338 lines
48 KiB
XML
338 lines
48 KiB
XML
<document id="9FD63F52B1E67DB4971B1A4B45BD7134" ID-CLB-Dataset="7105" ID-DOI="10.12782/specdiv.26.7" ID-GBIF-Dataset="5dc29412-7908-479d-93e6-1ecc88fdc1fd" ID-ISSN="2189-7301" ID-Zenodo-Dep="4585295" ID-ZooBank="368822BA-78A5-44BC-9C15-2DCB77047D7E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614947948901" checkinUser="felipe" docAuthor="Arai, Mikihito & Fujita, Toshihiko" docDate="2021" docId="A0103577FF88FFB4FED3D71D7EF30799" docLanguage="en" docName="SpeciesDiversity.26.1.7-21.pdf" docOrigin="Species Diversity 26 (1)" docSource="http://dx.doi.org/10.12782/specdiv.26.7" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Bathyferdina caelator Arai & Fujita 2021, sp. nov." docType="treatment" docUuid="28C6B460-B355-4601-AC8B-8E0940FD65CF" docUuidSource="ZooBank" docVersion="8" lastPageNumber="11" masterDocId="5C294D0FFF8BFFB0FFE7D1017D2A0208" masterDocTitle="Sea Stars of Families Ophidiasteridae and Goniasteridae (Echinodermata: Asteroidea) from the Mesophotic Zone of the Ogasawara Islands, Including Two New Species" masterLastPageNumber="21" masterPageNumber="7" pageNumber="10" updateTime="1698932841729" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="0CDEB9B2C7E5DF9D0BF76FAE029447F8">
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<mods:title id="CB6157C35415E068152F601848D991F7">Sea Stars of Families Ophidiasteridae and Goniasteridae (Echinodermata: Asteroidea) from the Mesophotic Zone of the Ogasawara Islands, Including Two New Species</mods:title>
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<mods:name id="9D7E6CC9FF343EB7F68380C0F9736683" type="personal">
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<mods:roleTerm id="E34AEBB553BD0D0D6792043EDAEFC7F3">Author</mods:roleTerm>
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<mods:namePart id="56E83BC45224E926C00710C6E4A8290B">Arai, Mikihito</mods:namePart>
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<mods:affiliation id="94F6DA76D49ABC39AB945B7F31D14427">Department of Biological Sciences, Graduate School of Science, the University of Tokyo, 7 - 3 - 1 Hongo, Bunkyo-ku, Tokyo 113 - 8654, Japan E-mail: arai @ kahaku. go. jp & National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan & Corresponding author</mods:affiliation>
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<mods:nameIdentifier id="8D9E352308CE1DF58C15F07BA2E60AC9" type="email">arai@kahaku.go.jp</mods:nameIdentifier>
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<mods:roleTerm id="02DC9CE191E0C4DC8266B56786F18386">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B8B072E69E91D966EA62F1369FCB8038">Fujita, Toshihiko</mods:namePart>
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<mods:affiliation id="DF64F5BCF52AE10C5E21A57254F6C6BF">Department of Biological Sciences, Graduate School of Science, the University of Tokyo, 7 - 3 - 1 Hongo, Bunkyo-ku, Tokyo 113 - 8654, Japan E-mail: arai @ kahaku. go. jp & National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan</mods:affiliation>
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<mods:nameIdentifier id="620B7AF8C398810A077D277FC2050381" type="email">arai@kahaku.go.jp</mods:nameIdentifier>
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<mods:typeOfResource id="D889F0273FD0F7E8287914C2CFF30669">text</mods:typeOfResource>
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<mods:titleInfo id="92538560CB6989BD0556C3949EE2217A">
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<mods:title id="AC296B0F5A8E29BA1B0704DDE421FDCD">Species Diversity</mods:title>
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</mods:titleInfo>
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<mods:part id="CE56BD89348E9B9A19A996EA72F7C069">
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<mods:date id="BD2168514D3E5F27C2B4F556BC230DA0">2021</mods:date>
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<mods:number id="CE67B12578A8A9A7EB0BAB2F5A83FF3A">2021-01-01</mods:number>
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<mods:number id="94062860221E4387851559186F6454CE">26</mods:number>
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<mods:url id="F979B315E8CF2219B8C86DEBF1F44ABC">http://dx.doi.org/10.12782/specdiv.26.7</mods:url>
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<treatment id="A0103577FF88FFB4FED3D71D7EF30799" ID-DOI="http://doi.org/10.5281/zenodo.4734680" ID-GBIF-Taxon="181851167" ID-Zenodo-Dep="4734680" LSID="urn:lsid:zoobank.org:act:28C6B460-B355-4601-AC8B-8E0940FD65CF" httpUri="http://treatment.plazi.org/id/A0103577FF88FFB4FED3D71D7EF30799" lastPageId="4" lastPageNumber="11" pageId="3" pageNumber="10">
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<subSubSection id="60A3D7EAFF88FFB3FED3D71D7F22047E" pageId="3" pageNumber="10" type="nomenclature">
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<paragraph id="28068461FF88FFB3FED3D71D7F42043C" blockId="3.[308,616,1564,1588]" box="[308,616,1564,1588]" pageId="3" pageNumber="10">
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<heading id="734E330DFF88FFB3FED3D71D7F42043C" box="[308,616,1564,1588]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="10" reason="2">
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<taxonomicName id="EFB9FFE2FF88FFB3FED3D71D7F3C043C" ID-CoL="846ZB" authority="Arai & Fujita, 2021" authorityName="Arai & Fujita" authorityYear="2021" box="[308,534,1564,1588]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="3" pageNumber="10" phylum="Echinodermata" rank="species" species="caelator" status="sp. nov.">
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<emphasis id="1ACD5873FF88FFB3FED3D71D7F3C043C" bold="true" box="[308,534,1564,1588]" italics="true" pageId="3" pageNumber="10">Bathyferdina caelator</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="01FEE508FF88FFB3FDFBD71C7F42043C" box="[540,616,1565,1588]" pageId="3" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="28068461FF88FFB3FF0AD73F7F22047E" blockId="3.[237,686,1597,1654]" pageId="3" pageNumber="10">
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[New Japanese name: Koyo-akamon-hitode] (
