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<document id="E46EF962E1055DBC145638126E12A018" ID-DOI="10.24199/j.mmv.2012.69.03" ID-ISSN="1447-2554" ID-Zenodo-Dep="10666102" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708005403175" checkinUser="felipe" docAuthor="Cartwright, David I." docDate="2012" docId="AF02CC7797470B4AE0BDF99E97873374" docLanguage="en" docName="MemMusVictoria.69.245-258.pdf" docOrigin="Memoirs of Museum Victoria 69" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-69-2012/pages-245-258/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Hydrobiosella tenuitas Cartwright 2012, sp. nov." docType="treatment" docVersion="3" lastPageNumber="256" masterDocId="533BB40F974C0B41E02EFFD59348377D" masterDocTitle="Two further species groups and new species among Australian Hydrobiosella Tillyard: new species from south-eastern Australia (Trichoptera: Philopotamidae)" masterLastPageNumber="258" masterPageNumber="245" pageNumber="256" updateTime="1708006338923" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6F79705D77F438B8CBD61A5199C15554">Two further species groups and new species among Australian Hydrobiosella Tillyard: new species from south-eastern Australia (Trichoptera: Philopotamidae)</mods:title>
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<treatment id="AF02CC7797470B4AE0BDF99E97873374" ID-DOI="http://doi.org/10.5281/zenodo.10666180" ID-Zenodo-Dep="10666180" LSID="urn:lsid:plazi:treatment:AF02CC7797470B4AE0BDF99E97873374" httpUri="http://treatment.plazi.org/id/AF02CC7797470B4AE0BDF99E97873374" lastPageNumber="256" pageId="11" pageNumber="256">
<subSubSection id="6FB12EEA97470B4AE0BDF99E929B311C" box="[147,467,1611,1633]" pageId="11" pageNumber="256" type="nomenclature">
<paragraph id="27147D6197470B4AE0BDF99E929B311C" blockId="11.[147,467,1611,1633]" box="[147,467,1611,1633]" pageId="11" pageNumber="256">
<heading id="7C5CCA0D97470B4AE0BDF99E929B311C" box="[147,467,1611,1633]" fontSize="9" level="2" pageId="11" pageNumber="256" reason="5">
<taxonomicName id="E0AB06E297470B4AE0BDF99E92CA311D" authority="Cartwright, 2012" authorityName="Cartwright" authorityYear="2012" box="[147,386,1611,1633]" class="Insecta" family="Philopotamidae" genus="Hydrobiosella" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="256" phylum="Arthropoda" rank="species" species="tenuitas" status="sp. nov.">
<emphasis id="15DFA17397470B4AE0BDF99E92CA311D" bold="true" box="[147,386,1611,1633]" italics="true" pageId="11" pageNumber="256">Hydrobiosella tenuitas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EEC1C0897470B4AE1A6F999929B311C" box="[392,467,1612,1633]" pageId="11" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6FB12EEA97470B4AE0BDF9A297873374" pageId="11" pageNumber="256" type="description">
<paragraph id="27147D6197470B4AE0BDF9A2926B31F1" blockId="11.[147,291,1655,1677]" box="[147,291,1655,1677]" pageId="11" pageNumber="256">
<figureCitation id="BF9061E497470B4AE0BDF9A2926B31F1" box="[147,291,1655,1677]" captionStart="Figures 2432" captionStartId="6.[140,208,1830,1849]" captionTargetBox="[148,1436,232,1683]" captionTargetId="figure-22@6.[140,1440,227,1688]" captionTargetPageId="6" captionText="Figures 2432. Hydrobiosella spp.; male genitalia in dorsal, lateral and part ventral views; 2426, Hydrobiosella incisura sp. nov.; 24, dorsal; 25, lateral; 26, ventral, mesodistal margin of segment IX; 2729, Hydrobiosella tenuitas sp. nov.; 27, dorsal; 28, lateral; 29, ventral, mesodistal margin of segment IX; 3032, Hydrobiosella exilatis sp. nov.; 30, dorsal; 31, lateral; 32, ventral, mesodistal margin of segment IX." figureDoi="http://doi.org/10.5281/zenodo.10666110" httpUri="https://zenodo.org/record/10666110/files/figure.png" pageId="11" pageNumber="256">Figures 2729</figureCitation>
</paragraph>
<paragraph id="27147D6197470B4AE097F97691943197" blockId="11.[147,779,1699,1771]" pageId="11" pageNumber="256">
<materialsCitation id="97C3773C97470B4AE097F97691943197" collectingDate="1981-11-01" collectingDateMax="1981-11-07" collectingDateMin="1981-11-01" collectionCode="NMV" elevation="1560" latitude="-17.266666" location="Bellenden Ker Range" longLatPrecision="1279" longitude="145.9" municipality="Male" pageId="11" pageNumber="256" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<typeStatus id="F810C3C397470B4AE097F976925831CB" box="[185,272,1699,1718]" pageId="11" pageNumber="256" type="holotype">Holotype</typeStatus>
.
