treatments-xml/data/D1/BB/27/D1BB273919275908918E5C3033B8423B.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="D1BB273919275908918E5C3033B8423B" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila huoyani Jochum, Slapnik &amp; Pall-Gergely 2014" docType="treatment" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1147</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>177</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
<mods:identifier type="Pensoft-UUID">FF2B6B317B505F9EA0E1000BDCD16CE7</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:D1BB273919275908918E5C3033B8423B" httpUri="http://treatment.plazi.org/id/D1BB273919275908918E5C3033B8423B" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="5V2MZ" LSID="D1BB2739-1927-5908-918E-5C3033B8423B" authority="Jochum, Slapnik &amp; Pall-Gergely, 2014" authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="huoyani">
Angustopila huoyani Jochum, Slapnik &amp; 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
, 2014
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 66" captionStartId="F66" captionText="Figure 66. Angustopila huoyani Jochum, Slapnik &amp; Pall-Gergely, 2014 (holotype, MCSMNH 50312 / 1). Apertural (A), apical (B), lateral (C) and ventral (D) sides of the shell; sculpture on the protoconch (E), aperture (F); ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure66" httpUri="https://binary.pensoft.net/fig/808875" pageId="0" pageNumber="1">Figs 66</figureCitation>
<figureCitation captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Conchological variability (upper plate) and Shell diameter (x axis, in mm) vs. shell height (y axis, in mm) (lower diagram) of Angustopila huoyani Jochum, Slapnik &amp; Pall-Gergely, 2014. Group 1 (A, B), Group 2 (C, D), Group 3 (E, F), Group 4 (G, H). Group 1 (dot), Group 2 (circle), Group 3 (empty square), Group 4 (filled square)." figureDoi="10.3897/zookeys.1147.93824.figure67" httpUri="https://binary.pensoft.net/fig/808876" pageId="0" pageNumber="1">, 67</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="huoyani">Angustopila huoyani</taxonomicName>
Jochum, Slapnik &amp; 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
in 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.410.7488" author="Jochum, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="23 - 40" refId="B30" refString="Jochum, A, Slapnik, R, Kampschulte, M, Martels, G, Heneka, M, Pall-Gergely, B, 2014. A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae). ZooKeys 410: 23 - 40, DOI: https://doi.org/10.3897/zookeys.410.7488" title="A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae)." url="https://doi.org/10.3897/zookeys.410.7488" volume="410" year="2014">Jochum et al. 2014</bibRefCitation>
: 27, figs 4, 5, video 1.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="huoyani">Angustopila huoyani</taxonomicName>
- 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.523.6114" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="31 - 64" refId="B39" refString="Pall-Gergely, B, Hunyadi, A, Jochum, A, Asami, T, 2015a. Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra). ZooKeys 523: 31 - 64, DOI: https://doi.org/10.3897/zookeys.523.6114" title="Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra)." url="https://doi.org/10.3897/zookeys.523.6114" volume="523" year="2015 a">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2015a
</bibRefCitation>
: 39, fig. 3.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
&quot;China, Hunan, Longshan (龙山县), Huoyan (火焰), Feihu Dong (飞虎洞), (
<geoCoordinate degrees="29" direction="north" minutes="12.53" orientation="latitude" precision="9" value="29.208834">29°12.53'N</geoCoordinate>
, 
<geoCoordinate degrees="109" direction="east" minutes="18.37" orientation="longitude" precision="9" value="109.30617">109°18.37'E</geoCoordinate>
, 550 m alt.)&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: China • 1 shell; Hunan, Longshan, Huoyan, Feihu Dong (&quot;Cave of the Wild Tiger&quot;), Kitajska; 13 Apr. 1997; Verovnik leg.; MCSMNH 50312/1.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: China • 1 shell; same data as for holotype; MCSMNH 50312/2-3 • 3 shells; same data as for holotype; MCSMNH 50312/4-6 • 2 shells; same data as for holotype; MCSMNH 50312/7-9.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<collectingCountry name="China">China</collectingCountry>
• 65 complete shells; same data as for holotype; MCSMNH 50312 (separated into 4 vials based on morphological groups, see remarks) • 48 broken shells (in 2 vials); same data as for 
<typeStatus>holotype</typeStatus>
; MCSMNH 50312.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A medium-sized to large, conical-globular 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with a very narrow umbilicus, a parietal tooth, and an upper palatal tooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
The morphologically closest congener with two apertural denticles is 
<taxonomicName lsidName="A. tamlod" pageId="0" pageNumber="1" rank="species" species="tamlod">
<emphasis italics="true" pageId="0" pageNumber="1">A. tamlod</emphasis>
</taxonomicName>
known from Thailand; however, 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="huoyani">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila huoyani</emphasis>
</taxonomicName>
is tightly coiled, has a more conical shell, bears more pronounced apertural dentition, and has stronger and denser spiral striation. The shell of 
<taxonomicName lsidName="A. dominikae" pageId="0" pageNumber="1" rank="species" species="dominikae">
<emphasis italics="true" pageId="0" pageNumber="1">A. dominikae</emphasis>
</taxonomicName>
is more compact and is 2/3 the size of 
<taxonomicName lsidName="A. huoyani" pageId="0" pageNumber="1" rank="species" species="huoyani">
<emphasis italics="true" pageId="0" pageNumber="1">A. huoyani</emphasis>
</taxonomicName>
. Though also bearing two denticles, 
<taxonomicName lsidName="A. uvula" pageId="0" pageNumber="1" rank="species" species="uvula">
<emphasis italics="true" pageId="0" pageNumber="1">A. uvula</emphasis>
</taxonomicName>
sp. nov. is comparatively loosely coiled, has larger whorls, a bulbous protoconch and a slightly downwards directed tuba and bears a very prominent parietal denticle opposed by a weak palatal one. 