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<figureCitation id="B08298E4FF88FFB3FE7ED75F7CC0047E" box="[409,490,1630,1654]" captionStart="Fig" captionStartId="2.[822,853,1874,1896]" captionTargetBox="[188,1402,207,2036]" captionTargetId="figure-6@2.[187,1400,205,2029]" captionTargetPageId="2" captionText="Fig. 2. Abactinal view of the live specimens (A–D) and ethanol-preserved specimen (E). A, Bathyferdina caelator sp. nov., NSMT E-8265, holotype; B, Fromia eusticha, NSMT E-10595; C, Fromia labeosa sp. nov., NSMT E-9297, holotype; D, Ogmaster capella, NSMT E-9312; E, Tamaria tenella, NSMT E-9277. Scale bars: 10mm." figureDoi="http://doi.org/10.5281/zenodo.4585299" httpUri="https://zenodo.org/record/4585299/files/figure.png" pageId="3" pageNumber="10">Figs 2A</figureCitation>
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,
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<figureCitation id="B08298E4FF88FFB3FE13D75F7F2B047E" box="[500,513,1630,1654]" captionStart="Fig" captionStartId="3.[157,188,1389,1411]" captionTargetBox="[188,1398,206,1350]" captionTargetId="figure-270@3.[187,1399,205,1351]" captionTargetPageId="3" captionText="Fig. 3. Bathyferdina caelator sp. nov., NSMT E-8265, holotype. A, actinal side; B, abactinal side, abactinal plates partly denuded. Arrows indicate pedicellariae on superomarginal plates; C, arm tip, lateral view. Arrow indicates terminal plate; D, SEM picture of a denuded abactinal plate at proximal portion of the arm; E, pedicellaria on actinal interradial plates; F, actinal, adambulacral and oral plates, partly denuded. Arrow indicates glassy boss on adambulacral plate. Scale bars: 10mm (A), 1mm (B, F), 5mm (C), 0.5mm (D, E). Abbreviations: ab, abactinal plates; sm, superomarginal plates; im, inferomarginal plates; ac, actinal plates; ad, adambulacral plates; op, oral plates." figureDoi="http://doi.org/10.5281/zenodo.4585301" httpUri="https://zenodo.org/record/4585301/files/figure.png" pageId="3" pageNumber="10">3</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="60A3D7EAFF88FFB3FF7BD79E7F6004BF" box="[156,586,1695,1719]" pageId="3" pageNumber="10" type="reference_group">
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<paragraph id="28068461FF88FFB3FF7BD79E7F6004BF" blockId="3.[156,586,1695,1719]" box="[156,586,1695,1719]" pageId="3" pageNumber="10">
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<treatmentCitationGroup id="08A9A34FFF88FFB3FF7BD79E7F6004BF" box="[156,586,1695,1719]" pageId="3" pageNumber="10">
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<taxonomicName id="EFB9FFE2FF88FFB3FF7BD79E7C3704BF" ID-CoL="7NSQG" authorityName="Mah" authorityYear="2017" box="[156,285,1695,1719]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="3" pageNumber="10" phylum="Echinodermata" rank="genus">
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<emphasis id="1ACD5873FF88FFB3FF7BD79E7C3704BF" box="[156,285,1695,1719]" italics="true" pageId="3" pageNumber="10">Bathyferdina</emphasis>
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</taxonomicName>
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sp.:
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<treatmentCitation id="A918A270FF88FFB3FEAED79E7F6F04BF" author="Arai, M. & Tanaka, Y. & Miyazaki, T. & Fujita, T." box="[329,581,1695,1719]" page="194" pageId="3" pageNumber="10" year="2018">
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<bibRefCitation id="4C28F990FF88FFB3FEAED79E7F6F04BF" author="Arai, M. & Tanaka, Y. & Miyazaki, T. & Fujita, T." box="[329,581,1695,1719]" pageId="3" pageNumber="10" pagination="191 - 203" refId="ref10506" refString="Arai, M., Tanaka, Y., Miyazaki, T., and Fujita, T. 2018. Valvatida (Echinodermata: Asteroidea) of the Ogasawara Islands, Japan. Memoirs of the National Museum of Nature and Science 52: 191 - 203. [In Japanese]" type="journal article" year="2018">Arai et al. 2018: 194–196</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="60A3D7EAFF88FFB3FF52D7E17CE10572" pageId="3" pageNumber="10" type="materials_examined">
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<paragraph id="28068461FF88FFB3FF52D7E17CE10572" blockId="3.[156,768,1760,2077]" pageId="3" pageNumber="10">
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<emphasis id="1ACD5873FF88FFB3FF52D7E17CA904F0" bold="true" box="[181,387,1760,1784]" pageId="3" pageNumber="10">Material examined.</emphasis>
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<materialsCitation id="98D18E3CFF88FFB3FE6CD7E17FA00510" ID-GBIF-Occurrence="3097624302" collectionCode="NSMT" elevation="136" location="Northwest of Ototo-jima Island" pageId="3" pageNumber="10" specimenCount="1" typeStatus="holotype">
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<typeStatus id="F7023AC3FF88FFB3FE6CD7E17CC604F0" box="[395,492,1760,1784]" pageId="3" pageNumber="10" type="holotype">Holotype</typeStatus>
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||
:
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<collectionCode id="4EA81CA4FF88FFB3FE11D7E17F1104F0" box="[502,571,1760,1784]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="3" pageNumber="10">NSMT</collectionCode>
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E-8265, KY-09-21,
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<location id="2D66D2BAFF88FFB3FF7BD6017CC30510" LSID="urn:lsid:plazi:treatment:A0103577FF88FFB4FED3D71D7EF30799:2D66D2BAFF88FFB3FF7BD6017CC30510" box="[156,489,1792,1816]" name="Northwest of Ototo-jima Island" pageId="3" pageNumber="10">Northwest of Ototo-jima Island</location>
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,
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<quantity id="EF412984FF88FFB3FE11D6017FA00510" box="[502,650,1792,1816]" metricMagnitude="2" metricUnit="m" metricValue="1.3565" metricValueMax="1.358" metricValueMin="1.355" pageId="3" pageNumber="10" unit="m" value="135.65" valueMax="135.8" valueMin="135.5">
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<elevation id="A3946352FF88FFB3FE11D6017FA00510" box="[502,650,1792,1816]" metricMagnitude="2" metricUnit="m" metricValue="1.3565" metricValueMax="1.358" metricValueMin="1.355" pageId="3" pageNumber="10" unit="m" value="135.65" valueMax="135.8" valueMin="135.5">135.5–135.8m</elevation>
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</quantity>
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</materialsCitation>
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||
.