<collectingMunicipality id="C770E71B97470B4AE108F976921D31CB" box="[294,341,1699,1718]" pageId="11" pageNumber="256">Male</collectingMunicipality>
(specimen PT-1038 figured),
<collectingRegion id="E56FB38397470B4AE2B7F976904E31CB" box="[665,774,1699,1718]" country="Australia" name="Queensland" pageId="11" pageNumber="256">Queensland</collectingRegion>
,
<location id="22742BBA97470B4AE0BDF968922231AD" LSID="urn:lsid:plazi:treatment:AF02CC7797470B4AE0BDF99E97873374:22742BBA97470B4AE0BDF968922231AD" box="[147,362,1725,1744]" latitude="-17.266666" longLatPrecision="1279" longitude="145.9" municipality="Male" name="Bellenden Ker Range" pageId="11" pageNumber="256" stateProvince="Queensland">Bellenden Ker Range</location>
, summit TV stn,
<quantity id="E053D08497470B4AE203F968912331AD" box="[557,619,1725,1744]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="11" pageNumber="256" unit="m" value="1560.0">
<elevation id="AC869A5297470B4AE203F968912331AD" box="[557,619,1725,1744]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="11" pageNumber="256" unit="m" value="1560.0">1560m</elevation>
</quantity>
(about
<geoCoordinate id="429F1BA697470B4AE2ECF968904E31AD" box="[706,774,1725,1744]" degrees="17" direction="south" minutes="16" orientation="latitude" pageId="11" pageNumber="256" precision="925" value="-17.266666">17°16'S</geoCoordinate>
,
<geoCoordinate id="429F1BA697470B4AE0BDF90293AE3197" box="[147,230,1751,1770]" degrees="145" direction="east" minutes="54" orientation="longitude" pageId="11" pageNumber="256" precision="925" value="145.9">145°54'E</geoCoordinate>
),
<date id="53155BA197470B4AE0DBF90292263197" box="[245,366,1751,1771]" pageId="11" pageNumber="256" value="1981-11-01" valueMax="1981-11-07" valueMin="1981-11-01">
<collectingDate id="4351A24997470B4AE0DBF90292263197" box="[245,366,1751,1771]" pageId="11" pageNumber="256" value="1981-11-01" valueMax="1981-11-07" valueMin="1981-11-01">1-7 Nov 1981</collectingDate>
</date>
, Earthwatch-Qld Mus. (
<collectionCode id="41BAE5A497470B4AE269F90291363197" box="[583,638,1751,1770]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="11" pageNumber="256" type="Museum">NMV</collectionCode>
, T-21479).
