCED4408731B2" author="Fauvel" box="[337,491,1426,1449]" page="52" pageId="1" pageNumber="2" year="1914">
<bibRefCitation id="4C724B00471BFF81FE9CCED4408731B2" author="Fauvel" box="[337,491,1426,1449]" pageId="1" pageNumber="17" refString="Fauvel, P. (1914) Annelides Polychetes non pelagiques provenant des Campagnes de l' &quot; Hirondelle &quot; et de la &quot; Princesse- Alice &quot; (1885 - 1910). Resultats des campagnes scientifiques accomplies sur son yacht par Albert I er Prince Souverain de Monaco, 46, 1 - 432." type="journal article" year="1914">Fauvel (1914)</bibRefCitation>
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: 52; 
<treatmentCitation id="A94210E0471BFF81FDD3CED4427C31B2" author="Uschakov" box="[542,784,1426,1449]" page="178" pageId="1" pageNumber="2" year="1982">
<bibRefCitation id="4C724B00471BFF81FDD3CED443B631B2" author="Uschakov" box="[542,730,1426,1449]" pageId="1" pageNumber="18" refString="Uschakov, P. V. (1982) Polychaetes of the suborder Aphroditiformia of the Arctic Ocean and the northwestern part of the Pacific. Families Aphroditidae and Polynoidae. Fauna of the USSR. Polychaetes, 2 (1), 1 - 271." type="journal article" year="1982">Uschakov (1982)</bibRefCitation>
: 178
</treatmentCitation>
.
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</paragraph>
<paragraph id="285C36F1471BFF81FF5ACEF443F531F2" blockId="1.[151,1095,1330,1673]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="08F311DF471BFF81FF5ACEF443F531F2" pageId="1" pageNumber="2">
<taxonomicName id="EFE34D72471BFF81FF5ACEF4435231D2" ID-CoL="6LCFB" authority="Ditlevsen (1917)" authorityName="Ditlevsen" authorityYear="1917" box="[151,574,1458,1481]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE3471BFF81FF5ACEF4401531D2" box="[151,377,1458,1481]" italics="true" pageId="1" pageNumber="2">Harmothoe globifera</emphasis>
: 
<treatmentCitation id="A94210E0471BFF81FE4ACEF4435231D2" author="Ditlevsen" box="[391,574,1458,1481]" page="9" pageId="1" pageNumber="2" year="1917">
<bibRefCitation id="4C724B00471BFF81FE4ACEF4435231D2" author="Ditlevsen" box="[391,574,1458,1481]" pageId="1" pageNumber="17" refString="Ditlevsen, H. (1917) Annelids. The Danish Ingolf Expedition, 4 (4), 1 - 71." type="journal article" year="1917">Ditlevsen (1917)</bibRefCitation>
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: 9; 
<treatmentCitation id="A94210E0471BFF81FDA9CEF4428331D2" author="Loshamn" box="[612,1007,1458,1481]" page="147" pageId="1" pageNumber="2" year="1980">
<bibRefCitation id="4C724B00471BFF81FDA9CEF442D731D2" author="Loshamn" box="[612,955,1458,1481]" pageId="1" pageNumber="17" refString="Loshamn, A. A. (1980) En systematisk og dyregeografisk Undersokelse over Skjellrygg-Gruppen (Familie Aphroditidae sensu Fauvel 1923). Dissertation. Zoologisk Institutt, Universitete i Oslo, 184 pp. [unpublished]." type="book" year="1980">Loshamn (1980, unpubl. thesis)</bibRefCitation>
: 147
</treatmentCitation>
, fig. 72. 
<taxonomicName id="EFE34D72471BFF81FF5ACE94437231F2" ID-CoL="6JXK3" authority="Augener (1929)" authorityName="Augener" authorityYear="1929" box="[151,542,1490,1513]" class="Polychaeta" family="Polynoidae" genus="Gattyana" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE3471BFF81FF5ACE94400F31F2" box="[151,355,1490,1513]" italics="true" pageId="1" pageNumber="2">Gattyana globifera</emphasis>
: 
<treatmentCitation id="A94210E0471BFF81FEBDCE94437231F2" author="Augener" box="[368,542,1490,1513]" page="693" pageId="1" pageNumber="2" year="1929">
<bibRefCitation id="4C724B00471BFF81FEBDCE94437231F2" author="Augener" box="[368,542,1490,1513]" pageId="1" pageNumber="17" refString="Augener, H. (1929) Die Polychaten von Spitzbergen. Fauna Arctica, 5, 649 - 834." type="journal article" year="1929">Augener (1929)</bibRefCitation>
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: 693 [part].
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<paragraph id="285C36F1471BFF81FF5ACEB442E23292" blockId="1.[151,1095,1330,1673]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="08F311DF471BFF81FF5ACEB442E23292" pageId="1" pageNumber="2">
<taxonomicName id="EFE34D72471BFF81FF5ACEB443733212" ID-CoL="3JRGV" authority="Augener (1913)" authorityName="Augener" authorityYear="1913" box="[151,543,1522,1545]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="nodosa">
<emphasis id="1A97EAE3471BFF81FF5ACEB440083212" box="[151,356,1522,1545]" italics="true" pageId="1" pageNumber="2">Harmothoe nodosa</emphasis>
: 
<treatmentCitation id="A94210E0471BFF81FEBFCEB443733212" author="Augener" box="[370,543,1522,1545]" page="210" pageId="1" pageNumber="2" year="1913">
<bibRefCitation id="4C724B00471BFF81FEBFCEB443733212" author="Augener" box="[370,543,1522,1545]" pageId="1" pageNumber="17" refString="Augener, H. (1913) Polychaeten von Franz-Joseph-Land I. Zoologischer Anzeiger, 41 (5), 202 - 220." type="journal article" year="1913">Augener (1913)</bibRefCitation>
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: 210 [part, not 
<taxonomicName id="EFE34D72471BFF81FD0DCEB445673212" ID-CoL="3CG47" authority="M. Sars, 1861" authorityName="M. Sars" authorityYear="1861" box="[704,1035,1522,1545]" class="Polychaeta" family="Polynoidae" genus="Eunoe" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="nodosa">
<emphasis id="1A97EAE3471BFF81FD0DCEB442343212" box="[704,856,1522,1545]" italics="true" pageId="1" pageNumber="2">Eunoe nodosa</emphasis>
(
<bibRefCitation id="4C724B00471BFF81FCAACEB4456F3212" author="Sars" box="[871,1027,1522,1545]" pageId="1" pageNumber="18" refString="Sars, M. (1861) Om de ved Norges Kyster forekommende Arter af Annelideslaegten Polynoe. Forhandlinger i Videnskabsselskabet i Kristiana, 54 - 62." type="book chapter" year="1861">M. Sars, 1861</bibRefCitation>
)
</taxonomicName>
]. 
<taxonomicName id="EFE34D72471BFF81FF5ACD54432C3232" ID-CoL="3CG47" authority="Barnich &amp; Fiege (2009)" authorityName="Barnich &amp; Fiege" authorityYear="2009" box="[151,576,1554,1577]" class="Polychaeta" family="Polynoidae" genus="Eunoe" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="nodosa">
<emphasis id="1A97EAE3471BFF81FF5ACD5440433232" box="[151,303,1554,1577]" italics="true" pageId="1" pageNumber="2">Eunoe nodosa</emphasis>
: Barnich &amp; Fiege (2009)
</taxonomicName>
: 71 [not 
<taxonomicName id="EFE34D72471BFF81FD50CD54428B3232" ID-CoL="3CG47" authority="M. Sars, 1861" authorityName="M. Sars" authorityYear="1861" box="[669,999,1554,1577]" class="Polychaeta" family="Polynoidae" genus="Eunoe" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="nodosa">
<emphasis id="1A97EAE3471BFF81FD50CD5442583232" box="[669,820,1554,1577]" italics="true" pageId="1" pageNumber="2">Eunoe nodosa</emphasis>
(
<bibRefCitation id="4C724B00471BFF81FC89CD5442B33232" author="Sars" box="[836,991,1554,1577]" pageId="1" pageNumber="18" refString="Sars, M. (1861) Om de ved Norges Kyster forekommende Arter af Annelideslaegten Polynoe. Forhandlinger i Videnskabsselskabet i Kristiana, 54 - 62." type="book chapter" year="1861">M. Sars, 1861</bibRefCitation>
)
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]. 
<taxonomicName id="EFE34D72471BFF81FF5ACD74434F3252" ID-CoL="9d2fc904-44be-4d8c-912c-52dda5b3ee94" authority="Hansen, 1880: 225" authorityName="Hansen" authorityPageNumber="225" authorityYear="1880" box="[151,547,1586,1609]" class="Polychaeta" family="Polynoidae" genus="Polynoe" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="assimilis">
<emphasis id="1A97EAE3471BFF81FF5ACD74403E3252" box="[151,338,1586,1609]" italics="true" pageId="1" pageNumber="2">Polynoe assimilis</emphasis>
<treatmentCitation id="A94210E0471BFF81FE95CD74434F3252" author="Hansen" box="[344,547,1586,1609]" page="225" pageId="1" pageNumber="2" year="1880">
<bibRefCitation id="4C724B00471BFF81FE95CD7440833252" author="Hansen" box="[344,495,1586,1609]" pageId="1" pageNumber="17" refString="Hansen, G. A. (1880) Annelider fra den norske Nordhavsexpedition i 1878. Nyt Magazin for Naturvidenskaberne, Oslo, 25, 224 - 234." type="journal article" year="1880">Hansen, 1880</bibRefCitation>
: 225
</treatmentCitation>
</taxonomicName>
, pl. 2 figs. 1–5; 
<treatmentCitation id="A94210E0471BFF81FD1DCD7442F53252" author="Hansen" box="[720,921,1586,1609]" page="27" pageId="1" pageNumber="2" year="1882">
<bibRefCitation id="4C724B00471BFF81FD1DCD74421D3252" author="Hansen" box="[720,881,1586,1609]" pageId="1" pageNumber="17" refString="Hansen, G. A. (1882) Annelida. The Norwegian North-Atlantic Expedition 1876 - 1878. Zoology, 3 (2), 1 - 53." type="journal article" year="1882">Hansen (1882)</bibRefCitation>
: 27
</treatmentCitation>
. 