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<typeStatus id="F7023AC3FF88FFB3FD70D6007FD10511" box="[663,763,1793,1817]" pageId="3" pageNumber="10" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="4EA81CA4FF88FFB3FF7BD6207DCB0531" box="[156,225,1825,1849]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="3" pageNumber="10">NSMT</collectionCode>
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E-8266, 1 individual, KT-09-2-TW1-1, West of Chichi-jima Island,
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<quantity id="EF412984FF88FFB3FE89D6407F2B0551" box="[366,513,1857,1881]" metricMagnitude="2" metricUnit="m" metricValue="1.4189999999999998" metricValueMax="1.452" metricValueMin="1.386" pageId="3" pageNumber="10" unit="m" value="141.89999999999998" valueMax="145.2" valueMin="138.6">138.6–145.2 m</quantity>
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;
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<collectionCode id="4EA81CA4FF88FFB3FDF7D6437F7F0552" box="[528,597,1858,1882]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="3" pageNumber="10">NSMT</collectionCode>
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E-8267, 1 individual, dry, locality unknown.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="60A3D7EAFF88FFB3FF52D6837ED1049E" pageId="3" pageNumber="10" type="diagnosis">
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||
<paragraph id="28068461FF88FFB3FF52D6837ED1049E" blockId="3.[156,768,1760,2077]" lastBlockId="3.[821,1432,1565,2077]" pageId="3" pageNumber="10">
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||
<emphasis id="1ACD5873FF88FFB3FF52D6837C0F0592" bold="true" box="[181,293,1922,1946]" pageId="3" pageNumber="10">Diagnosis.</emphasis>
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A species of
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<taxonomicName id="EFB9FFE2FF88FFB3FE5FD6827F130593" authorityName="Mah" authorityYear="2017" box="[440,569,1923,1947]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="3" pageNumber="10" phylum="Echinodermata" rank="genus">
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<emphasis id="1ACD5873FF88FFB3FE5FD6827F130593" box="[440,569,1923,1947]" italics="true" pageId="3" pageNumber="10">Bathyferdina</emphasis>
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</taxonomicName>
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with glassy bosses on abactinal, actinal and adambulacral plates. Disc broad with R/r 2.0–2.3. Abactinal plates are homogeneous in size and shape. Superomarginal plates rectangular to barrel-like in outline, and regularly decreasing in size from the interradius to the tip of arms. Glassy boss/ridge present on abactinal and actinal plates. One or two glassy bosses on the center of each adambulacral plate. Three furrow spines on each adambulacral plate.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="60A3D7EAFF88FFB4FCA9D79E7C1705F4" lastPageId="4" lastPageNumber="11" pageId="3" pageNumber="10" type="description">
|
||
<paragraph id="28068461FF88FFB3FCA9D79E78690531" blockId="3.[821,1432,1565,2077]" pageId="3" pageNumber="10">
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<emphasis id="1ACD5873FF88FFB3FCA9D79E797F04BF" bold="true" box="[846,1109,1695,1719]" pageId="3" pageNumber="10">
|
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Description of
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||
<typeStatus id="F7023AC3FF88FFB3FC15D79E797B04BF" box="[1010,1105,1695,1719]" pageId="3" pageNumber="10" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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R=
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<quantity id="EF412984FF88FFB3FB64D79E79F604BE" box="[1155,1244,1695,1719]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="3" pageNumber="10" unit="mm" value="23.2">23.2 mm</quantity>
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||
, r=
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||
<quantity id="EF412984FF88FFB3FAE0D79E787E04BE" box="[1287,1364,1695,1719]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="3" pageNumber="10" unit="mm" value="9.9">9.9 mm</quantity>
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||
, R/r= 2.3, width of arm is
|
||
<quantity id="EF412984FF88FFB3FBE9D7BE794F04DF" box="[1038,1125,1727,1751]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="3" pageNumber="10" unit="mm" value="11.1">11.1 mm</quantity>
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||
at base,
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||
<quantity id="EF412984FF88FFB3FB22D7BE782504DF" box="[1221,1295,1727,1751]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="3" pageNumber="10" unit="mm" value="6.6">6.6 mm</quantity>
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||
at half of R, and
|
||
<quantity id="EF412984FF88FFB3FC82D7E17E9A04FF" box="[869,944,1760,1784]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="10" unit="mm" value="3.1">3.1 mm</quantity>
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at 1/10 R from the tip. Body is flat on both abactinal and actinal sides. Arms are five and tapering more greatly near the disc than near the arm tip (
|
||
<figureCitation id="B08298E4FF88FFB3FB0ED620781D0531" box="[1257,1335,1825,1849]" captionStart="Fig" captionStartId="3.[157,188,1389,1411]" captionTargetBox="[188,1398,206,1350]" captionTargetId="figure-270@3.[187,1399,205,1351]" captionTargetPageId="3" captionText="Fig. 3. Bathyferdina caelator sp. nov., NSMT E-8265, holotype. A, actinal side; B, abactinal side, abactinal plates partly denuded. Arrows indicate pedicellariae on superomarginal plates; C, arm tip, lateral view. Arrow indicates terminal plate; D, SEM picture of a denuded abactinal plate at proximal portion of the arm; E, pedicellaria on actinal interradial plates; F, actinal, adambulacral and oral plates, partly denuded. Arrow indicates glassy boss on adambulacral plate. Scale bars: 10mm (A), 1mm (B, F), 5mm (C), 0.5mm (D, E). Abbreviations: ab, abactinal plates; sm, superomarginal plates; im, inferomarginal plates; ac, actinal plates; ad, adambulacral plates; op, oral plates." figureDoi="http://doi.org/10.5281/zenodo.4585301" httpUri="https://zenodo.org/record/4585301/files/figure.png" pageId="3" pageNumber="10">Fig. 3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="28068461FF88FFB4FCA9D6407FC4000B" blockId="3.[821,1432,1565,2077]" lastBlockId="4.[156,768,198,2077]" lastPageId="4" lastPageNumber="11" pageId="3" pageNumber="10">