</materialsCitation>
</paragraph>
<paragraph id="27147D6197470B4AE0BDF8D791013012" blockId="11.[147,778,1794,1903]" pageId="11" pageNumber="256">
<emphasis id="15DFA17397470B4AE0BDF8D793B1306A" box="[147,249,1794,1815]" italics="true" pageId="11" pageNumber="256">Diagnosis</emphasis>
. Males of
<taxonomicName id="E0AB06E297470B4AE145F8D7911D306A" authorityName="Cartwright" authorityYear="2012" box="[363,597,1794,1815]" class="Insecta" family="Philopotamidae" genus="Hydrobiosella" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="256" phylum="Arthropoda" rank="species" species="tenuitas">
<emphasis id="15DFA17397470B4AE145F8D7911D306A" box="[363,597,1794,1815]" italics="true" pageId="11" pageNumber="256">Hydrobiosella tenuitas</emphasis>
</taxonomicName>
can be separated from those of other species in the group by the slender segment X, also slightly dilated apically in lateral view, and the long and slender harpago of the inferior appendages.
</paragraph>
<paragraph id="27147D6197470B4AE31FFF3397693665" blockId="11.[817,1448,229,280]" pageId="11" pageNumber="256">
<emphasis id="15DFA17397470B4AE31FFF3390E13786" box="[817,937,230,251]" italics="true" pageId="11" pageNumber="256">Description</emphasis>
. Wings similar to
<taxonomicName id="E0AB06E297470B4AE451FF3397843787" box="[1151,1228,229,251]" class="Insecta" family="Philopotamidae" genus="Hydrobiosella" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="256" phylum="Arthropoda" rank="species" species="letti">
<emphasis id="15DFA17397470B4AE451FF3397843787" box="[1151,1228,229,251]" italics="true" pageId="11" pageNumber="256">H. letti</emphasis>
</taxonomicName>
(
<figureCitation id="BF9061E497470B4AE4CEFF3096663786" box="[1248,1326,229,251]" captionStart="Figures 1723" captionStartId="5.[147,216,1830,1849]" captionTargetBox="[168,1423,230,1808]" captionTargetId="figure-6@5.[162,1432,227,1813]" captionTargetPageId="5" captionText="Figures 1723. Hydrobiosella spp.; 17, Hydrobiosella letti Korboot., wings. 1823, Hydrobiosella spp., male genitalia in dorsal, lateral and part ventral views; 1820, Hydrobiosella fibra sp. nov.; 18, dorsal; 19, lateral; 20, ventral, mesodistal margin of segment IX; 2123, Hydrobiosella bifurca sp. nov.; 21, dorsal; 22, lateral; 23, ventral, mesodistal margin of segment IX." figureDoi="http://doi.org/10.5281/zenodo.10666108" httpUri="https://zenodo.org/record/10666108/files/figure.png" pageId="11" pageNumber="256">Fig. 17</figureCitation>
), length of forewing: male
<quantity id="E053D08497470B4AE3E1FED797553665" box="[975,1053,258,280]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="11" pageNumber="256" unit="mm" value="7.6">7.6 mm</quantity>
.