<taxonomicName id="EFE34D72471BFF81FF5ACD1443443272" authority="McIntosh, 1885: 77" authorityName="McIntosh" authorityPageNumber="77" authorityYear="1885" box="[151,552,1618,1641]" class="Magnoliopsida" family="Achariaceae" genus="Dasylepis" kingdom="Animalia" order="Malpighiales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="species" species="equitis">
<emphasis id="1A97EAE3471BFF81FF5ACD1440223272" box="[151,334,1618,1641]" italics="true" pageId="1" pageNumber="2">Dasylepis equitis</emphasis>
<treatmentCitation id="A94210E0471BFF81FE99CD1443443272" author="McIntosh" box="[340,552,1618,1641]" page="77" pageId="1" pageNumber="2" year="1885">
<bibRefCitation id="4C724B00471BFF81FE99CD14436D3272" author="McIntosh" box="[340,513,1618,1641]" pageId="1" pageNumber="17" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. &quot; Challenger &quot; during the years 1873 - 76. Reports on the scientific results of the voyage of H. M. S. &quot; Challenger &quot;, Zoology, 12, 1 - 554." type="journal article" year="1885">McIntosh, 1885</bibRefCitation>
: 77
</treatmentCitation>
</taxonomicName>
, pl. 32 fig. 7, pl. 32A figs. 1, 2 [new synonymy]. 
<taxonomicName id="EFE34D72471BFF81FF5ACD34431E3292" ID-CoL="8TNL" authority="Barnich et al. (2000)" authorityName="Barnich et al." authorityYear="2000" box="[151,626,1650,1673]" class="Polychaeta" family="Polynoidae" genus="Acanthicolepis" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="equitis">
<emphasis id="1A97EAE3471BFF81FF5ACD3440EF3292" box="[151,387,1650,1673]" italics="true" pageId="1" pageNumber="2">Acanthicolepis equitis</emphasis>
: 
<treatmentCitation id="A94210E0471BFF81FE5CCD34431E3292" author="" box="[401,626,1650,1673]" page="316" pageId="1" pageNumber="2">
<bibRefCitation id="4C724B00471BFF81FE5CCD34431E3292" box="[401,626,1650,1673]" pageId="1" pageNumber="17" refString="Barnich, R., Gil, J. &amp; Fiege, D. (2000) Revision of the genus Acanthicolepis Norman in McIntosh, 1900 (Polychaeta, Polynoidae, Polynoinae). Sarsia, 85, 313 - 319." type="journal article">Barnich et al. (2000)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
: 316 [as doubtful species].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="60F9657A471BFF81FF5ACDF1454B3359" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="285C36F1471BFF81FF5ACDF1409632EC" blockId="1.[151,1436,1719,2007]" pageId="1" pageNumber="2">
<emphasis id="1A97EAE3471BFF81FF5ACDF1402232CA" bold="true" box="[151,334,1719,1745]" pageId="1" pageNumber="2">
<typeStatus id="F7588853471BFF81FF5ACDF141B932CA" box="[151,213,1719,1745]" pageId="1" pageNumber="2">Type</typeStatus>
material.
</emphasis>
<taxonomicName id="EFE34D72471BFF81FE9BCDFE437232CA" box="[342,542,1720,1745]" class="Polychaeta" family="Polynoidae" genus="Nychia" higherTaxonomySource="CoL" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE3471BFF81FE9BCDFE437232CA" box="[342,542,1720,1745]" italics="true" pageId="1" pageNumber="2">Nychia globifera</emphasis>
</taxonomicName>
: 
<typeStatus id="F7588853471BFF81FDE3CDFE43F932C9" box="[558,661,1720,1746]" pageId="1" pageNumber="2" type="syntype">syntypes</typeStatus>
lost; NE Atlantic, off &quot;Storeggen&quot; in great depths and at Lofoten Islands in 300 to 400 fathoms.
</paragraph>
<paragraph id="285C36F1471BFF81FF08CC45454B3359" blockId="1.[151,1436,1719,2007]" pageId="1" pageNumber="2">
<taxonomicName id="EFE34D72471BFF81FF08CC4540FF3307" box="[197,403,1795,1820]" class="Polychaeta" family="Polynoidae" genus="Polynoe" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="assimilis">
<emphasis id="1A97EAE3471BFF81FF08CC4540FF3307" box="[197,403,1795,1820]" italics="true" pageId="1" pageNumber="2">Polynoe assimilis</emphasis>
</taxonomicName>
: 4 
<typeStatus id="F7588853471BFF81FE7BCC4443713307" box="[438,541,1794,1820]" pageId="1" pageNumber="2" type="syntype">syntypes</typeStatus>
(2 cs, 2 af, 1 mf, 1pf, several elytra free in vial), 
<collectionCode id="4EF2AE34471BFF81FB9BCC4445C73307" box="[1110,1195,1794,1820]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="1" pageNumber="2">ZMBN</collectionCode>
1989, Arctic Ocean, Norske Nordhavsexpedition 1878, St. 363, 
<geoCoordinate id="4DD75036471BFF81FD5FCC6E43DE3359" box="[658,690,1832,1858]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="80.05">80</geoCoordinate>
°3'N 
<geoCoordinate id="4DD75036471BFF81FD26CC6E42513359" box="[747,829,1832,1858]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="8.466666">8°28'E</geoCoordinate>
, bluish clay, 
<specimenCount id="3EE5FD78471BFF81FC18CC6E454B3359" box="[981,1063,1832,1858]" pageId="1" pageNumber="2" type="generic">
<quantity id="EF1B9B14471BFF81FC18CC6E45483359" box="[981,1060,1832,1858]" metricMagnitude="2" metricUnit="m" metricValue="4.75" pageId="1" pageNumber="2" unit="m" value="475.0">475 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="60F9657A471BFF81FF08CC0B401A3397" pageId="1" pageNumber="2" type="description">
<paragraph id="285C36F1471BFF81FF08CC0B401A3397" blockId="1.[151,1436,1719,2007]" pageId="1" pageNumber="2">
<taxonomicName id="EFE34D72471BFF81FF08CC0B40E2337D" box="[197,398,1869,1894]" class="Magnoliopsida" family="Achariaceae" genus="Dasylepis" kingdom="Animalia" order="Malpighiales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="species" species="equitis">
<emphasis id="1A97EAE3471BFF81FF08CC0B40E2337D" box="[197,398,1869,1894]" italics="true" pageId="1" pageNumber="2">Dasylepis equitis</emphasis>
</taxonomicName>
: 
<typeStatus id="F7588853471BFF81FE53CC0B4369337C" box="[414,517,1869,1895]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
(pf) lost; NE Atlantic, &quot;Knight Errant&quot; St. 8, 
<collectingCountry id="50F47661471BFF81FBD0CC0B450C337C" box="[1053,1120,1869,1895]" name="Faroe Islands" pageId="1" pageNumber="2">Faroe</collectingCountry>
Channel, 
<date id="5C5D1031471BFF81FB15CC0B44F4337C" box="[1240,1432,1869,1895]" pageId="1" pageNumber="2" value="1880-08-17">17 August 1880</date>
, 540 fathoms, ooze.
</paragraph>
</subSubSection>
<subSubSection id="60F9657A471BFF82FF0BCCD140D936F9" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="285C36F1471BFF82FF0BCCD140D936F9" blockId="1.[151,1436,1719,2007]" lastBlockId="2.[151,1437,152,2007]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<emphasis id="1A97EAE3471BFF81FF0BCCD140AA33AA" bold="true" box="[198,454,1943,1969]" pageId="1" pageNumber="2">Additional material.</emphasis>
1 spm. (cs), Akvaplan-Niva collection, NE Atlantic, Hydro St. 3-3, 
<date id="5C5D1031471BFF81FB38CCDE447E33A9" box="[1269,1298,1944,1970]" pageId="1" pageNumber="2" value="2003-06-13">13</date>
June 2003, 
<geoCoordinate id="4DD75036471BFF81FF5ACCFB41DA33CC" box="[151,182,1981,2007]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="1" value="72.56129">72</geoCoordinate>
°33.677’N 
<geoCoordinate id="4DD75036471BFF81FEF4CCFB40D933CC" box="[313,437,1981,2007]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="9" value="21.019833">21°1.19’E</geoCoordinate>
, 
<specimenCount id="3EE5FD78471BFF81FE0FCCFB437933CC" box="[450,533,1981,2007]" pageId="1" pageNumber="2" type="generic">
<quantity id="EF1B9B14471BFF81FE0FCCFB437E33CC" box="[450,530,1981,2007]" metricMagnitude="2" metricUnit="m" metricValue="3.8299999999999996" pageId="1" pageNumber="2" unit="m" value="383.0">383 m</quantity>
.
</specimenCount>
1 spm. (juv.), 
<collectionCode id="4EF2AE34471BFF81FD09CCFB439333CC" LSID="urn:lsid:biocol.org:col:34838" box="[708,767,1981,2007]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="2">SMF</collectionCode>
18881, NE Atlantic, Nucula St. 