|
||
Abactinal plates are polygonal to elliptical in shape, not lobed, variable in size, and tessellated so that only small spaces are left for papulae around the corners of these plates (
|
||
<figureCitation id="B08298E4FF88FFB3FCDBD6A27EA005B3" box="[828,906,1955,1979]" captionStart="Fig" captionStartId="3.[157,188,1389,1411]" captionTargetBox="[188,1398,206,1350]" captionTargetId="figure-270@3.[187,1399,205,1351]" captionTargetPageId="3" captionText="Fig. 3. Bathyferdina caelator sp. nov., NSMT E-8265, holotype. A, actinal side; B, abactinal side, abactinal plates partly denuded. Arrows indicate pedicellariae on superomarginal plates; C, arm tip, lateral view. Arrow indicates terminal plate; D, SEM picture of a denuded abactinal plate at proximal portion of the arm; E, pedicellaria on actinal interradial plates; F, actinal, adambulacral and oral plates, partly denuded. Arrow indicates glassy boss on adambulacral plate. Scale bars: 10mm (A), 1mm (B, F), 5mm (C), 0.5mm (D, E). Abbreviations: ab, abactinal plates; sm, superomarginal plates; im, inferomarginal plates; ac, actinal plates; ad, adambulacral plates; op, oral plates." figureDoi="http://doi.org/10.5281/zenodo.4585301" httpUri="https://zenodo.org/record/4585301/files/figure.png" pageId="3" pageNumber="10">Fig. 3B</figureCitation>
|
||
). Those on the disc are about 1.5–2.0 times larger than those on the arms. The arrangement of abactinal plates is regular, and median rows of plates are conspicuous. Between two upper interradial corners of first superomarginal plates on a ray, there are seven to nine abactinal plates. The entire abactinal surface of abactinal plates is rough with glassy bosses which are circular around the center of the plate and elongated around the periphery (
|
||
<figureCitation id="B08298E4FF8FFFB4FDB6D0297F8B0348" box="[593,673,296,320]" captionStart="Fig" captionStartId="3.[157,188,1389,1411]" captionTargetBox="[188,1398,206,1350]" captionTargetId="figure-270@3.[187,1399,205,1351]" captionTargetPageId="3" captionText="Fig. 3. Bathyferdina caelator sp. nov., NSMT E-8265, holotype. A, actinal side; B, abactinal side, abactinal plates partly denuded. Arrows indicate pedicellariae on superomarginal plates; C, arm tip, lateral view. Arrow indicates terminal plate; D, SEM picture of a denuded abactinal plate at proximal portion of the arm; E, pedicellaria on actinal interradial plates; F, actinal, adambulacral and oral plates, partly denuded. Arrow indicates glassy boss on adambulacral plate. Scale bars: 10mm (A), 1mm (B, F), 5mm (C), 0.5mm (D, E). Abbreviations: ab, abactinal plates; sm, superomarginal plates; im, inferomarginal plates; ac, actinal plates; ad, adambulacral plates; op, oral plates." figureDoi="http://doi.org/10.5281/zenodo.4585301" httpUri="https://zenodo.org/record/4585301/files/figure.png" pageId="4" pageNumber="11">Fig. 3D</figureCitation>
|
||
). Madreporite is single,
|
||
<quantity id="EF412984FF8FFFB4FEA6D0487C530368" box="[321,377,329,353]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="11" unit="mm" value="1.0">1 mm</quantity>
|
||
in diameter, flat and pentagonal with rounded corners; gyri extend radially from the central area where nine discontinuous, short gyri lie in parallel to one another. Terminal plates are conical with a broad base and a narrow rounded apex, smaller than the distalmost marginal plates, smooth and bare without skin or granules (
|
||
<figureCitation id="B08298E4FF8FFFB4FD72D0EA7FCB000B" box="[661,737,491,515]" captionStart="Fig" captionStartId="3.[157,188,1389,1411]" captionTargetBox="[188,1398,206,1350]" captionTargetId="figure-270@3.[187,1399,205,1351]" captionTargetPageId="3" captionText="Fig. 3. Bathyferdina caelator sp. nov., NSMT E-8265, holotype. A, actinal side; B, abactinal side, abactinal plates partly denuded. Arrows indicate pedicellariae on superomarginal plates; C, arm tip, lateral view. Arrow indicates terminal plate; D, SEM picture of a denuded abactinal plate at proximal portion of the arm; E, pedicellaria on actinal interradial plates; F, actinal, adambulacral and oral plates, partly denuded. Arrow indicates glassy boss on adambulacral plate. Scale bars: 10mm (A), 1mm (B, F), 5mm (C), 0.5mm (D, E). Abbreviations: ab, abactinal plates; sm, superomarginal plates; im, inferomarginal plates; ac, actinal plates; ad, adambulacral plates; op, oral plates." figureDoi="http://doi.org/10.5281/zenodo.4585301" httpUri="https://zenodo.org/record/4585301/files/figure.png" pageId="4" pageNumber="11">Fig. 3C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="28068461FF8FFFB4FF52D30D7FD90140" blockId="4.[156,768,198,2077]" pageId="4" pageNumber="11">
|
||
Superomarginal and inferomarginal plates correspond in number and size. There are six (seven on one arm) plates to each side of an arm. Superomarginal plates are rectangular in outline, longer than width, rounded at corners, and conspicuous on the abactinal side; first superomarginal plates are about
|
||
<quantity id="EF412984FF8FFFB4FEE4D3AF7C1100CE" box="[259,315,686,710]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="11" unit="mm" value="3.0">3 mm</quantity>
|
||
in length and
|
||
<quantity id="EF412984FF8FFFB4FE34D3AF7F2100CE" box="[467,523,686,710]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="11" unit="mm" value="2.0">2 mm</quantity>
|
||
in width on the abactinal surface. They gradually decrease in size toward the arm tip where the distalmost superomarginal plates are more squarish, measuring about
|
||
<quantity id="EF412984FF8FFFB4FE27D2117F0E012F" box="[448,548,784,808]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="4" pageNumber="11" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
|
||
in both length and width. No glassy boss was observed on any marginal plates.