</paragraph>
<paragraph id="27147D6197470B4AE31FFEFB9777346F" blockId="11.[817,1448,302,786]" pageId="11" pageNumber="256">
<emphasis id="15DFA17397470B4AE31FFEFB902D363E" box="[817,869,302,323]" italics="true" pageId="11" pageNumber="256">Male</emphasis>
. Segment IX in lateral view, length about 1.2 times width, sub-rectangular, not projecting and tapered basally, projecting slightly distally (
<figureCitation id="BF9061E497470B4AE470FEBD97E63603" box="[1118,1198,360,382]" captionStart="Figures 2432" captionStartId="6.[140,208,1830,1849]" captionTargetBox="[148,1436,232,1683]" captionTargetId="figure-22@6.[140,1440,227,1688]" captionTargetPageId="6" captionText="Figures 2432. Hydrobiosella spp.; male genitalia in dorsal, lateral and part ventral views; 2426, Hydrobiosella incisura sp. nov.; 24, dorsal; 25, lateral; 26, ventral, mesodistal margin of segment IX; 2729, Hydrobiosella tenuitas sp. nov.; 27, dorsal; 28, lateral; 29, ventral, mesodistal margin of segment IX; 3032, Hydrobiosella exilatis sp. nov.; 30, dorsal; 31, lateral; 32, ventral, mesodistal margin of segment IX." figureDoi="http://doi.org/10.5281/zenodo.10666110" httpUri="https://zenodo.org/record/10666110/files/figure.png" pageId="11" pageNumber="256">Fig. 28</figureCitation>
), with a shallow, wide V-shaped notch medially on distal margin (
<figureCitation id="BF9061E497470B4AE4DCFE50967536E6" box="[1266,1341,389,411]" captionStart="Figures 2432" captionStartId="6.[140,208,1830,1849]" captionTargetBox="[148,1436,232,1683]" captionTargetId="figure-22@6.[140,1440,227,1688]" captionTargetPageId="6" captionText="Figures 2432. Hydrobiosella spp.; male genitalia in dorsal, lateral and part ventral views; 2426, Hydrobiosella incisura sp. nov.; 24, dorsal; 25, lateral; 26, ventral, mesodistal margin of segment IX; 2729, Hydrobiosella tenuitas sp. nov.; 27, dorsal; 28, lateral; 29, ventral, mesodistal margin of segment IX; 3032, Hydrobiosella exilatis sp. nov.; 30, dorsal; 31, lateral; 32, ventral, mesodistal margin of segment IX." figureDoi="http://doi.org/10.5281/zenodo.10666110" httpUri="https://zenodo.org/record/10666110/files/figure.png" pageId="11" pageNumber="256">Fig. 29</figureCitation>
). Segment X slender, almost parallel sided in basal two-thirds, length about 2.8-2.9 times width, narrowed in apical third (
<figureCitation id="BF9061E497470B4AE57BFE6B9004368F" captionStart="Figures 2432" captionStartId="6.[140,208,1830,1849]" captionTargetBox="[148,1436,232,1683]" captionTargetId="figure-22@6.[140,1440,227,1688]" captionTargetPageId="6" captionText="Figures 2432. Hydrobiosella spp.; male genitalia in dorsal, lateral and part ventral views; 2426, Hydrobiosella incisura sp. nov.; 24, dorsal; 25, lateral; 26, ventral, mesodistal margin of segment IX; 2729, Hydrobiosella tenuitas sp. nov.; 27, dorsal; 28, lateral; 29, ventral, mesodistal margin of segment IX; 3032, Hydrobiosella exilatis sp. nov.; 30, dorsal; 31, lateral; 32, ventral, mesodistal margin of segment IX." figureDoi="http://doi.org/10.5281/zenodo.10666110" httpUri="https://zenodo.org/record/10666110/files/figure.png" pageId="11" pageNumber="256">Figs 27, 28</figureCitation>
), in dorsal view, narrow tongue-shaped, tapered slightly in distal third, narrowly rounded apex (