<taxonomicName authorityName="Pall-Gergely &amp; Jochum" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila bidentata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bidentata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila bidentata</emphasis>
</taxonomicName>
sp. nov. has a narrower umbilicus and more oblique aperture. 
<taxonomicName authorityName="Pall-Gergely &amp; Dumrongrojwattana" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila erawanica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="erawanica">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila erawanica</emphasis>
</taxonomicName>
sp. nov. has a more horizontally oriented aperture that is impressed at the parietal lamella.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.96-1.31; D = 0.82-1.1, H/D*100 = 109.1-143.6 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 67), RUD = 14.3-18.8 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 6).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known from two caves, one located in Guangxi, and the other in Hunan, China (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure19" httpUri="https://binary.pensoft.net/fig/808828" pageId="0" pageNumber="1">19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The original description of the type material describes the holotype and eight paratypes. However, results in 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.410.7488" author="Jochum, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="23 - 40" refId="B30" refString="Jochum, A, Slapnik, R, Kampschulte, M, Martels, G, Heneka, M, Pall-Gergely, B, 2014. A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae). ZooKeys 410: 23 - 40, DOI: https://doi.org/10.3897/zookeys.410.7488" title="A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae)." url="https://doi.org/10.3897/zookeys.410.7488" volume="410" year="2014">Jochum et al. (2014</bibRefCitation>
: table 2) were based on 30 specimens. For this revision, and because we intended to re-investigate this species and compare it with other congeners, we borrowed the type material of 
<taxonomicName lsidName="A. huoyani" pageId="0" pageNumber="1" rank="species" species="huoyani">
<emphasis italics="true" pageId="0" pageNumber="1">A. huoyani</emphasis>
</taxonomicName>
from its home museum in Ljubljana. The examination revealed that in addition to the holotype and the paratypes, there were 65 shells (previously unknown to 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.410.7488" author="Jochum, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="23 - 40" refId="B30" refString="Jochum, A, Slapnik, R, Kampschulte, M, Martels, G, Heneka, M, Pall-Gergely, B, 2014. A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae). ZooKeys 410: 23 - 40, DOI: https://doi.org/10.3897/zookeys.410.7488" title="A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae)." url="https://doi.org/10.3897/zookeys.410.7488" volume="410" year="2014">Jochum et al. 2014</bibRefCitation>
) in perfect condition and several others that were somewhat damaged. The 65 shells showed extraordinary variation in terms of shell size and shape, while the positions of the apertural teeth and the sculpture did not vary. At first sight, the shells could be separated into four more or less clearly distinguishable groups as follows: (1) shell large, rounded, aperture ovoid, whorls: 4.75-5.5 (mean: 5.15); (2) shell large, with many whorls, aperture depressed (narrow), whorls: 5.5-6.5 (mean: 5.89) (3) shell very slender, aperture ovoid, whorls: 5-6 (mean: 5.47); and (4) shell small, rounded, aperture ovoid, whorls: 4.75-5.5 (mean: 5.26). After we measured all shells, it became evident that none of these groups are distinct based on shell measurements alone (see Fig. 
<figureCitation captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Conchological variability (upper plate) and Shell diameter (x axis, in mm) vs. shell height (y axis, in mm) (lower diagram) of Angustopila huoyani Jochum, Slapnik &amp; Pall-Gergely, 2014. Group 1 (A, B), Group 2 (C, D), Group 3 (E, F), Group 4 (G, H). Group 1 (dot), Group 2 (circle), Group 3 (empty square), Group 4 (filled square)." figureDoi="10.3897/zookeys.1147.93824.figure67" httpUri="https://binary.pensoft.net/fig/808876" pageId="0" pageNumber="1">67</figureCitation>
). Although the grouping of the shells into morphological groups was possible using a stereo microscope, these do not likely belong to a biologically distinct species. It is, however, possible that the soil sample containing all 
<taxonomicName lsidName="A. huoyani" pageId="0" pageNumber="1" rank="species" species="huoyani">
<emphasis italics="true" pageId="0" pageNumber="1">A. huoyani</emphasis>
</taxonomicName>
shells originated from multiple subpopulations living under different environmental conditions. Thus, the extreme variability we observed is explained by phenotypic plasticity.
</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure66" httpUri="https://binary.pensoft.net/fig/808875" pageId="0" pageNumber="1" start="Figure 66" startId="F66">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 66.</emphasis>
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="huoyani">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila huoyani</emphasis>
</taxonomicName>
Jochum, Slapnik &amp; 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
, 2014 (holotype, MCSMNH 50312/1). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) sides of the shell; sculpture on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
), aperture (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
); ventral (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure67" httpUri="https://binary.pensoft.net/fig/808876" pageId="0" pageNumber="1" start="Figure 67" startId="F67">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 67.</emphasis>
Conchological variability (upper plate) and Shell diameter (x axis, in mm) vs. shell height (y axis, in mm) (lower diagram) of 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="huoyani">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila huoyani</emphasis>
</taxonomicName>
Jochum, Slapnik &amp; 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
, 2014. Group 1 (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
), Group 2 (
<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
), Group 3 (
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
), Group 4 (
<emphasis bold="true" pageId="0" pageNumber="1">G, H</emphasis>
). Group 1 (dot), Group 2 (circle), Group 3 (empty square), Group 4 (filled square).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">Measurements of the holotype (H: 1.14 mm, D: 0.94 mm) differ from those published in the original description (H: 1.09 mm, D: 0.87 mm).</paragraph>
</subSubSection>
</treatment>
</document>