|
||
</paragraph>
|
||
<paragraph id="28068461FF8FFFB4FF52D2507FF601E3" blockId="4.[156,768,198,2077]" pageId="4" pageNumber="11">
|
||
Actinal plates are quadrangular, regularly arranged, and normally flat except several small plates slightly convex between larger actinal plates (
|
||
<figureCitation id="B08298E4FF8FFFB4FE28D2937F3501A2" box="[463,543,914,938]" captionStart="Fig" captionStartId="3.[157,188,1389,1411]" captionTargetBox="[188,1398,206,1350]" captionTargetId="figure-270@3.[187,1399,205,1351]" captionTargetPageId="3" captionText="Fig. 3. Bathyferdina caelator sp. nov., NSMT E-8265, holotype. A, actinal side; B, abactinal side, abactinal plates partly denuded. Arrows indicate pedicellariae on superomarginal plates; C, arm tip, lateral view. Arrow indicates terminal plate; D, SEM picture of a denuded abactinal plate at proximal portion of the arm; E, pedicellaria on actinal interradial plates; F, actinal, adambulacral and oral plates, partly denuded. Arrow indicates glassy boss on adambulacral plate. Scale bars: 10mm (A), 1mm (B, F), 5mm (C), 0.5mm (D, E). Abbreviations: ab, abactinal plates; sm, superomarginal plates; im, inferomarginal plates; ac, actinal plates; ad, adambulacral plates; op, oral plates." figureDoi="http://doi.org/10.5281/zenodo.4585301" httpUri="https://zenodo.org/record/4585301/files/figure.png" pageId="4" pageNumber="11">Fig. 3F</figureCitation>
|
||
) and inferomarginal plates are slightly convex. The surface of actinal plates is rough with glassy bosses when denuded of granular skin.
|
||
</paragraph>
|
||
<paragraph id="28068461FF8FFFB4FF52D2F57F7806C7" blockId="4.[156,768,198,2077]" pageId="4" pageNumber="11">
|
||
Adambulacral armature is composed of three (distally two) furrow spines on each adambulacral plate (
|
||
<figureCitation id="B08298E4FF8FFFB4FD42D5157FDE0624" box="[677,756,1044,1068]" captionStart="Fig" captionStartId="3.[157,188,1389,1411]" captionTargetBox="[188,1398,206,1350]" captionTargetId="figure-270@3.[187,1399,205,1351]" captionTargetPageId="3" captionText="Fig. 3. Bathyferdina caelator sp. nov., NSMT E-8265, holotype. A, actinal side; B, abactinal side, abactinal plates partly denuded. Arrows indicate pedicellariae on superomarginal plates; C, arm tip, lateral view. Arrow indicates terminal plate; D, SEM picture of a denuded abactinal plate at proximal portion of the arm; E, pedicellaria on actinal interradial plates; F, actinal, adambulacral and oral plates, partly denuded. Arrow indicates glassy boss on adambulacral plate. Scale bars: 10mm (A), 1mm (B, F), 5mm (C), 0.5mm (D, E). Abbreviations: ab, abactinal plates; sm, superomarginal plates; im, inferomarginal plates; ac, actinal plates; ad, adambulacral plates; op, oral plates." figureDoi="http://doi.org/10.5281/zenodo.4585301" httpUri="https://zenodo.org/record/4585301/files/figure.png" pageId="4" pageNumber="11">Fig. 3F</figureCitation>
|
||
). Furrow spines are truncated at the tip, prismatic, and quadrangular or triangular in a cross section. The adambulacral plates are rectangular with slightly rounded corners. On the proximal first to tenth adambulacral plates, there are one or two glassy bosses at the center of the plates.
|
||
</paragraph>
|
||
<paragraph id="28068461FF8FFFB4FF52D5D67C3F049E" blockId="4.[156,768,198,2077]" pageId="4" pageNumber="11">
|
||
Abactinal, actinal and adambulacral plates are covered with a thin skin concealed by fine granules. On the marginal plates, the skin and granules are limited at the periphery, leaving the rest of the plate surface exposed (
|
||
<figureCitation id="B08298E4FF8FFFB4FD42D4387FD90759" box="[677,755,1337,1361]" captionStart="Fig" captionStartId="3.[157,188,1389,1411]" captionTargetBox="[188,1398,206,1350]" captionTargetId="figure-270@3.[187,1399,205,1351]" captionTargetPageId="3" captionText="Fig. 3. Bathyferdina caelator sp. nov., NSMT E-8265, holotype. A, actinal side; B, abactinal side, abactinal plates partly denuded. Arrows indicate pedicellariae on superomarginal plates; C, arm tip, lateral view. Arrow indicates terminal plate; D, SEM picture of a denuded abactinal plate at proximal portion of the arm; E, pedicellaria on actinal interradial plates; F, actinal, adambulacral and oral plates, partly denuded. Arrow indicates glassy boss on adambulacral plate. Scale bars: 10mm (A), 1mm (B, F), 5mm (C), 0.5mm (D, E). Abbreviations: ab, abactinal plates; sm, superomarginal plates; im, inferomarginal plates; ac, actinal plates; ad, adambulacral plates; op, oral plates." figureDoi="http://doi.org/10.5281/zenodo.4585301" httpUri="https://zenodo.org/record/4585301/files/figure.png" pageId="4" pageNumber="11">Fig. 3B</figureCitation>
|
||
). Papulae are isolated, confined around the abactinal plates except in the abactinal interradial areas where they are absent. There are no papulae on the actinal surface. Most papulae occur in spaces where corners of three abactinal plates meet. There are pedicellariae on some of the first or second superomarginal plate and actinal interradial plates (
|
||
<figureCitation id="B08298E4FF8FFFB4FD40D4FD7FD9041C" box="[679,755,1532,1556]" captionStart="Fig" captionStartId="3.[157,188,1389,1411]" captionTargetBox="[188,1398,206,1350]" captionTargetId="figure-270@3.[187,1399,205,1351]" captionTargetPageId="3" captionText="Fig. 3. Bathyferdina caelator sp. nov., NSMT E-8265, holotype. A, actinal side; B, abactinal side, abactinal plates partly denuded. Arrows indicate pedicellariae on superomarginal plates; C, arm tip, lateral view. Arrow indicates terminal plate; D, SEM picture of a denuded abactinal plate at proximal portion of the arm; E, pedicellaria on actinal interradial plates; F, actinal, adambulacral and oral plates, partly denuded. Arrow indicates glassy boss on adambulacral plate. Scale bars: 10mm (A), 1mm (B, F), 5mm (C), 0.5mm (D, E). Abbreviations: ab, abactinal plates; sm, superomarginal plates; im, inferomarginal plates; ac, actinal plates; ad, adambulacral plates; op, oral plates." figureDoi="http://doi.org/10.5281/zenodo.4585301" httpUri="https://zenodo.org/record/4585301/files/figure.png" pageId="4" pageNumber="11">Fig. 3B</figureCitation>
|
||
). Each pedicellaria with a single fan-shaped valve rather than two or more valves normally observed in
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FDABD73C7F96045D" authorityName="Mah" authorityYear="2017" box="[588,700,1597,1621]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="subFamily" subFamily="Ferdininae">Ferdininae</taxonomicName>
|
||
. Some alveoli lacking a fan-shaped valve are present on some marginal plates.