<figureCitation id="BF9061E497470B4AE480FE2D97B23573" box="[1198,1274,504,526]" captionStart="Figures 2432" captionStartId="6.[140,208,1830,1849]" captionTargetBox="[148,1436,232,1683]" captionTargetId="figure-22@6.[140,1440,227,1688]" captionTargetPageId="6" captionText="Figures 2432. Hydrobiosella spp.; male genitalia in dorsal, lateral and part ventral views; 2426, Hydrobiosella incisura sp. nov.; 24, dorsal; 25, lateral; 26, ventral, mesodistal margin of segment IX; 2729, Hydrobiosella tenuitas sp. nov.; 27, dorsal; 28, lateral; 29, ventral, mesodistal margin of segment IX; 3032, Hydrobiosella exilatis sp. nov.; 30, dorsal; 31, lateral; 32, ventral, mesodistal margin of segment IX." figureDoi="http://doi.org/10.5281/zenodo.10666110" httpUri="https://zenodo.org/record/10666110/files/figure.png" pageId="11" pageNumber="256">Fig. 27</figureCitation>
); in lateral view, narrowed at about apical third, dilated slightly distally with bulbous apex (
<figureCitation id="BF9061E497470B4AE3EAFDE797453535" box="[964,1037,562,584]" captionStart="Figures 2432" captionStartId="6.[140,208,1830,1849]" captionTargetBox="[148,1436,232,1683]" captionTargetId="figure-22@6.[140,1440,227,1688]" captionTargetPageId="6" captionText="Figures 2432. Hydrobiosella spp.; male genitalia in dorsal, lateral and part ventral views; 2426, Hydrobiosella incisura sp. nov.; 24, dorsal; 25, lateral; 26, ventral, mesodistal margin of segment IX; 2729, Hydrobiosella tenuitas sp. nov.; 27, dorsal; 28, lateral; 29, ventral, mesodistal margin of segment IX; 3032, Hydrobiosella exilatis sp. nov.; 30, dorsal; 31, lateral; 32, ventral, mesodistal margin of segment IX." figureDoi="http://doi.org/10.5281/zenodo.10666110" httpUri="https://zenodo.org/record/10666110/files/figure.png" pageId="11" pageNumber="256">Fig. 28</figureCitation>
). Phallus generally tube-like; with a pair of slightly curved, slender lateral parameres arising from the phallus near middle (
<figureCitation id="BF9061E497470B4AE433FDB997D435FF" box="[1053,1180,620,642]" captionStart="Figures 2432" captionStartId="6.[140,208,1830,1849]" captionTargetBox="[148,1436,232,1683]" captionTargetId="figure-22@6.[140,1440,227,1688]" captionTargetPageId="6" captionText="Figures 2432. Hydrobiosella spp.; male genitalia in dorsal, lateral and part ventral views; 2426, Hydrobiosella incisura sp. nov.; 24, dorsal; 25, lateral; 26, ventral, mesodistal margin of segment IX; 2729, Hydrobiosella tenuitas sp. nov.; 27, dorsal; 28, lateral; 29, ventral, mesodistal margin of segment IX; 3032, Hydrobiosella exilatis sp. nov.; 30, dorsal; 31, lateral; 32, ventral, mesodistal margin of segment IX." figureDoi="http://doi.org/10.5281/zenodo.10666110" httpUri="https://zenodo.org/record/10666110/files/figure.png" pageId="11" pageNumber="256">Figs 27, 28</figureCitation>
). Inferior appendages in lateral view, with basal segment sub-trapezoidal, length about 1.4 times maximum width; harpago longer, length about 1.6 times length basal segment, more slender, length about 3.8-3.9 times maximum width, tapered slightly distally, broadly rounded apically (
<figureCitation id="BF9061E497470B4AE3C4FD29977B346F" box="[1002,1075,764,786]" captionStart="Figures 2432" captionStartId="6.[140,208,1830,1849]" captionTargetBox="[148,1436,232,1683]" captionTargetId="figure-22@6.[140,1440,227,1688]" captionTargetPageId="6" captionText="Figures 2432. Hydrobiosella spp.; male genitalia in dorsal, lateral and part ventral views; 2426, Hydrobiosella incisura sp. nov.; 24, dorsal; 25, lateral; 26, ventral, mesodistal margin of segment IX; 2729, Hydrobiosella tenuitas sp. nov.; 27, dorsal; 28, lateral; 29, ventral, mesodistal margin of segment IX; 3032, Hydrobiosella exilatis sp. nov.; 30, dorsal; 31, lateral; 32, ventral, mesodistal margin of segment IX." figureDoi="http://doi.org/10.5281/zenodo.10666110" httpUri="https://zenodo.org/record/10666110/files/figure.png" pageId="11" pageNumber="256">Fig. 28</figureCitation>
).
</paragraph>
<paragraph id="27147D6197470B4AE31FFCFD90BA3443" blockId="11.[817,1010,808,830]" box="[817,1010,808,830]" pageId="11" pageNumber="256">
<emphasis id="15DFA17397470B4AE31FFCFD90343440" box="[817,892,808,829]" italics="true" pageId="11" pageNumber="256">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="27147D6197470B4AE31FFC81973134FA" blockId="11.[817,1448,852,903]" pageId="11" pageNumber="256">
<emphasis id="15DFA17397470B4AE31FFC8190D53414" box="[817,925,852,873]" italics="true" pageId="11" pageNumber="256">Etymology</emphasis>
.
<emphasis id="15DFA17397470B4AE39CFC8097403417" box="[946,1032,853,874]" italics="true" pageId="11" pageNumber="256">Tenuitas</emphasis>
Latin for slender (segment X and harpago of inferior appendages).
</paragraph>
<paragraph id="27147D6197470B4AE31FFC4897873374" blockId="11.[817,1448,925,1033]" pageId="11" pageNumber="256">
<emphasis id="15DFA17397470B4AE31FFC4890C134CF" box="[817,905,925,946]" italics="true" pageId="11" pageNumber="256">Remarks</emphasis>
. The single male specimen of
<taxonomicName id="E0AB06E297470B4AE491FC4896E034CE" authorityName="Cartwright" authorityYear="2012" box="[1215,1448,925,947]" class="Insecta" family="Philopotamidae" genus="Hydrobiosella" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="256" phylum="Arthropoda" rank="species" species="tenuitas">
<emphasis id="15DFA17397470B4AE491FC4896E034CE" box="[1215,1448,925,947]" italics="true" pageId="11" pageNumber="256">Hydrobiosella tenuitas</emphasis>
</taxonomicName>
was collected from north-eastern
<collectingRegion id="E56FB38397470B4AE452FC6F97BB34AD" box="[1148,1267,954,976]" country="Australia" name="Queensland" pageId="11" pageNumber="256">Queensland</collectingRegion>
(latitude
<geoCoordinate id="429F1BA697470B4AE57EFC6F96D534AD" box="[1360,1437,954,976]" degrees="17" direction="south" minutes="16" orientation="latitude" pageId="11" pageNumber="256" precision="925" value="-17.266666">17°16'S</geoCoordinate>
).
<taxonomicName id="E0AB06E297470B4AE31FFC0297523491" authorityName="Cartwright" authorityYear="2012" box="[817,1050,983,1004]" class="Insecta" family="Philopotamidae" genus="Hydrobiosella" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="256" phylum="Arthropoda" rank="species" species="tenuitas">
<emphasis id="15DFA17397470B4AE31FFC0297523491" box="[817,1050,983,1004]" italics="true" pageId="11" pageNumber="256">Hydrobiosella tenuitas</emphasis>
</taxonomicName>
was cited in a checklist (
<bibRefCitation id="433A009097470B4AE530FC02902B3374" author="Walker, K. &amp; Neboiss, A. &amp; Dean, J. &amp; Cartwright, D." pageId="11" pageNumber="256" pagination="19 - 31" refId="ref11081" refString="Walker, K., Neboiss, A., Dean, J. and Cartwright, D. 1995. A preliminary investigation of the Caddis-flies (Insecta: Trichoptera) of the Queensland Wet Tropics. Australian Entomologist 22: 19 - 31." type="journal article" year="1995">Walker et al., 1995</bibRefCitation>
) as
<taxonomicName id="E0AB06E297470B4AE3A2FC2197553374" authority="Cartwright, 2012" authorityName="Cartwright" authorityYear="2012" box="[908,1053,1012,1033]" class="Insecta" family="Philopotamidae" genus="Hydrobiosella" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="256" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="15DFA17397470B4AE3A2FC2197553374" box="[908,1053,1012,1033]" italics="true" pageId="11" pageNumber="256">Hydrobiosella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EEC1C0897470B4AE40DFC2197273374" box="[1059,1135,1012,1033]" pageId="11" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
PT-1038.
</paragraph>
</subSubSection>
</treatment>
</document>