<date id="5C5D1031471BFF81FB49CCFB45C233CC" box="[1156,1198,1981,2007]" pageId="1" pageNumber="2" value="2006-09">5-3</date>
, 
<date id="5C5D1031471BFF81FB71CCFB44FB33CC" box="[1212,1431,1981,2007]" pageId="1" pageNumber="2" value="2006-09-06">6 September 2006</date>
, 
<geoCoordinate id="4DD750364718FF82FF5ACBDE406034A9" box="[151,268,152,178]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="71.545">71.545°N</geoCoordinate>
<geoCoordinate id="4DD750364718FF82FED9CBDE401634A9" box="[276,378,152,178]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="555" value="25.24">25.24°E</geoCoordinate>
, 
<quantity id="EF1B9B144718FF82FE4ACBDE40BB34A9" box="[391,471,152,178]" metricMagnitude="2" metricUnit="m" metricValue="2.92" pageId="2" pageNumber="3" unit="m" value="292.0">292 m</quantity>
, ded. A. Sikorski. 1 spm., 
<collectionCode id="4EF2AE344718FF82FCD3CBDE423634A9" LSID="urn:lsid:biocol.org:col:34838" box="[798,858,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="3">SMF</collectionCode>
18882, NE Atlantic, Snøhvit St. N3-2, 
<date id="5C5D10314718FF82FAF3CBDE443034A9" box="[1342,1372,152,178]" pageId="2" pageNumber="3" value="2007-06-19">19</date>
June 2007, 
<geoCoordinate id="4DD750364718FF82FF2DCBFB403D34CC" box="[224,337,189,215]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="71.49">71.490°N</geoCoordinate>
<geoCoordinate id="4DD750364718FF82FE95CBFB40A634CC" box="[344,458,189,215]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="55" value="21.087">21.087°E</geoCoordinate>
, 
<quantity id="EF1B9B144718FF82FE18CBFB434E34CC" box="[469,546,189,215]" metricMagnitude="2" metricUnit="m" metricValue="3.22" pageId="2" pageNumber="3" unit="m" value="322.0">322 m</quantity>
, ded. A. Sikorski. 1 spm. (juv.), 
<collectionCode id="4EF2AE344718FF82FC56CBFB42BA34CC" LSID="urn:lsid:biocol.org:col:34838" box="[923,982,189,215]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="3">SMF</collectionCode>
18883, NE Atlantic, Snøhvit St. N3-3, 
<date id="5C5D10314718FF82FF5ACBA441D834E7" box="[151,180,226,252]" pageId="2" pageNumber="3" value="2007-06-19">19</date>
June 2007, 
<geoCoordinate id="4DD750364718FF82FE8FCBA440DF34E7" box="[322,435,226,252]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="71.49">71.490°N</geoCoordinate>
<geoCoordinate id="4DD750364718FF82FE77CBA4434034E7" box="[442,556,226,252]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="55" value="21.087">21.087°E</geoCoordinate>
, 
<quantity id="EF1B9B144718FF82FDFACBA443E934E7" box="[567,645,226,252]" metricMagnitude="2" metricUnit="m" metricValue="3.22" pageId="2" pageNumber="3" unit="m" value="322.0">322 m</quantity>
, ded. A. Sikorski. 2 spms. (juv.), 
<collectionCode id="4EF2AE344718FF82FBC6CBA4452A34E7" LSID="urn:lsid:biocol.org:col:34838" box="[1035,1094,226,252]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="3">SMF</collectionCode>
18884, NE Atlantic, Snøhvit St. N4-1, 
<date id="5C5D10314718FF82FEC6CA4E40443539" box="[267,296,264,290]" pageId="2" pageNumber="3" value="2007-06-19">19</date>
June 2007, 
<geoCoordinate id="4DD750364718FF82FE74CA4E43473539" box="[441,555,264,290]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="71.491">71.491°N</geoCoordinate>
<geoCoordinate id="4DD750364718FF82FDFECA4E43C93539" box="[563,677,264,290]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="55" value="21.091">21.091°E</geoCoordinate>
, 
<quantity id="EF1B9B144718FF82FD7FCA4E426C3539" box="[690,768,264,290]" metricMagnitude="2" metricUnit="m" metricValue="3.22" pageId="2" pageNumber="3" unit="m" value="322.0">322 m</quantity>
, ded. A. Sikorski. 1 spm. (1 af, 1 mf, juv.), 
<collectionCode id="4EF2AE344718FF82FACACA4E442E3539" LSID="urn:lsid:biocol.org:col:34838" box="[1287,1346,264,290]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="3">SMF</collectionCode>
18885, NE Atlantic, Snøhvit St. N4-3, 
<date id="5C5D10314718FF82FDCACA6B4348355C" box="[519,548,301,327]" pageId="2" pageNumber="3" value="2007-06-19">19</date>
June 2007, 
<geoCoordinate id="4DD750364718FF82FD7ECA6B4249355C" box="[691,805,301,327]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="71.491">71.491°N</geoCoordinate>
<geoCoordinate id="4DD750364718FF82FCE1CA6B42F1355C" box="[812,925,301,327]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="55" value="21.091">21.091°E</geoCoordinate>
, 
<quantity id="EF1B9B144718FF82FC64CA6B429B355C" box="[937,1015,301,327]" metricMagnitude="2" metricUnit="m" metricValue="3.22" pageId="2" pageNumber="3" unit="m" value="322.0">322 m</quantity>
, ded. A. Sikorski. 1 spm. (cs), 
<collectionCode id="4EF2AE344718FF82FAACCA6B44F0355C" LSID="urn:lsid:biocol.org:col:34838" box="[1377,1436,301,327]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="3">SMF</collectionCode>
18886, NE Atlantic, Snøhvit St. SD4-5, 
<date id="5C5D10314718FF82FDBFCA1443E33577" box="[626,655,338,364]" pageId="2" pageNumber="3" value="2003-06-12">12</date>
June 2003, 
<geoCoordinate id="4DD750364718FF82FCD2CA1442503577" box="[799,828,338,364]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="71.5943">71</geoCoordinate>
°35.658’N 
<geoCoordinate id="4DD750364718FF82FC0DCA1445343577" box="[960,1112,338,364]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="21.286833">21°17.210’E</geoCoordinate>
, 
<quantity id="EF1B9B144718FF82FBA9CA1445DE3577" box="[1124,1202,338,364]" metricMagnitude="2" metricUnit="m" metricValue="3.4299999999999997" pageId="2" pageNumber="3" unit="m" value="343.0">343 m</quantity>
, ded. A. Sikorski. 1 spm. (cs), 
<collectionCode id="4EF2AE344718FF82FEDACA3E403F3589" LSID="urn:lsid:biocol.org:col:34838" box="[279,339,376,402]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="3">SMF</collectionCode>
18887, NE Atlantic, Tornerose St. T 
<date id="5C5D10314718FF82FCD4CA3E422F3589" box="[793,835,376,402]" pageId="2" pageNumber="3" value="2006-04">6-4</date>
, 
<date id="5C5D10314718FF82FC9DCA3E428B3589" box="[848,999,376,402]" pageId="2" pageNumber="3" value="2006-06-03">3 June 2006</date>
, 
<geoCoordinate id="4DD750364718FF82FC39CA3E45053589" box="[1012,1129,376,402]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="71.586">71.586°N</geoCoordinate>
<geoCoordinate id="4DD750364718FF82FBBFCA3E45B43589" box="[1138,1240,376,402]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="555" value="22.85">22.85°E</geoCoordinate>
, 
<quantity id="EF1B9B144718FF82FB28CA3E44593589" box="[1253,1333,376,402]" metricMagnitude="2" metricUnit="m" metricValue="4.12" pageId="2" pageNumber="3" unit="m" value="412.0">412 m</quantity>
, ded. A. Sikorski. 2 spms (2 af, 2 mf), 
<collectionCode id="4EF2AE344718FF82FDC1CADB432535AC" LSID="urn:lsid:biocol.org:col:34838" box="[524,585,413,439]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="3">SMF</collectionCode>
18888, NE Atlantic, Tornerose II St. 
<date id="5C5D10314718FF82FBD1CADB452B35AC" box="[1052,1095,413,439]" pageId="2" pageNumber="3" value="2006-04">2-4</date>
, 
<date id="5C5D10314718FF82FB9BCADB450935AC" box="[1110,1125,413,439]" pageId="2" pageNumber="3" value="2006-10-03">3</date>
October 2006, 
<geoCoordinate id="4DD750364718FF82FAEBCADB44F035AC" box="[1318,1436,413,439]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="71.588">71.588°N</geoCoordinate>
<geoCoordinate id="4DD750364718FF82FF5ACA84419535C7" box="[151,249,450,476]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="555" value="22.86">22.86°E</geoCoordinate>
, 
<quantity id="EF1B9B144718FF82FECBCA84403835C7" box="[262,340,450,476]" metricMagnitude="2" metricUnit="m" metricValue="4.12" pageId="2" pageNumber="3" unit="m" value="412.0">412 m</quantity>
, ded. A. Sikorski. 1 spm. (af, juv.), 
<collectionCode id="4EF2AE344718FF82FD34CA84425835C7" LSID="urn:lsid:biocol.org:col:34838" box="[761,820,450,476]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="3">SMF</collectionCode>
18889, NE Atlantic, Tornerose II St. 
<date id="5C5D10314718FF82FB3CCA84447635C7" box="[1265,1306,450,476]" pageId="2" pageNumber="3" value="2006-10">3-2</date>
, 
<date id="5C5D10314718FF82FAEACA8441B83619" pageId="2" pageNumber="3" value="2006-10-03">3 October 2006</date>
, 
<geoCoordinate id="4DD750364718FF82FF2DCAAE403E3619" box="[224,338,488,514]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="71.589">71.589°N</geoCoordinate>
<geoCoordinate id="4DD750364718FF82FE94CAAE40D73619" box="[345,443,488,514]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="555" value="22.87">22.87°E</geoCoordinate>
, 
<quantity id="EF1B9B144718FF82FE0ACAAE43783619" box="[455,532,488,514]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="3" unit="m" value="410.0">410 m</quantity>
, ded. A. Sikorski. 1 spm. (juv.), 
<collectionCode id="4EF2AE344718FF82FC5DCAAE42A73619" LSID="urn:lsid:biocol.org:col:34838" box="[912,971,488,514]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="3">SMF</collectionCode>
18890, NE Atlantic, Tornerose II St. 
<date id="5C5D10314718FF82FA4ECAAE41C4363C" pageId="2" pageNumber="3" value="2006-04">5- 4</date>
, 
<date id="5C5D10314718FF82FF7BC94B4019363C" box="[182,373,525,551]" pageId="2" pageNumber="3" value="2006-10-04">4 October 2006</date>
, 
<geoCoordinate id="4DD750364718FF82FE4FC94B4099363C" box="[386,501,525,551]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="71.585">71.585°N</geoCoordinate>
<geoCoordinate id="4DD750364718FF82FE30C94B430E363C" box="[509,610,525,551]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="555" value="22.86">22.86°E</geoCoordinate>
, 
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, ded. A. Sikorski. 1 spm. (af, mf), 
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18891, NE Atlantic, Tornerose II St. 
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, 
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, 
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, 
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, ded. A. Sikorski. 2 spms. (juv.), 
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18892, NE Atlantic, Tornerose II St. 
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, 
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, 
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, 
<quantity id="EF1B9B144718FF82FC02C91E45703669" box="[975,1052,600,626]" metricMagnitude="2" metricUnit="m" metricValue="4.12" pageId="2" pageNumber="3" unit="m" value="412.0">412 m</quantity>
, ded. A. Sikorski. 1 spm. (juv., 1 af, 1 mf), 
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18893, NE Atlantic, Tornerose II St. 
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, 
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, 
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, 
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, ded. A. Sikorski. 1 spm. (cs, juv.), 
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18894, NE Atlantic, Tornerose II St. 
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, 
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, 
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, 
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, ded. A. Sikorski.
</paragraph>
</subSubSection>
<subSubSection id="60F9657A4718FF82FF0BC9AB44FE376C" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="285C36F14718FF82FF0BC9AB44FE376C" blockId="2.[151,1437,152,2007]" pageId="2" pageNumber="3">
<emphasis id="1A97EAE34718FF82FF0BC9AB402A371C" bold="true" box="[198,326,749,775]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Anterior pair of eyes dorsolateral at widest part of prostomium. Antennae and cirri densely papillate. Elytra densely papillate at outer lateral margin, posterior margin with scattered, short papillae; micro- and macrotubercles globose to club-shaped, covered by numerous nodular papillae, microtubercles getting larger towards outer lateral and posterior margin, macrotubercles few, in a row near posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="60F9657A4718FF82FF0BC8C4418A32C9" pageId="2" pageNumber="3" type="description">
<paragraph id="285C36F14718FF82FF0BC8C443A23029" blockId="2.[151,1437,152,2007]" pageId="2" pageNumber="3">
<emphasis id="1A97EAE34718FF82FF0BC8C4403B3787" bold="true" box="[198,343,898,924]" pageId="2" pageNumber="3">Description</emphasis>
(based on specimen in good condition, SMF 18887; 
<typeStatus id="F75888534718FF82FC36C8C445413787" box="[1019,1069,898,924]" pageId="2" pageNumber="3">type</typeStatus>
material lost). Body with 36 segments. At anterior end (
<figureCitation id="B0D82A744718FF82FE1EC8EE437937D9" box="[467,533,936,962]" captionStart="FIGURE 1" captionStartId="4.[151,255,1880,1904]" captionTargetBox="[240,1348,202,1850]" captionTargetId="figure@4.[231,1353,194,1865]" captionTargetPageId="4" captionText="FIGURE 1. Harmothoe globifera (SMF 18887). (A) anterior end; (B) right anterior elytron from unknown segment, free in vial; (C) micro- and macrotubercles from same; (D) left elytragerous parapodium from segment 5, posterior view; (E) distal half of long notochaeta; (F) distal part of middle, unidentate neurochaeta; (G) tip of same; (H) tip of upper bidentate neurochaeta with fragile secondary tooth; (I) tip of upper bidentate neurochaeta with abraded secondary tooth; (J) tip of lower unidentate neurochaeta. (Scales: A, B = 2 mm; C = 250 µm; D = 500 µm; E, F = 100 µm; G – J = 50 µm)." httpUri="https://zenodo.org/record/196400/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
A), prostomium bilobed, with distinct cephalic peaks; ceratophore of median antenna in anterior notch, lateral antennae inserted ventrally, styles of antennae densely papillate, tapering; anterior pair of eyes situated dorsolaterally at widest part of prostomium, posterior pair dorsally near hind margin of prostomium; palps papillate, tapering.
</paragraph>
<paragraph id="285C36F14718FF82FF08CF7B441F30B9" blockId="2.[151,1437,152,2007]" pageId="2" pageNumber="3">Tentaculophores inserted laterally to prostomium, each with two to three notochaetae and a dorsal and ventral tentacular cirrus, styles of cirri densely papillate, tapering. Second segment with first pair of elytra, biramous parapodia, and long buccal cirri. Following segments with short, tapering, papillate ventral cirri.</paragraph>
<paragraph id="285C36F14718FF82FF08CFEB427331BC" blockId="2.[151,1437,152,2007]" pageId="2" pageNumber="3">
Fifteen pairs of elytra, covering dorsum, on segments 2, 4, 5, 7, then on every second segment to 23, 26, 29, 32; last four segments cirrigerous; elytra densely papillate at outer lateral margin, posterior margin with scattered, short papillae; micro- and macrotubercles globose to club-shaped, covered by numerous nodular papillae, microtubercles getting larger towards outer lateral and posterior margin, macrotubercles few, in a row near posterior margin; in anterior elytra occasionally one or two branched microtubercles present near outer lateral margin. (
<figureCitation id="B0D82A744718FF82FE64CE2E409D3199" box="[425,497,1384,1410]" captionStart="FIGURE 1" captionStartId="4.[151,255,1880,1904]" captionTargetBox="[240,1348,202,1850]" captionTargetId="figure@4.[231,1353,194,1865]" captionTargetPageId="4" captionText="FIGURE 1. Harmothoe globifera (SMF 18887). (A) anterior end; (B) right anterior elytron from unknown segment, free in vial; (C) micro- and macrotubercles from same; (D) left elytragerous parapodium from segment 5, posterior view; (E) distal half of long notochaeta; (F) distal part of middle, unidentate neurochaeta; (G) tip of same; (H) tip of upper bidentate neurochaeta with fragile secondary tooth; (I) tip of upper bidentate neurochaeta with abraded secondary tooth; (J) tip of lower unidentate neurochaeta. (Scales: A, B = 2 mm; C = 250 µm; D = 500 µm; E, F = 100 µm; G – J = 50 µm)." httpUri="https://zenodo.org/record/196400/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
B,C). Cirrigerous segments with distinct dorsal tubercles; dorsal cirri with cylindrical cirrophore, style densely papillate, tapering.
</paragraph>
<paragraph id="285C36F14718FF82FF08CEF440E2329C" blockId="2.[151,1437,152,2007]" pageId="2" pageNumber="3">
Parapodia biramous; notopodia with elongate acicular lobe; neuropodia with elongate prechaetal acicular lobe with digitiform supra-acicular process; neuropodial postchaetal lobe shorter than prechaetal lobe, rounded; tips of noto- and neuroacicula penetrating epidermis (
<figureCitation id="B0D82A744718FF82FC18CEBB454D320C" box="[981,1057,1533,1559]" captionStart="FIGURE 1" captionStartId="4.[151,255,1880,1904]" captionTargetBox="[240,1348,202,1850]" captionTargetId="figure@4.[231,1353,194,1865]" captionTargetPageId="4" captionText="FIGURE 1. Harmothoe globifera (SMF 18887). (A) anterior end; (B) right anterior elytron from unknown segment, free in vial; (C) micro- and macrotubercles from same; (D) left elytragerous parapodium from segment 5, posterior view; (E) distal half of long notochaeta; (F) distal part of middle, unidentate neurochaeta; (G) tip of same; (H) tip of upper bidentate neurochaeta with fragile secondary tooth; (I) tip of upper bidentate neurochaeta with abraded secondary tooth; (J) tip of lower unidentate neurochaeta. (Scales: A, B = 2 mm; C = 250 µm; D = 500 µm; E, F = 100 µm; G – J = 50 µm)." httpUri="https://zenodo.org/record/196400/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
D). Notochaetae stouter than neurochaetae, with distinct rows of spines and blunt tip (
<figureCitation id="B0D82A744718FF82FCF5CD6442173227" box="[824,891,1570,1596]" captionStart="FIGURE 1" captionStartId="4.[151,255,1880,1904]" captionTargetBox="[240,1348,202,1850]" captionTargetId="figure@4.[231,1353,194,1865]" captionTargetPageId="4" captionText="FIGURE 1. Harmothoe globifera (SMF 18887). (A) anterior end; (B) right anterior elytron from unknown segment, free in vial; (C) micro- and macrotubercles from same; (D) left elytragerous parapodium from segment 5, posterior view; (E) distal half of long notochaeta; (F) distal part of middle, unidentate neurochaeta; (G) tip of same; (H) tip of upper bidentate neurochaeta with fragile secondary tooth; (I) tip of upper bidentate neurochaeta with abraded secondary tooth; (J) tip of lower unidentate neurochaeta. (Scales: A, B = 2 mm; C = 250 µm; D = 500 µm; E, F = 100 µm; G – J = 50 µm)." httpUri="https://zenodo.org/record/196400/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
E); neurochaetae with distinct rows of spines, tips mainly unidentate, some upper neurochaetae minutely bidentate with fragile secondary tooth (often abraded) (
<figureCitation id="B0D82A744718FF82FEC0CD2B4022329C" box="[269,334,1645,1671]" captionStart="FIGURE 1" captionStartId="4.[151,255,1880,1904]" captionTargetBox="[240,1348,202,1850]" captionTargetId="figure@4.[231,1353,194,1865]" captionTargetPageId="4" captionText="FIGURE 1. Harmothoe globifera (SMF 18887). (A) anterior end; (B) right anterior elytron from unknown segment, free in vial; (C) micro- and macrotubercles from same; (D) left elytragerous parapodium from segment 5, posterior view; (E) distal half of long notochaeta; (F) distal part of middle, unidentate neurochaeta; (G) tip of same; (H) tip of upper bidentate neurochaeta with fragile secondary tooth; (I) tip of upper bidentate neurochaeta with abraded secondary tooth; (J) tip of lower unidentate neurochaeta. (Scales: A, B = 2 mm; C = 250 µm; D = 500 µm; E, F = 100 µm; G – J = 50 µm)." httpUri="https://zenodo.org/record/196400/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
F–J).
</paragraph>
<paragraph id="285C36F14718FF82FF0BCDD4418A32C9" blockId="2.[151,1437,152,2007]" pageId="2" pageNumber="3">
<emphasis id="1A97EAE34718FF82FF0BCDD440E932B7" bold="true" box="[198,389,1682,1708]" pageId="2" pageNumber="3">Measurements.</emphasis>
Specimen SMF 18887: cs in two fragments, L 
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, W 
<quantity id="EF1B9B144718FF82FB81CDD445FE32B7" box="[1100,1170,1682,1708]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="3" unit="mm" value="5.0">5 mm</quantity>
for 36 segments (
<figureCitation id="B0D82A744718FF82FAA0CDD441CD32C9" captionStart="FIGURE 1" captionStartId="4.[151,255,1880,1904]" captionTargetBox="[240,1348,202,1850]" captionTargetId="figure@4.[231,1353,194,1865]" captionTargetPageId="4" captionText="FIGURE 1. Harmothoe globifera (SMF 18887). (A) anterior end; (B) right anterior elytron from unknown segment, free in vial; (C) micro- and macrotubercles from same; (D) left elytragerous parapodium from segment 5, posterior view; (E) distal half of long notochaeta; (F) distal part of middle, unidentate neurochaeta; (G) tip of same; (H) tip of upper bidentate neurochaeta with fragile secondary tooth; (I) tip of upper bidentate neurochaeta with abraded secondary tooth; (J) tip of lower unidentate neurochaeta. (Scales: A, B = 2 mm; C = 250 µm; D = 500 µm; E, F = 100 µm; G – J = 50 µm)." httpUri="https://zenodo.org/record/196400/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
A–J).
</paragraph>
</subSubSection>
<subSubSection id="60F9657A4718FF83FF0BCD9B458137FC" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="discussion">
<paragraph id="285C36F14718FF82FF0BCD9B4259337C" blockId="2.[151,1437,152,2007]" pageId="2" pageNumber="3">
<emphasis id="1A97EAE34718FF82FF0BCD9B402C32EC" bold="true" box="[198,320,1757,1783]" pageId="2" pageNumber="3">Remarks.</emphasis>
Although the 
<typeStatus id="F75888534718FF82FE39CD9B433732EC" box="[500,603,1757,1783]" pageId="2" pageNumber="3" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="EFE34D724718FF82FD46CD9B423932ED" box="[651,853,1757,1782]" class="Polychaeta" family="Polynoidae" genus="Nychia" higherTaxonomySource="CoL" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="3" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE34718FF82FD46CD9B423932ED" box="[651,853,1757,1782]" italics="true" pageId="2" pageNumber="3">Nychia globifera</emphasis>
</taxonomicName>
are lost, G.O. Sars' (1873) description is rather detailed, especially regarding the densely papillate dorsal cirri and elytral margin, and the elytral tubercles covered by nodular papillae. These characters, which have been first figured by 
<bibRefCitation id="4C724B004718FF82FBBECC6E44473359" author="Hansen" box="[1139,1323,1832,1858]" pageId="2" pageNumber="17" refString="Hansen, G. A. (1882) Annelida. The Norwegian North-Atlantic Expedition 1876 - 1878. Zoology, 3 (2), 1 - 53." type="journal article" year="1882">Hansen (1882)</bibRefCitation>
, are very typical and allow the certain identification of the species.
</paragraph>
<paragraph id="285C36F14718FF83FF08CC34456535C7" blockId="2.[151,1437,152,2007]" lastBlockId="3.[151,1437,152,1484]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<bibRefCitation id="4C724B004718FF82FF08CC3440E33397" author="Sars" box="[197,399,1906,1932]" pageId="2" pageNumber="18" refString="Sars, G. O. (1873) Bidrag til kundskaben om dyrelivet paa vore havbanker. Forhandlinger i Videnskabsselskabet i Kristiana, 73 - 119." type="book chapter" year="1873">G.O. Sars (1873)</bibRefCitation>
originally assigned the species to 
<taxonomicName id="EFE34D724718FF82FCE3CC35453F3397" authority="Malmgren, 1866" authorityName="Malmgren" authorityYear="1866" box="[814,1107,1906,1932]" class="Polychaeta" family="Polynoidae" genus="Nychia" higherTaxonomySource="CoL" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="3" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34718FF82FCE3CC3542EF3397" box="[814,899,1907,1932]" italics="true" pageId="2" pageNumber="3">Nychia</emphasis>
<bibRefCitation id="4C724B004718FF82FC46CC34453F3397" author="Malmgren" box="[907,1107,1906,1932]" pageId="2" pageNumber="17" refString="Malmgren, A. J. (1866) Nordiska Hafs-Annulata. Ofversigt af Konglia Vetenskaps-Akademiens Forhhandlingar Stockholm, 1, 51 - 110 [read 1865, printed 1866]." type="journal article" year="1866">Malmgren, 1866</bibRefCitation>
</taxonomicName>
(name pre-occupied, today 
<taxonomicName id="EFE34D724718FF82FF5ACCDE40A233A9" authority="McIntosh, 1897" authorityName="McIntosh" authorityYear="1897" box="[151,462,1944,1970]" class="Polychaeta" family="Polynoidae" genus="Gattyana" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="3" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34718FF82FF5ACCDE406933AA" box="[151,261,1944,1969]" italics="true" pageId="2" pageNumber="3">Gattyana</emphasis>
<bibRefCitation id="4C724B004718FF82FEC0CCDE40A233A9" author="McIntosh" box="[269,462,1944,1970]" pageId="2" pageNumber="17" refString="McIntosh, W. C. (1897) Notes from the Gatty Marine Laboratory, St. Andrews, No. XVIII. Annals and Magazine of Natural History, series 6, 20, 167 - 178." type="journal article" year="1897">McIntosh, 1897</bibRefCitation>
</taxonomicName>
). It was later considered as 
<taxonomicName id="EFE34D724718FF82FCD6CCDE428933A9" authority="Savigny" authorityName="Savigny" box="[795,997,1944,1970]" class="Polychaeta" family="Polynoidae" genus="Polynoe" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="3" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34718FF82FCD6CCDE421033AA" box="[795,892,1944,1969]" italics="true" pageId="2" pageNumber="3">Polynoe</emphasis>
Savigny
</taxonomicName>
in 
<bibRefCitation id="4C724B004718FF82FBC1CCDE45AC33A9" author="Lamarck" box="[1036,1216,1944,1970]" pageId="2" pageNumber="17" refString="Lamarck, C. de (1818) Histoire naturelle des animaux sans vertebres. 5. Deterville et Verdiere, Paris, 612 pp." type="book" year="1818">Lamarck, 1818</bibRefCitation>
by 
<bibRefCitation id="4C724B004718FF82FB23CCDE44FB33A9" author="Hansen" box="[1262,1431,1944,1970]" pageId="2" pageNumber="17" refString="Hansen, G. A. (1880) Annelider fra den norske Nordhavsexpedition i 1878. Nyt Magazin for Naturvidenskaberne, Oslo, 25, 224 - 234." type="journal article" year="1880">Hansen (1880</bibRefCitation>
, 
<bibRefCitation id="4C724B004718FF82FF5ACCFB41BB33CC" author="Hansen" box="[151,215,1981,2007]" pageId="2" pageNumber="17" refString="Hansen, G. A. (1882) Annelida. The Norwegian North-Atlantic Expedition 1876 - 1878. Zoology, 3 (2), 1 - 53." type="journal article" year="1882">1882</bibRefCitation>
), and as 
<taxonomicName id="EFE34D724718FF82FE8FCCFB431F33CC" authority="Leach, 1816" authorityName="Leach" authorityYear="1816" box="[322,627,1981,2007]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="3" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34718FF82FE8FCCFB40BA33CD" box="[322,470,1981,2006]" italics="true" pageId="2" pageNumber="3">Lepidonotus</emphasis>
<bibRefCitation id="4C724B004718FF82FE13CCFB431F33CC" author="Leach" box="[478,627,1981,2007]" pageId="2" pageNumber="17" refString="Leach, W. E. (1816) Annulosa. In: Supplement to the 4 th, 5 th, and 6 th editions of the Encyclopaedia Britannica, 1 (2), 401 - 453." type="journal article" year="1816">Leach, 1816</bibRefCitation>
</taxonomicName>
by 
<bibRefCitation id="4C724B004718FF82FD6FCCFB421D33CC" author="Bidenkap" box="[674,881,1981,2007]" pageId="2" pageNumber="17" refString="Bidenkap, O. (1894) Systematisk oversigt over Norges Annulata Polychaeta. Christiania Videnskabs-Selskabs Forhandlinger, 10, 1 - 142." type="journal article" year="1894">Bidenkap (1894)</bibRefCitation>
. 
<bibRefCitation id="4C724B004718FF82FCB0CCFB455133CC" author="Augener" box="[893,1085,1981,2007]" pageId="2" pageNumber="17" refString="Augener, H. (1913) Polychaeten von Franz-Joseph-Land I. Zoologischer Anzeiger, 41 (5), 202 - 220." type="journal article" year="1913">Augener (1913)</bibRefCitation>
put the species in synonymy with 
<taxonomicName id="EFE34D724719FF83FF1BCBDE43E334A9" authority="M. Sars, 1861" authorityName="M. Sars" authorityYear="1861" box="[214,655,152,178]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="nodosa">
<emphasis id="1A97EAE34719FF83FF1BCBDE40AF34AA" box="[214,451,152,177]" italics="true" pageId="3" pageNumber="4">Harmothoe nodosa</emphasis>
(
<bibRefCitation id="4C724B004719FF83FE18CBDE43EA34A9" author="Sars" box="[469,646,152,178]" pageId="3" pageNumber="18" refString="Sars, M. (1861) Om de ved Norges Kyster forekommende Arter af Annelideslaegten Polynoe. Forhandlinger i Videnskabsselskabet i Kristiana, 54 - 62." type="book chapter" year="1861">M. Sars, 1861</bibRefCitation>
)
</taxonomicName>
together with 
<taxonomicName id="EFE34D724719FF83FC88CBDE45B834A9" authority="Hansen, 1880" authorityName="Hansen" authorityYear="1880" box="[837,1236,152,178]" class="Polychaeta" family="Polynoidae" genus="Polynoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="assimilis">
<emphasis id="1A97EAE34719FF83FC88CBDE457334AA" box="[837,1055,152,177]" italics="true" pageId="3" pageNumber="4">Polynoe assimilis</emphasis>
<bibRefCitation id="4C724B004719FF83FBE5CBDE45B834A9" author="Hansen" box="[1064,1236,152,178]" pageId="3" pageNumber="17" refString="Hansen, G. A. (1880) Annelider fra den norske Nordhavsexpedition i 1878. Nyt Magazin for Naturvidenskaberne, Oslo, 25, 224 - 234." type="journal article" year="1880">Hansen, 1880</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="EFE34D724719FF83FB2FCBDE400B34CC" authority="Malmgren, 1866" authorityName="Malmgren" authorityYear="1866" class="Polychaeta" family="Polynoidae" genus="Eunoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="oerstedi">
<emphasis id="1A97EAE34719FF83FB2FCBDE44F134AA" box="[1250,1437,152,177]" italics="true" pageId="3" pageNumber="4">Eunoe oerstedi</emphasis>
<bibRefCitation id="4C724B004719FF83FF5ACBFB400B34CC" author="Malmgren" box="[151,359,189,215]" pageId="3" pageNumber="17" refString="Malmgren, A. J. (1866) Nordiska Hafs-Annulata. Ofversigt af Konglia Vetenskaps-Akademiens Forhhandlingar Stockholm, 1, 51 - 110 [read 1865, printed 1866]." type="journal article" year="1866">Malmgren, 1866</bibRefCitation>
</taxonomicName>
and some other species, but without discussing this decision. 
<bibRefCitation id="4C724B004719FF83FBB5CBFB444534CC" author="Fauvel" box="[1144,1321,189,215]" pageId="3" pageNumber="17" refString="Fauvel, P. (1914) Annelides Polychetes non pelagiques provenant des Campagnes de l' &quot; Hirondelle &quot; et de la &quot; Princesse- Alice &quot; (1885 - 1910). Resultats des campagnes scientifiques accomplies sur son yacht par Albert I er Prince Souverain de Monaco, 46, 1 - 432." type="journal article" year="1914">Fauvel (1914)</bibRefCitation>
however considered 
<taxonomicName id="EFE34D724719FF83FEECCBA440E334E0" box="[289,399,226,251]" class="Polychaeta" family="Polynoidae" genus="Nychia" higherTaxonomySource="CoL" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE34719FF83FEECCBA440E334E0" box="[289,399,226,251]" italics="true" pageId="3" pageNumber="4">globifera</emphasis>
</taxonomicName>
valid, and was the first to discuss the generic placement. He put the species in 
<taxonomicName id="EFE34D724719FF83FA9DCBA4400D3539" authority="Malmgren, 1866" authorityName="Malmgren" authorityYear="1866" class="Polychaeta" family="Polynoidae" genus="Eunoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34719FF83FA9DCBA444F034E0" box="[1360,1436,226,251]" italics="true" pageId="3" pageNumber="4">Eunoe</emphasis>
<bibRefCitation id="4C724B004719FF83FF5ACA4E400D3539" author="Malmgren" box="[151,353,264,290]" pageId="3" pageNumber="17" refString="Malmgren, A. J. (1866) Nordiska Hafs-Annulata. Ofversigt af Konglia Vetenskaps-Akademiens Forhhandlingar Stockholm, 1, 51 - 110 [read 1865, printed 1866]." type="journal article" year="1866">Malmgren, 1866</bibRefCitation>
</taxonomicName>
because of the unidentate neurochaetae and exclusively stout notochaetae (in contrast to 
<taxonomicName id="EFE34D724719FF83FF5ACA6B4068355D" box="[151,260,301,326]" class="Polychaeta" family="Polynoidae" genus="Gattyana" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34719FF83FF5ACA6B4068355D" box="[151,260,301,326]" italics="true" pageId="3" pageNumber="4">Gattyana</emphasis>
</taxonomicName>
which has two kinds of notochaetae: stout with blunt and slender with capillary tip; see also Barnich &amp; Fiege 2009). 
<bibRefCitation id="4C724B004719FF83FE9FCA14437B3577" author="Ditlevsen" box="[338,535,338,364]" pageId="3" pageNumber="17" refString="Ditlevsen, H. (1917) Annelids. The Danish Ingolf Expedition, 4 (4), 1 - 71." type="journal article" year="1917">Ditlevsen (1917)</bibRefCitation>
finally checked the 
<typeStatus id="F75888534719FF83FCC5CA14422A3577" box="[776,838,338,364]" pageId="3" pageNumber="4">types</typeStatus>
of 
<taxonomicName id="EFE34D724719FF83FCA0CA1445503570" box="[877,1084,338,363]" class="Polychaeta" family="Polynoidae" genus="Polynoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="assimilis">
<emphasis id="1A97EAE34719FF83FCA0CA1445503570" box="[877,1084,338,363]" italics="true" pageId="3" pageNumber="4">Polynoe assimilis</emphasis>
</taxonomicName>
and confirmed the synonymy with 
<taxonomicName id="EFE34D724719FF83FF18CA3E4029358A" box="[213,325,376,401]" class="Polychaeta" family="Polynoidae" genus="Nychia" higherTaxonomySource="CoL" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE34719FF83FF18CA3E4029358A" box="[213,325,376,401]" italics="true" pageId="3" pageNumber="4">globifera</emphasis>
</taxonomicName>
, but he moved the species to 
<taxonomicName id="EFE34D724719FF83FD79CA3E4252358A" box="[692,830,376,401]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34719FF83FD79CA3E4252358A" box="[692,830,376,401]" italics="true" pageId="3" pageNumber="4">Harmothoe</emphasis>
</taxonomicName>
. The validity of 
<taxonomicName id="EFE34D724719FF83FBC0CA3E4511358A" box="[1037,1149,376,401]" class="Polychaeta" family="Polynoidae" genus="Nychia" higherTaxonomySource="CoL" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE34719FF83FBC0CA3E4511358A" box="[1037,1149,376,401]" italics="true" pageId="3" pageNumber="4">globifera</emphasis>
</taxonomicName>
and its synonymy with 
<taxonomicName id="EFE34D724719FF83FF5ACADB419235AD" box="[151,254,413,438]" class="Polychaeta" family="Polynoidae" genus="Polynoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="assimilis">
<emphasis id="1A97EAE34719FF83FF5ACADB419235AD" box="[151,254,413,438]" italics="true" pageId="3" pageNumber="4">assimilis</emphasis>
</taxonomicName>
was finally accepted by 
<bibRefCitation id="4C724B004719FF83FDD3CADB43B735AC" author="Augener" box="[542,731,413,439]" pageId="3" pageNumber="17" refString="Augener, H. (1929) Die Polychaten von Spitzbergen. Fauna Arctica, 5, 649 - 834." type="journal article" year="1929">Augener (1929)</bibRefCitation>
, but he still considered the species to belong to 
<taxonomicName id="EFE34D724719FF83FAC5CADB441935AD" box="[1288,1397,413,438]" class="Polychaeta" family="Polynoidae" genus="Gattyana" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34719FF83FAC5CADB441935AD" box="[1288,1397,413,438]" italics="true" pageId="3" pageNumber="4">Gattyana</emphasis>
</taxonomicName>
. In his unpublished thesis 
<bibRefCitation id="4C724B004719FF83FE6CCA84430F35C7" author="Loshamn" box="[417,611,450,476]" pageId="3" pageNumber="17" refString="Loshamn, A. A. (1980) En systematisk og dyregeografisk Undersokelse over Skjellrygg-Gruppen (Familie Aphroditidae sensu Fauvel 1923). Dissertation. Zoologisk Institutt, Universitete i Oslo, 184 pp. [unpublished]." type="book" year="1980">Loshamn (1980)</bibRefCitation>
finally placed it within 
<taxonomicName id="EFE34D724719FF83FCB1CA84456D35C0" box="[892,1025,450,475]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34719FF83FCB1CA84456D35C0" box="[892,1025,450,475]" italics="true" pageId="3" pageNumber="4">Harmothoe</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="285C36F14719FF83FF08CAAE458137FC" blockId="3.[151,1437,152,1484]" pageId="3" pageNumber="4">
The main reason for this disagreement in the literature is the shape of the tip in the neurochaetae. G.O. Sars describes them as being all unidentate; Loshamn however mentions some upper neurochaetae being minutely bidentate, a fact which we found confirmed in the recently examined specimens listed above. Since the secondary tooth is often abraded, the bidentate neurochaetae are easily overlooked, and thus the species could be mistaken for a 
<taxonomicName id="EFE34D724719FF83FE7DC93B4097368D" box="[432,507,637,662]" class="Polychaeta" family="Polynoidae" genus="Eunoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34719FF83FE7DC93B4097368D" box="[432,507,637,662]" italics="true" pageId="3" pageNumber="4">Eunoe</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="4C724B004719FF83FDF4C93B438E368C" author="Augener" box="[569,738,637,663]" pageId="3" pageNumber="17" refString="Augener, H. (1913) Polychaeten von Franz-Joseph-Land I. Zoologischer Anzeiger, 41 (5), 202 - 220." type="journal article" year="1913">Augener 1913</bibRefCitation>
, 
<bibRefCitation id="4C724B004719FF83FD23C93B42EE368C" author="Fauvel" box="[750,898,637,663]" pageId="3" pageNumber="17" refString="Fauvel, P. (1914) Annelides Polychetes non pelagiques provenant des Campagnes de l' &quot; Hirondelle &quot; et de la &quot; Princesse- Alice &quot; (1885 - 1910). Resultats des campagnes scientifiques accomplies sur son yacht par Albert I er Prince Souverain de Monaco, 46, 1 - 432." type="journal article" year="1914">Fauvel 1914</bibRefCitation>
, Barnich &amp; Fiege 2009). But our examination of several young and adult specimens showed, that a few bidentate neurochaetae are always present and even when abraded, this tip is different from a unidentate tip (see 
<figureCitation id="B0D82A744719FF83FC5FC98E42BB36F9" box="[914,983,712,738]" captionStart="FIGURE 1" captionStartId="4.[151,255,1880,1904]" captionTargetBox="[240,1348,202,1850]" captionTargetId="figure@4.[231,1353,194,1865]" captionTargetPageId="4" captionText="FIGURE 1. Harmothoe globifera (SMF 18887). (A) anterior end; (B) right anterior elytron from unknown segment, free in vial; (C) micro- and macrotubercles from same; (D) left elytragerous parapodium from segment 5, posterior view; (E) distal half of long notochaeta; (F) distal part of middle, unidentate neurochaeta; (G) tip of same; (H) tip of upper bidentate neurochaeta with fragile secondary tooth; (I) tip of upper bidentate neurochaeta with abraded secondary tooth; (J) tip of lower unidentate neurochaeta. (Scales: A, B = 2 mm; C = 250 µm; D = 500 µm; E, F = 100 µm; G – J = 50 µm)." httpUri="https://zenodo.org/record/196400/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
G–J). Therefore, we finally confirm Loshamn's opinion and place the species within 
<taxonomicName id="EFE34D724719FF83FCCCC9AB42FC371D" box="[769,912,749,774]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34719FF83FCCCC9AB42FC371D" box="[769,912,749,774]" italics="true" pageId="3" pageNumber="4">Harmothoe</emphasis>
</taxonomicName>
and not within 
<taxonomicName id="EFE34D724719FF83FB97C9AB45C6371D" box="[1114,1194,749,774]" class="Polychaeta" family="Polynoidae" genus="Eunoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34719FF83FB97C9AB45C6371D" box="[1114,1194,749,774]" italics="true" pageId="3" pageNumber="4">Eunoe</emphasis>
</taxonomicName>
. The most striking characters of 
<taxonomicName id="EFE34D724719FF83FEF9C85443403730" box="[308,556,786,811]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE34719FF83FEF9C85443403730" box="[308,556,786,811]" italics="true" pageId="3" pageNumber="4">Harmothoe globifera</emphasis>
</taxonomicName>
are found in the elytra with a densely papillate outer lateral margin, micro- and macrotubercles covered by nodular papillae, and in the densely papillate antennae and cirri (
<figureCitation id="B0D82A744719FF83FAC0C87E44233749" box="[1293,1359,824,850]" captionStart="FIGURE 1" captionStartId="4.[151,255,1880,1904]" captionTargetBox="[240,1348,202,1850]" captionTargetId="figure@4.[231,1353,194,1865]" captionTargetPageId="4" captionText="FIGURE 1. Harmothoe globifera (SMF 18887). (A) anterior end; (B) right anterior elytron from unknown segment, free in vial; (C) micro- and macrotubercles from same; (D) left elytragerous parapodium from segment 5, posterior view; (E) distal half of long notochaeta; (F) distal part of middle, unidentate neurochaeta; (G) tip of same; (H) tip of upper bidentate neurochaeta with fragile secondary tooth; (I) tip of upper bidentate neurochaeta with abraded secondary tooth; (J) tip of lower unidentate neurochaeta. (Scales: A, B = 2 mm; C = 250 µm; D = 500 µm; E, F = 100 µm; G – J = 50 µm)." httpUri="https://zenodo.org/record/196400/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
A–C). At first glance, the species could be confused with 
<taxonomicName id="EFE34D724719FF83FCC9C81B4514376C" authority="M. Sars, 1861" authorityName="M. Sars" authorityYear="1861" box="[772,1144,861,887]" class="Polychaeta" family="Polynoidae" genus="Eunoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="nodosa">
<emphasis id="1A97EAE34719FF83FCC9C81B42C3376D" box="[772,943,861,886]" italics="true" pageId="3" pageNumber="4">Eunoe nodosa</emphasis>
(
<bibRefCitation id="4C724B004719FF83FC0DC81B4502376C" author="Sars" box="[960,1134,861,887]" pageId="3" pageNumber="18" refString="Sars, M. (1861) Om de ved Norges Kyster forekommende Arter af Annelideslaegten Polynoe. Forhandlinger i Videnskabsselskabet i Kristiana, 54 - 62." type="book chapter" year="1861">M. Sars, 1861</bibRefCitation>
)
</taxonomicName>
, which has also nodular macrotubercles. But, 
<taxonomicName id="EFE34D724719FF83FE59C8C443663780" box="[404,522,898,923]" class="Polychaeta" family="Polynoidae" genus="Eunoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="nodosa">
<emphasis id="1A97EAE34719FF83FE59C8C443663780" box="[404,522,898,923]" italics="true" pageId="3" pageNumber="4">E. nodosa</emphasis>
</taxonomicName>
differs by the exclusively unidentate neurochaetae, the less densely papillate elytral margin, antennae and cirri, the microtubercles which are mostly low semiglobose and distally nodular, and the macrotubercles with nodular papillae only distally (
<figureCitation id="B0D82A744719FF83FC9DC88B42F837FC" box="[848,916,973,999]" captionStart="FIGURE 2" captionStartId="6.[151,256,1838,1862]" captionTargetBox="[231,1355,204,1812]" captionTargetId="figure@6.[227,1361,194,1825]" captionTargetPageId="6" captionText="FIGURE 2. Eunoe nodosa (ZMO C 5244). (A) anterior end, styles of median antenna, left tentacular and right dorsal tentacular cirri, and dorsal cirri missing; (B) right anterior elytron from unknown segment, free in vial; (C) micro- and macrotubercles at posterior margin from same; (D) micro-and macrotubercles at outer lateral margin from same; (E) right elytragerous parapodium from segment 21, posterior view; (F) distal half of long notochaeta; (G) tip of same; (H) distal part of middle neurochaeta; (I) tip of same; (J) tip of upper neurochaeta; (K) tip of lower neurochaeta. (Scales: A, B = 2 mm; C, D, F, H = 250 µm; E = 500 µm; G, I – K = 100 µm)." httpUri="https://zenodo.org/record/196401/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
A–K; see description below).
</paragraph>
</subSubSection>
<subSubSection id="60F9657A4719FF83FF08C8B444023199" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="285C36F14719FF83FF08C8B4419F3067" blockId="3.[151,1437,152,1484]" pageId="3" pageNumber="4">
<taxonomicName id="EFE34D724719FF83FF08C8B440D23010" ID-CoL="6LCFB" box="[197,446,1010,1035]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE34719FF83FF08C8B440D23010" box="[197,446,1010,1035]" italics="true" pageId="3" pageNumber="4">Harmothoe globifera</emphasis>
</taxonomicName>
could also be confused with 
<taxonomicName id="EFE34D724719FF83FCDEC8B445093017" ID-CoL="3JRDP" authority="M. Sars, 1863" authorityName="M. Sars" authorityYear="1863" box="[787,1125,1010,1036]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="clavigera">
<emphasis id="1A97EAE34719FF83FCDEC8B442C93010" box="[787,933,1010,1035]" italics="true" pageId="3" pageNumber="4">H. clavigera</emphasis>
(
<bibRefCitation id="4C724B004719FF83FC7BC8B445313017" author="Sars" box="[950,1117,1010,1036]" pageId="3" pageNumber="18" refString="Sars, M. (1863) Geologiske og zoologiske Iagttagelser, anstillede paa en Reise i en Deel af Trondhjems Stift i Sommeren 1862. II. Zoologiske Notitser fra Christiansund og Bejan. Nyt Magazin for Naturvidenskaberne, 12 (3), 273 - 340." type="journal article" year="1863">M. Sars, 1863</bibRefCitation>
)
</taxonomicName>
which was redescribed by Barnich &amp; Fiege (2009). In 
<taxonomicName id="EFE34D724719FF83FDC4CF5E43C2302A" ID-CoL="3JRDP" box="[521,686,1048,1073]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="clavigera">
<emphasis id="1A97EAE34719FF83FDC4CF5E43C2302A" box="[521,686,1048,1073]" italics="true" pageId="3" pageNumber="4">H. clavigera</emphasis>
</taxonomicName>
, however, the neurochaetae are mainly bidentate, elytral microtubercles are conical, and macrotubercles are rather large, club-shaped with nodular papillae only distally.
</paragraph>
<paragraph id="285C36F14719FF83FF08CFCE44023199" blockId="3.[151,1437,152,1484]" pageId="3" pageNumber="4">
<taxonomicName id="EFE34D724719FF83FF08CFCE40FD30BA" box="[197,401,1160,1185]" class="Magnoliopsida" family="Achariaceae" genus="Dasylepis" kingdom="Animalia" order="Malpighiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="equitis">
<emphasis id="1A97EAE34719FF83FF08CFCE40FD30BA" box="[197,401,1160,1185]" italics="true" pageId="3" pageNumber="4">Dasylepis equitis</emphasis>
</taxonomicName>
is only known from a posterior fragment described by 
<bibRefCitation id="4C724B004719FF83FBE8CFCE459D30B9" author="McIntosh" box="[1061,1265,1160,1186]" pageId="3" pageNumber="17" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. &quot; Challenger &quot; during the years 1873 - 76. Reports on the scientific results of the voyage of H. M. S. &quot; Challenger &quot;, Zoology, 12, 1 - 554." type="journal article" year="1885">McIntosh (1885)</bibRefCitation>
; this fragment seems to be lost, since the vial labelled &quot; 
<taxonomicName id="EFE34D724719FF83FD41CFEB423330DD" box="[652,863,1197,1222]" class="Magnoliopsida" family="Achariaceae" genus="Dasylepis" kingdom="Animalia" order="Malpighiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="equitis">
<emphasis id="1A97EAE34719FF83FD41CFEB423330DD" box="[652,863,1197,1222]" italics="true" pageId="3" pageNumber="4">Dasylepis equitis</emphasis>
</taxonomicName>
, 
<typeStatus id="F75888534719FF83FCA2CFEB42B130DC" box="[879,989,1197,1223]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
, BMNH 1885.12.1.57&quot; contains two opheliids. Unfortunately, McIntosh did not explain why he assigned his species to 
<taxonomicName id="EFE34D724719FF83FB90CF9544F730F7" authority="Malmgren, 1867" authorityName="Malmgren" authorityYear="1867" box="[1117,1435,1234,1260]" class="Magnoliopsida" family="Achariaceae" genus="Dasylepis" kingdom="Animalia" order="Malpighiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="1A97EAE34719FF83FB90CF9545BC30F7" box="[1117,1232,1235,1260]" italics="true" pageId="3" pageNumber="4">Dasylepis</emphasis>
<bibRefCitation id="4C724B004719FF83FB14CF9444F730F7" author="Malmgren" box="[1241,1435,1234,1260]" pageId="3" pageNumber="17" refString="Malmgren, A. J. (1867) Annulata polychaeta Spetsbergiae, Groenlandiae, Islandiae et Scandinaviae hactenus cognita. Ofversigt af Kongliga Vetenskaps-Akademiens Forhhandlingar Stockholm, 4, 127 - 235." type="journal article" year="1867">Malmgren, 1867</bibRefCitation>
</taxonomicName>
(name pre-occupied, replaced by 
<taxonomicName id="EFE34D724719FF83FDFECFBE42393109" ID-CoL="84L2T" authority="Norman" authorityName="Norman" box="[563,853,1272,1298]" class="Polychaeta" family="Polynoidae" genus="Acanthicolepis" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34719FF83FDFECFBE4385310A" box="[563,745,1272,1297]" italics="true" pageId="3" pageNumber="4">Acanthicolepis</emphasis>
Norman
</taxonomicName>
in 
<bibRefCitation id="4C724B004719FF83FCB3CFBE45293109" author="McIntosh" box="[894,1093,1272,1298]" pageId="3" pageNumber="17" refString="McIntosh, W. C. (1900) Polychaeta. Amphinomidae to Sigalionidae. A Monograph of the British Annelids, 1 (2), 215 - 444." type="journal article" year="1900">McIntosh, 1900</bibRefCitation>
). Already in our revision of 
<taxonomicName id="EFE34D724719FF83FF5ACE5B4339312C" ID-CoL="84L2T" authority="Barnich et al. 2000" authorityName="Barnich et al." authorityYear="2000" box="[151,597,1309,1335]" class="Polychaeta" family="Polynoidae" genus="Acanthicolepis" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="1A97EAE34719FF83FF5ACE5B4027312D" box="[151,331,1309,1334]" italics="true" pageId="3" pageNumber="4">Acanthicolepis</emphasis>
(
<bibRefCitation id="4C724B004719FF83FE91CE5B4320312C" author="Barnich" box="[348,588,1309,1335]" pageId="3" pageNumber="17" refString="Barnich, R., Gil, J. &amp; Fiege, D. (2000) Revision of the genus Acanthicolepis Norman in McIntosh, 1900 (Polychaeta, Polynoidae, Polynoinae). Sarsia, 85, 313 - 319." type="journal article" year="2000">Barnich et al. 2000</bibRefCitation>
)
</taxonomicName>
, we considered 
<taxonomicName id="EFE34D724719FF83FCD4CE5B4286312D" box="[793,1002,1309,1334]" class="Magnoliopsida" family="Achariaceae" genus="Dasylepis" kingdom="Animalia" order="Malpighiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="equitis">
<emphasis id="1A97EAE34719FF83FCD4CE5B4286312D" box="[793,1002,1309,1334]" italics="true" pageId="3" pageNumber="4">Dasylepis equitis</emphasis>
</taxonomicName>
a doubtful species, close to 
<taxonomicName id="EFE34D724719FF83FA8ACE5B40683147" class="Polychaeta" family="Polynoidae" genus="Nychia" higherTaxonomySource="CoL" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE34719FF83FA8ACE5B40683147" italics="true" pageId="3" pageNumber="4">Nychia globifera</emphasis>
</taxonomicName>
. Having now examined a number of specimens of 
<taxonomicName id="EFE34D724719FF83FCA7CE0542903147" box="[874,1020,1347,1372]" class="Polychaeta" family="Polynoidae" genus="Nychia" higherTaxonomySource="CoL" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE34719FF83FCA7CE0542903147" box="[874,1020,1347,1372]" italics="true" pageId="3" pageNumber="4">H. globifera</emphasis>
</taxonomicName>
and compared them to McIntosh's description of the dorsal cirri and elytra, we finally confirm the synonymy of 
<taxonomicName id="EFE34D724719FF83FBEECE2E45F9319A" box="[1059,1173,1384,1409]" class="Magnoliopsida" family="Achariaceae" genus="Dasylepis" kingdom="Animalia" order="Malpighiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="equitis">
<emphasis id="1A97EAE34719FF83FBEECE2E45F9319A" box="[1059,1173,1384,1409]" italics="true" pageId="3" pageNumber="4">D. equitis</emphasis>
</taxonomicName>
with 
<taxonomicName id="EFE34D724719FF83FB1ACE2E440A319A" box="[1239,1382,1384,1409]" class="Polychaeta" family="Polynoidae" genus="Nychia" higherTaxonomySource="CoL" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="globifera">
<emphasis id="1A97EAE34719FF83FB1ACE2E440A319A" box="[1239,1382,1384,1409]" italics="true" pageId="3" pageNumber="4">H. globifera</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="60F9657A4719FF83FF08CECB40AB31D7" pageId="3" pageNumber="4" type="distribution">
<paragraph id="285C36F14719FF83FF08CECB40AB31D7" blockId="3.[151,1437,152,1484]" pageId="3" pageNumber="4">
<emphasis id="1A97EAE34719FF83FF08CECB409031BC" bold="true" box="[197,508,1421,1447]" pageId="3" pageNumber="4">Distribution and habitat.</emphasis>
Arctic Ocean and Northeast Atlantic; on mud and clay, in 
<quantity id="EF1B9B144719FF83FB67CECB443831BC" box="[1194,1364,1421,1447]" metricMagnitude="3" metricUnit="m" metricValue="1.825" metricValueMax="3.4" metricValueMin="0.25" pageId="3" pageNumber="4" unit="m" value="1825.0" valueMax="3400.0" valueMin="250.0">250 to 3400 m</quantity>
depth (see also 
<bibRefCitation id="4C724B004719FF83FEC9CEF440D631D7" author="Ditlevsen" box="[260,442,1458,1484]" pageId="3" pageNumber="17" refString="Ditlevsen, H. (1917) Annelids. The Danish Ingolf Expedition, 4 (4), 1 - 71." type="journal article" year="1917">Ditlevsen 1917</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>