|
||
</paragraph>
|
||
<paragraph id="28068461FF8FFFB4FF52D79E7C7404F0" blockId="4.[156,768,198,2077]" pageId="4" pageNumber="11">
|
||
Color in life is brownish on the abactinal surface and white on the actinal. Bare areas on marginal plates are darker brown (
|
||
<figureCitation id="B08298E4FF8FFFB4FEE3D7E17C7904F0" box="[260,339,1760,1784]" captionStart="Fig" captionStartId="2.[822,853,1874,1896]" captionTargetBox="[188,1402,207,2036]" captionTargetId="figure-6@2.[187,1400,205,2029]" captionTargetPageId="2" captionText="Fig. 2. Abactinal view of the live specimens (A–D) and ethanol-preserved specimen (E). A, Bathyferdina caelator sp. nov., NSMT E-8265, holotype; B, Fromia eusticha, NSMT E-10595; C, Fromia labeosa sp. nov., NSMT E-9297, holotype; D, Ogmaster capella, NSMT E-9312; E, Tamaria tenella, NSMT E-9277. Scale bars: 10mm." figureDoi="http://doi.org/10.5281/zenodo.4585299" httpUri="https://zenodo.org/record/4585299/files/figure.png" pageId="4" pageNumber="11">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="28068461FF8FFFB4FF52D6017CB10592" blockId="4.[156,768,198,2077]" pageId="4" pageNumber="11">
|
||
<emphasis id="1ACD5873FF8FFFB4FF52D6017CA70510" bold="true" box="[181,397,1792,1816]" pageId="4" pageNumber="11">
|
||
Notes on
|
||
<typeStatus id="F7023AC3FF8FFFB4FEF8D6017CA20510" box="[287,392,1792,1816]" pageId="4" pageNumber="11" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<materialsCitation id="98D18E3CFF8FFFB4FE70D6017DC50572" ID-GBIF-Occurrence="3097624303" collectionCode="R, NSMT" pageId="4" pageNumber="11" specimenCount="1" typeStatus="paratype">
|
||
The body dimensions in the
|
||
<typeStatus id="F7023AC3FF8FFFB4FD2ED6007DFB0531" pageId="4" pageNumber="11" type="paratype">paratypes</typeStatus>
|
||
are:
|
||
<collectionCode id="4EA81CA4FF8FFFB4FEF7D6207C0A0530" box="[272,288,1825,1848]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="11">R</collectionCode>
|
||
=10.0 mm, r=5.0 mm, and
|
||
<collectionCode id="4EA81CA4FF8FFFB4FDA8D6207F4A0531" box="[591,608,1825,1849]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="11">R</collectionCode>
|
||
/r=2.0 (
|
||
<collectionCode id="4EA81CA4FF8FFFB4FD5FD6207FD50531" box="[696,767,1825,1849]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="4" pageNumber="11">NSMT</collectionCode>
|
||
E-8266);
|
||
<collectionCode id="4EA81CA4FF8FFFB4FEE4D6437C390551" box="[259,275,1858,1881]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="11">R</collectionCode>
|
||
=
|
||
<quantity id="EF412984FF8FFFB4FECFD6407CAB0551" box="[296,385,1857,1881]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="4" pageNumber="11" unit="mm" value="20.4">20.4 mm</quantity>
|
||
, r=
|
||
<quantity id="EF412984FF8FFFB4FE53D6407F2A0551" box="[436,512,1857,1881]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="4" pageNumber="11" unit="mm" value="9.7">9.7 mm</quantity>
|
||
, and
|
||
<collectionCode id="4EA81CA4FF8FFFB4FDABD6407F770551" box="[588,605,1857,1881]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="11">R</collectionCode>
|
||
/r=2.1 (
|
||
<collectionCode id="4EA81CA4FF8FFFB4FD5FD6407FD50551" box="[696,767,1857,1881]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="4" pageNumber="11">NSMT</collectionCode>
|
||
E-8267)
|
||
</materialsCitation>
|
||
. The other characters are consistent with the description of the
|
||
<typeStatus id="F7023AC3FF8FFFB4FEDBD6837CB20592" box="[316,408,1922,1946]" pageId="4" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="28068461FF8FFFB4FF52D6A27C1705F4" blockId="4.[156,768,198,2077]" pageId="4" pageNumber="11">
|
||
<emphasis id="1ACD5873FF8FFFB4FF52D6A27C7005B3" bold="true" box="[181,346,1955,1979]" pageId="4" pageNumber="11">DNA sequence.</emphasis>
|
||
A partial sequence of COI (655 bp) was obtained from NSMT E-8265 and deposited in DDBJ (Acc. No.
|
||
<accessionNumber id="37EA1982FF8FFFB4FF21D6E57C1805F4" box="[198,306,2020,2044]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC427072" pageId="4" pageNumber="11">LC427072</accessionNumber>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="60A3D7EAFF8FFFB4FF52D90479790623" pageId="4" pageNumber="11" type="discussion">
|
||
<paragraph id="28068461FF8FFFB4FF52D90479A300AD" blockId="4.[156,768,198,2077]" lastBlockId="4.[820,1432,198,1425]" pageId="4" pageNumber="11">
|
||
<emphasis id="1ACD5873FF8FFFB4FF52D9047C330A15" bold="true" box="[181,281,2053,2077]" pageId="4" pageNumber="11">Remarks.</emphasis>
|
||
<bibRefCitation id="4C28F990FF8FFFB4FEF8D9057CBD0A15" author="Mah, C. L." box="[287,407,2052,2077]" pageId="4" pageNumber="11" pagination="1 - 72" refId="ref11582" refString="Mah, C. L. 2017. Overview of the Ferdina - like Goniasteridae (Echinodermata: Asteroidea) including a new subfamily, three new genera and fourteen new species. Zootaxa 4271: 1 - 72." type="journal article" year="2017">Mah (2017)</bibRefCitation>
|
||
established
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FDF5D9057FB90A14" authorityName="Mah" authorityYear="2017" box="[530,659,2052,2076]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="1ACD5873FF8FFFB4FDF5D9057FB90A14" box="[530,659,2052,2076]" italics="true" pageId="4" pageNumber="11">Bathyferdina</emphasis>
|
||
</taxonomicName>
|
||
with a description of
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FC53D1C6793202D6" authorityName="Mah" authorityYear="2017" box="[948,1048,199,222]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="species" species="aireyae">
|
||
<emphasis id="1ACD5873FF8FFFB4FC53D1C6793202D6" box="[948,1048,199,222]" italics="true" pageId="4" pageNumber="11">B. aireyae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FBC0D1C779D402D6" authority="Arai & Fujita, 2021" authorityName="Arai & Fujita" authorityYear="2021" box="[1063,1278,198,222]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="species" species="caelator" status="sp. nov.">
|
||
<emphasis id="1ACD5873FF8FFFB4FBC0D1C779D402D6" box="[1063,1278,198,222]" italics="true" pageId="4" pageNumber="11">Bathyferdina caelator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01FEE508FF8FFFB4FAE0D1C6787C02D6" box="[1287,1366,199,222]" pageId="4" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is devoid of any subambulacral spines or enlarged granules while it shows small R/r, a uniform granular skin covering abactinal and actinal surface, marginal plates continuous in size, large quadrate bare area on marginal plates, homogenous abactinal plates completely covered with skin, and glassy bosses on abactinal plates. These characters support placement of the present species in
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FB6ED0AB782003CA" authorityName="Mah" authorityYear="2017" box="[1161,1290,426,450]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="1ACD5873FF8FFFB4FB6ED0AB782003CA" box="[1161,1290,426,450]" italics="true" pageId="4" pageNumber="11">Bathyferdina</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="4C28F990FF8FFFB4FAFCD0AB78BD03CA" author="Mah, C. L." box="[1307,1431,426,450]" pageId="4" pageNumber="11" pagination="1 - 72" refId="ref11582" refString="Mah, C. L. 2017. Overview of the Ferdina - like Goniasteridae (Echinodermata: Asteroidea) including a new subfamily, three new genera and fourteen new species. Zootaxa 4271: 1 - 72." type="journal article" year="2017">Mah (2017)</bibRefCitation>
|
||
mentioned that
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FC3BD0CB797703EA" authorityName="Mah" authorityYear="2017" box="[988,1117,458,482]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="1ACD5873FF8FFFB4FC3BD0CB797703EA" box="[988,1117,458,482]" italics="true" pageId="4" pageNumber="11">Bathyferdina</emphasis>
|
||
</taxonomicName>
|
||
has no glassy bosses on marginal plates in his key of
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FBDBD0EA7983000B" authorityName="Mah" authorityYear="2017" box="[1084,1193,491,515]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="subFamily" subFamily="Ferdininae">Ferdininae</taxonomicName>
|
||
but he also stated that it has glassy bosses on both abactinal and marginal plates. According to Mah (personal communication),
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FAFFD32C7851004C" authorityName="Mah" authorityYear="2017" box="[1304,1403,557,580]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="species" species="aireyae">
|
||
<emphasis id="1ACD5873FF8FFFB4FAFFD32C7851004C" box="[1304,1403,557,580]" italics="true" pageId="4" pageNumber="11">B. aireyae</emphasis>
|
||
</taxonomicName>
|
||
in fact has glassy bosses on the abactinal and marginal surfaces but not on the actinal surface. We have emended the diagnosis of
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FC6FD38C792300AD" authorityName="Mah" authorityYear="2017" box="[904,1033,653,677]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="1ACD5873FF8FFFB4FC6FD38C792300AD" box="[904,1033,653,677]" italics="true" pageId="4" pageNumber="11">Bathyferdina</emphasis>
|
||
</taxonomicName>
|
||
accordingly.
|
||
</paragraph>
|
||
<paragraph id="28068461FF8FFFB4FCA9D3AF79790623" blockId="4.[820,1432,198,1425]" pageId="4" pageNumber="11">
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FCA9D3AF790100CE" authorityName="Arai & Fujita" authorityYear="2021" box="[846,1067,686,710]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="species" species="caelator">
|
||
<emphasis id="1ACD5873FF8FFFB4FCA9D3AF790100CE" box="[846,1067,686,710]" italics="true" pageId="4" pageNumber="11">Bathyferdina caelator</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from the single congeneric species,
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FBE1D3CE794600EE" authorityName="Mah" authorityYear="2017" box="[1030,1132,719,742]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="species" species="aireyae">
|
||
<emphasis id="1ACD5873FF8FFFB4FBE1D3CE794600EE" box="[1030,1132,719,742]" italics="true" pageId="4" pageNumber="11">B. aireyae</emphasis>
|
||
</taxonomicName>
|
||
, in having glassy bosses on not only abactinal but actinal and adambulacral plates while lacking them on marginal plates.
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FB6ED21178750120" authorityName="Arai & Fujita" authorityYear="2021" box="[1161,1375,784,808]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="species" species="caelator">
|
||
<emphasis id="1ACD5873FF8FFFB4FB6ED21178750120" box="[1161,1375,784,808]" italics="true" pageId="4" pageNumber="11">Bathyferdina caelator</emphasis>
|
||
</taxonomicName>
|
||
lacks granules or tubercles on the inferomarginal plates, while they are occasionally present in
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FB64D25079CD0160" authorityName="Mah" authorityYear="2017" box="[1155,1255,849,872]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="species" species="aireyae">
|
||
<emphasis id="1ACD5873FF8FFFB4FB64D25079CD0160" box="[1155,1255,849,872]" italics="true" pageId="4" pageNumber="11">B. aireyae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="4C28F990FF8FFFB4FB12D25078440161" author="Mah, C. L." box="[1269,1390,849,873]" pageId="4" pageNumber="11" pagination="1 - 72" refId="ref11582" refString="Mah, C. L. 2017. Overview of the Ferdina - like Goniasteridae (Echinodermata: Asteroidea) including a new subfamily, three new genera and fourteen new species. Zootaxa 4271: 1 - 72." type="journal article" year="2017">Mah (2017)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FC62D2737ECD0181" authorityName="Mah" authorityYear="2017" box="[901,999,882,905]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="species" species="aireyae">
|
||
<emphasis id="1ACD5873FF8FFFB4FC62D2737ECD0181" box="[901,999,882,905]" italics="true" pageId="4" pageNumber="11">
|
||
B.
|
||
<materialsCitation id="98D18E3CFF8FFFB4FC47D2737ECD0181" ID-GBIF-Occurrence="3097624306" box="[928,999,882,905]" collectionCode="CASIZ" pageId="4" pageNumber="11" specimenCode="CASIZ 219696" specimenCount="1" typeStatus="paratype">aireyae</materialsCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
as lacking any pedicellariae but one of the
|
||
<typeStatus id="F7023AC3FF8FFFB4FCD3D2937EBF01A2" box="[820,917,914,938]" pageId="4" pageNumber="11" type="paratype">paratypes</typeStatus>
|
||
reexamined in this study (
|
||
<specimenCode id="781F2C1AFF8FFFB4FB5CD293787301A2" box="[1211,1369,914,938]" collectionCode="CASIZ" pageId="4" pageNumber="11">CASIZ 219696</specimenCode>
|
||
) possesses a large number of pedicellariae on marginal and actinal plates. Although these pedicellariae varied in number of valves from one to five, the shape of the valves closely resembled those in
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FC02D51479670624" authorityName="Arai & Fujita" authorityYear="2021" box="[997,1101,1044,1068]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="species" species="caelator">
|
||
<emphasis id="1ACD5873FF8FFFB4FC02D51479670624" box="[997,1101,1044,1068]" italics="true" pageId="4" pageNumber="11">B. caelator</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="60A3D7EAFF8FFFB4FCA9D5357E510665" pageId="4" pageNumber="11" type="distribution">
|
||
<paragraph id="28068461FF8FFFB4FCA9D5357E510665" blockId="4.[820,1432,198,1425]" pageId="4" pageNumber="11">
|
||
<emphasis id="1ACD5873FF8FFFB4FCA9D5357EF30644" bold="true" box="[846,985,1076,1100]" pageId="4" pageNumber="11">Distribution.</emphasis>
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FC0DD53579980644" authority="Islands" authorityName="Islands" box="[1002,1202,1076,1100]" class="Insecta" family="Cerambycidae" genus="Ogasawara" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="11" phylum="Arthropoda" rank="genus">Ogasawara Islands</taxonomicName>
|
||
,
|
||
<quantity id="EF412984FF8FFFB4FB20D53578730644" box="[1223,1369,1076,1100]" metricMagnitude="2" metricUnit="m" metricValue="1.4035" metricValueMax="1.452" metricValueMin="1.355" pageId="4" pageNumber="11" unit="m" value="140.35" valueMax="145.2" valueMin="135.5">135.5–145.2 m</quantity>
|
||
(this study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="60A3D7EAFF8FFFB4FCA9D5747EF30799" pageId="4" pageNumber="11" type="etymology">
|
||
<paragraph id="28068461FF8FFFB4FCA9D574791A06E7" blockId="4.[820,1432,198,1425]" pageId="4" pageNumber="11">
|
||
<emphasis id="1ACD5873FF8FFFB4FCA9D5747EEF0685" bold="true" box="[846,965,1141,1165]" pageId="4" pageNumber="11">Etymology.</emphasis>
|
||
The specific epithet,
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FB45D57479DA0685" authorityName="Arai & Fujita" authorityYear="2021" box="[1186,1264,1141,1165]" class="Asteroidea" family="Goniasteridae" genus="Bathyferdina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="species" species="caelator">
|
||
<emphasis id="1ACD5873FF8FFFB4FB45D57479DA0685" box="[1186,1264,1141,1165]" italics="true" pageId="4" pageNumber="11">caelator</emphasis>
|
||
</taxonomicName>
|
||
, is a Latin masculine noun in apposition meaning sculptor, and alluding to the complex patterns of glassy bosses on abactinal, actinal and adambulacral plates.
|
||
</paragraph>
|
||
<paragraph id="28068461FF8FFFB4FCA9D5F97EF30799" blockId="4.[820,1432,198,1425]" pageId="4" pageNumber="11">
|
||
<emphasis id="1ACD5873FF8FFFB4FCA9D5F97ED90718" bold="true" box="[846,1011,1272,1296]" pageId="4" pageNumber="11">Japanese name.</emphasis>
|
||
<emphasis id="1ACD5873FF8FFFB4FC1AD5F979050707" box="[1021,1071,1272,1295]" italics="true" pageId="4" pageNumber="11">Koyo</emphasis>
|
||
is taken from R/V
|
||
<emphasis id="1ACD5873FF8FFFB4FAE3D5F9781C0707" box="[1284,1334,1272,1295]" italics="true" pageId="4" pageNumber="11">Koyo</emphasis>
|
||
, the vessel which collected the
|
||
<typeStatus id="F7023AC3FF8FFFB4FBC1D41979AB0738" box="[1062,1153,1304,1328]" pageId="4" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
, and
|
||
<emphasis id="1ACD5873FF8FFFB4FB5ED419787A0738" box="[1209,1360,1304,1328]" italics="true" pageId="4" pageNumber="11">akamon-hitode</emphasis>
|
||
comes from the Japanese name of
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FBADD43878B90759" authority="(Gray, 1840)" baseAuthorityName="Gray" baseAuthorityYear="1840" box="[1098,1427,1337,1361]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="species" species="cumingi">
|
||
<emphasis id="1ACD5873FF8FFFB4FBADD43878270758" box="[1098,1293,1337,1361]" italics="true" pageId="4" pageNumber="11">Neoferdina cumingi</emphasis>
|
||
(Gray, 1840)
|
||
</taxonomicName>
|
||
, another species of
|
||
<taxonomicName id="EFB9FFE2FF8FFFB4FC19D45879410779" authorityName="Mah" authorityYear="2017" box="[1022,1131,1369,1393]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="11" phylum="Echinodermata" rank="subFamily" subFamily="Ferdininae">Ferdininae</taxonomicName>
|
||
which occurs commonly in Japanese waters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |