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<mods:title id="B26346071D09DB2C6E5D3AFF729D9044">A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)</mods:title>
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<subSubSection id="16AEF3826D0E79D040CEB964E677FFD9" pageId="168" pageNumber="169" type="nomenclature">
<paragraph id="EF66DD076488294DB126BFB682182166" pageId="168" pageNumber="169">
<taxonomicName id="6C85D5FAF94F2EE2AFA0551643F241EF" ID-CoL="7WS8G" authority="Lewis, 1901" authorityName="Lewis" authorityYear="1901" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) viridipennis" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="viridipennis" subGenus="Saprinus">Saprinus (Saprinus) viridipennis Lewis, 1901</taxonomicName>
Figs 603, 604-611, 612-618, 765
</paragraph>
</subSubSection>
<subSubSection id="95E16CED33ED0718A9886748C74DA0C3" pageId="168" pageNumber="169" type="reference_group">
<paragraph id="610CD35DC46872E660C56C8ECAD9FE93" pageId="168" pageNumber="169">
<taxonomicName id="8D415306DAA8F82D182073C71D043AF6" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus viridipennis" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="viridipennis">Saprinus viridipennis</taxonomicName>
Lewis, 1901: 245.
</paragraph>
<paragraph id="2FD955375CE66D1FCCBCEBB512BA4264" pageId="168" pageNumber="169">
<taxonomicName id="5852D7CF63E3921D65281819672542F4" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus desbordesi" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="desbordesi">Saprinus desbordesi</taxonomicName>
Auzat, 1916: 32 - Synonymized by
<bibRefCitation id="606D04EF7D41FD2830E08068A564BDBD" author="Dahlgren, G" journalOrPublisher="Opuscula Entomologica" pageId="214" pageNumber="215" pagination="213 - 224" title="Beitraege zur Kenntnis der Gattung Saprinus (Col. Histeridae)." volume="32" year="1967">Dahlgren (1967)</bibRefCitation>
: 220.
</paragraph>
</subSubSection>
<subSubSection id="4896B380E81736E066D33E55121CDD0D" pageId="168" pageNumber="169" type="type locality">
<paragraph id="FB60E3E15F3E01257E41D778B4ECBD49" pageId="168" pageNumber="169">Type locality.</paragraph>
<paragraph id="925A4ED804935BD3F24EEC0B29568CDA" pageId="168" pageNumber="169">Australia.</paragraph>
<caption id="F1570685C2DC36B72BEA9732452BEE78" pageId="168" pageNumber="169">
<paragraph id="DE408F68B3E42D3C393FAEE5455627DA" pageId="168" pageNumber="169">
Figure 603.
<taxonomicName id="F84C8164782FC65C6CF70856FCC99925" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) viridipennis" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="viridipennis" subGenus="Saprinus">Saprinus (Saprinus) viridipennis</taxonomicName>
Lewis, 1901 habitus, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="635B3C2172779FBF35EB76DC307CA715" pageId="168" pageNumber="169" type="type material">
<paragraph id="080724FE480B9A3261F90BD3AF89AB45" pageId="168" pageNumber="169">Type material examined.</paragraph>
<paragraph id="6C5C254B9CC6E5DCB9A8DD9AB35BE0FB" pageId="168" pageNumber="169">
<taxonomicName id="89CAFCDAD4766139443154DE70AD1446" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus viridipennis" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="viridipennis">Saprinus viridipennis</taxonomicName>
Lewis, 1901: Lectotype, present designation: ♀, side-mounted on a triangular mounting card with genitalia extracted and mounted on another mounting card under the specimen, except for left mesotarsus all tarsi are broken off to some degree, with the following labels: &quot;Australie / CH FRENCH&quot; (written); followed by: &quot;Prosternum / without striae&quot; (written); followed by: &quot;
<taxonomicName id="7642733845FE32339EA2B172CD2FEEB1" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="A11DA8CF2265E72292C630A83246C566" class="Insecta" family="Corydiidae" genus="Histeroidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Histeroidea viridipennis" order="Blattodea" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="viridipennis">viridipennis</taxonomicName>
/ Type Lewis&quot; (written); followed by: &quot;G. Lewis Coll. / B.M. 1926-369&quot; (printed); followed by:
<normalizedToken id="BFC40463B78BD5D30E9C5057416B5807" originalValue="“Type”">&quot;Type&quot;</normalizedToken>
(round, printed red-margined label); followed by:
<normalizedToken id="8F84F62D053AD9B5832C99C0B8B3DFB9" originalValue="“08-078”">&quot;08-078&quot;</normalizedToken>
(yellow, pencil-written label, added by the senior author); followed by: &quot;LECTOTYPE /
<taxonomicName id="F89740FF1EC7EA13278C2C53FB746E26" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus viridipennis" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="viridipennis">Saprinus viridipennis</taxonomicName>
/ Lewis, 1901 / Designated by / T. Lackner, 2008&quot; (red label, written) (BMNH). This species has been described from numerous specimens (&quot;Examples in the Belgium Museum and in my own cabinet.&quot; -
<bibRefCitation id="77096B96F26C6AB93641B1EA52274C6D" pageId="168" pageNumber="169">Lewis 1901</bibRefCitation>
: 245) and the lectotype designation fixes the identity of species. 1 paralectotype (present designation), ♀, with blue round label, followed by:
<normalizedToken id="7B06764048E04597D57A8F0878A35848" originalValue="“Co-type”">&quot;Co-type&quot;</normalizedToken>
(round, printed, yellow-margined label); followed by: &quot;Australie / CH FRENCH&quot; (written); followed by: &quot;
<taxonomicName id="9E1500DD40250E0641741B6E18C59AEE" lsidName="S. viridipennis" pageId="168" pageNumber="169" rank="species" species="viridipennis">S. viridipennis</taxonomicName>
/ Cotype Lewis&quot; (BMNH).
</paragraph>
<paragraph id="5C0D5C1E2731DBFAF1D63CE92171D0D8" pageId="168" pageNumber="169">
<taxonomicName id="0BDBADDBD9627FE565D00A06C088D3B6" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus desbordesi" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="desbordesi">Saprinus desbordesi</taxonomicName>
Auzat, 1916: 32: Lectotype, present designation: ♂, all tarsi and right mesotibia missing, male genitalia extracted, glued on the same mounting card as the specimen, with following labels: &quot;
<taxonomicName id="1C366DAE4D8981690224033FF330EE1D" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/ Desbordesi / Typus / Dr. Auzat det 1916&quot; (printed-written); followed by:
<normalizedToken id="335F8160339BFBF3FF5F9E134C811E2E" originalValue="“MUSÉUM">&quot;MUSEUM</normalizedToken>
PARIS / 1933 / Coll. DESBORDES&quot; (printed); followed by:
<normalizedToken id="453DF0433D40A48D6B71449662DEAA97" originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
(red label, printed); followed by: &quot;Sapr. VIRIDIPEN- / NIS LEW. / G Dahlgren det&quot; (printed-written); followed by: &quot;
<taxonomicName id="49E57E2226F16DEF53C227D4E2C39A78" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus desbordesi" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="desbordesi">Saprinus desbordesi</taxonomicName>
/ Auzat, 1916 / LECTOTYPE 2014 / des. T. Lackner&quot; (red label, written) (MNHN). Paralectotype (present designation): ♀, with genitalia extracted, glued onto the same label as specimen, all tarsi and left metatibia broken off, with the following labels: &quot;Desbordesi / Typus / dr. Auzat det. 1916&quot; (printed-written); followed by:
<normalizedToken id="3E89FB35E73F7704F4223E0EC395D91C" originalValue="“MUSÉUM">&quot;MUSEUM</normalizedToken>
PARIS / 1933 / Coll. DESBORDES&quot; (printed); followed by:
<normalizedToken id="1D3B6840DBBBC1476415188DE64A6A23" originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
(red label, printed); followed by: &quot;Sapr. VIRIDIPEN- / NIS LEW. / G Dahlgren det&quot; (printed-written); followed by: &quot;
<taxonomicName id="C2204CDDC183F8E02B51DEA3B28980EC" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus desbordesi" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="desbordesi">Saprinus desbordesi</taxonomicName>
/ Auzat, 1916 / PARALECTOTYPE / des. T. Lackner 2014&quot; (red label, written) (MNHN). Paralectotype (present designation), ♀, pinned, two segments of right mesotarsus, entire left mesotarsus and two segments of left metatarsus missing, with the following labels:
<normalizedToken id="643469BA11072CE8FF788C9A44CA1D89" originalValue="“♀”">&quot;&quot;</normalizedToken>
(printed); followed by: &quot;S. Austral.&quot; (written); followed by: &quot;S. Austral. / coll.
<normalizedToken id="922EC5C4BB10C741E4E50319CE7585FF" originalValue="Théry”">Thery&quot;</normalizedToken>
(written); followed by: &quot;
<taxonomicName id="4917BFC83365D194F8AFCBFBF2386083" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/ Desbordesi / Typus / D. Auzat det 1916&quot; (printed-written); followed by:
<normalizedToken id="2225618C7C32D7C0B4794DD001D7FAAD" originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
(red label, printed); followed by: &quot;
<taxonomicName id="B1FECA660517E5E3A09C2475B22C240D" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="E6CE36785DCB49B0925077D1A38FA8BD" class="Insecta" family="Corydiidae" genus="Histeroidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Histeroidea desbordesi" order="Blattodea" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="desbordesi">desbordesi</taxonomicName>
/ Auzat, 1916 / PARALECTOTYPE / des. T.
<bibRefCitation id="AB7BF206561D643FD6BA0ED2D0BE6478" author="Lackner, T" journalOrPublisher="ZooKeys" pageId="216" pageNumber="217" title="Redescription of the enigmatic neotropical inquiline Paramyrmetesfoveipennis Bruch, 1929 (Coleoptera: Histeridae) with notes on myrmecophily in the Saprininae subfamily" url="http://dx.doi.org.10.3897/zookeys.675.12690" volume="675" year="2017">Lackner 2017</bibRefCitation>
&quot; (red label, written) (MNHN, coll.
<normalizedToken id="6245B0AA5074FE7BFEE809C2CE9A175D" originalValue="Thérond">Therond</normalizedToken>
). This species has been described from three specimens; two of them were from Desbordes collection and third was from
<normalizedToken id="BDA61FD13E27988DD7300C33FE0FE8E5" originalValue="Auzats">Auzat's</normalizedToken>
collection (
<bibRefCitation id="AD83129C68E21AD2CB812C2613346E76" author="Auzat, V" journalOrPublisher="Bulletin de la Societe entomologique de France" pageId="213" pageNumber="214" pagination="132 - 133" title="Description d'un Saprinus nouveau d'Australie." volume="1916" year="1916">Auzat 1916</bibRefCitation>
). The lectotype designation fixes the species identity.
</paragraph>
</subSubSection>
<subSubSection id="FA2666FF9005053BAFF2EE5C2FFAF485" lastPageId="169" lastPageNumber="170" pageId="168" pageNumber="169" type="materials_examined">
<paragraph id="7E5A4165EB821249B02832A10B0F6939" pageId="168" pageNumber="169">Additional material examined.</paragraph>
<paragraph id="F48050759C591B0694E360E2772A8C1C" lastPageId="169" lastPageNumber="170" pageId="168" pageNumber="169">
AUSTRALIA. Victoria: 1 ♂, Merinqur, 3.i.1931, C.E. Clarke (AMNZ); 2 specs., Melbourne, without further data (ZMHUB). South Australia: 1 spec., Gawler Geb., without further data (ZMHUB); 1 ♂ &amp; 1 ♀, S. Australia, without further data (BMNH); 1 spec., Maralinga, viii.-x.1956, F.L. Hill (BMNH); 2 ♂♂ &amp; 1 ♀, Killalpanima, 100 miles of L. Eyre, H.J. Hillier (BMNH); 1
<pageBreakToken id="5ED2EF02A2790C611C0384E480D150D2" pageId="169" pageNumber="170" start="start">spec</pageBreakToken>
., Kingonya, R. Harvey (SAMA); 1 spec., Oodnadatta, Blackburn (SAMA); 1 spec., Purple Downs (SAMA); 2 spec., Innamincka, W. Lamb (SAMA); 5 specs.,
<normalizedToken id="FBBA8F85D5DFDD411559656FDA9BEB25" originalValue="Coopers">Cooper's</normalizedToken>
Creek, J.G. Reuther (SAMA); 3 specs., Mt. Serle, N Flinders Ra., Hale &amp; Tindale (SAMA). Northern Territory: 1 ♂ &amp; 3 ♀♀, Hermansburg, H.J. Hillier, 1908-177 (BMNH); Coniston Station, near Alice Springs, M.W. Mules (SAMA). New South Wales: 2 ♂♂, Alexandria, New South Wales, W. Stalker, x.[19]05 (BMNH); 4 specs., Lake Callabona, A. Zietz (SAMA). Queensland: 2 specs., Cunnamulla, H. Hardcastle (both exs. SAMA). Western Australia: 1 ♂, Nicol Bay Distr., Dr. Clement, 1900-220; 1 ♀, West Australia, 95-64 (BMNH).
</paragraph>
<paragraph id="2E2C10C6494CFA08A5E820DFF89C13FC" pageId="169" pageNumber="170">Unknown localities: 3 specs., Australia occid. 1192, without further data (HNMH).</paragraph>
</subSubSection>
<subSubSection id="F63C151E13ECF39838B322A44D9716E8" pageId="169" pageNumber="170" type="biology_ecology">
<paragraph id="D01D08D459F6B5041A6FA44108CA7AF9" pageId="169" pageNumber="170">Biology.</paragraph>
<paragraph id="CC4A1FDE06048B4A831D61287AE735DF" pageId="169" pageNumber="170">Presumably similar to congeners; an uncommon species.</paragraph>
</subSubSection>
<subSubSection id="14FF703326CF0667279F08E06D18B78E" pageId="169" pageNumber="170" type="distribution">
<paragraph id="A1F6369343EC518D9E8ED835E5F43AF1" pageId="169" pageNumber="170">Distribution.</paragraph>
<paragraph id="9E912E5FA7354AA0DFF221EE5397FA72" pageId="169" pageNumber="170">Australia: Victoria, South Australia, Western Australia, Northern Territory, New South Wales &amp; Queensland (Fig. 765).</paragraph>
</subSubSection>
<subSubSection id="E361C56D0BD5B1F960DBB006ABE0C137" pageId="169" pageNumber="170" type="remarks">
<paragraph id="240F34E963F10B6A8BE0266A53CC922A" pageId="169" pageNumber="170">Remarks.</paragraph>
<paragraph id="B2B6C82EA5FC09839783D03ADEFE5DC3" pageId="169" pageNumber="170">
A rare species, easily distinguishable from the most similar
<taxonomicName id="F98FDAAFD2AF9C24FB79808FF869FF97" lsidName="S. cyaneus" pageId="169" pageNumber="170" rank="species" species="cyaneus">S. cyaneus</taxonomicName>
by the presence of long amber setae of pronotal hypomeron easily visible from above, the complete absence of carinal prosternal striae and setose lateral prosternal striae. The protibiae differ from
<taxonomicName id="824FED684D1F96671C0485ECDE9F2F24" lsidName="S. cyaneus" pageId="169" pageNumber="170" rank="species" species="cyaneus">S. cyaneus</taxonomicName>
or
<taxonomicName id="AACC7269C335EEE1E1635857B4955277" lsidName="S. laetus" pageId="169" pageNumber="170" rank="species" species="laetus">S. laetus</taxonomicName>
by their larger triangular teeth.
</paragraph>
</subSubSection>
<subSubSection id="6020630E8D9440ED13F85C92E9BC57CF" lastPageId="172" lastPageNumber="173" pageId="169" pageNumber="170" type="description">
<paragraph id="DC35B2E4756F09799738FDA677605EF5" pageId="169" pageNumber="170">Re-description.</paragraph>
<paragraph id="1A00EB54E2C66BF7304AA465E205CEE6" pageId="169" pageNumber="170">Body length: PEL: 4.25-5.50 mm; APW: 1.60-2.00 mm; PPW: 3.50-4.70 mm; EL: 2.60-3.50 mm; EW: 3.75-5.00 mm. Body (Fig. 603) rectangular oval, convex, elytra widest at humeri, elytra stark green, shining, metallic; pronotum purple, metallic; legs, mouthparts and antennal scape castaneous; antennal club light brown.</paragraph>
<paragraph id="05AB3ADBF0B41867A210DC78339FC1B5" pageId="169" pageNumber="170">Antennal scape (Fig. 604) triangularly thickened, with several long setae; antennal club large, wider than long, depressed dorso-ventrally, somewhat truncate apically, dorsally on lower fourth asetose, rest of club covered with sparse to dense short sensilla intermingled with sparse longer erect setae; ventrally four large oval sensory structures of antennal club (Figs 605) easily observable; vesicle(s) not examined.</paragraph>
<caption id="4E4052064B63316AE7457271F4C6EE7A" pageId="169" pageNumber="170">
<paragraph id="61D8948D67F9897E013DB6D28E52F498" pageId="169" pageNumber="170">
Figures 604-611. 604
<taxonomicName id="0F260873D42FFA8B3487BB86932145CD" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) viridipennis" order="Coleoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="viridipennis" subGenus="Saprinus">Saprinus (Saprinus) viridipennis</taxonomicName>
Lewis, 1901 head, dorsal view 605 antennal club, ventral view 606 mentum, ventral view 607 clypeus 608 propygidium + pygidium 609 prosternum 610 mesoventrite 611 lateral disc of metaventrite + metepisternum.
</paragraph>
</caption>
<paragraph id="75C48E8B70F27E5F95C24987960060CF" pageId="169" pageNumber="170">Mandibles dorso-laterally punctuate, rounded, stout, outer margin slightly carinate, mandibular apex acute, sub-apical tooth on left mandible obtuse; labrum finely and sparsely punctuate, convex, broadly depressed medially; labral pits present, each with two short labral setae; terminal labial palpomere elongate, pointed apically, approximately twice as long as pen-ultimate, its length about three times its width; terminal maxillary palpomere elongate, about twice as long as pen-ultimate, approximately three times as long as wide, pointed apically; mentum (Fig. 606) sub-trapezoid, laterally with a row of short ramose setae, several short setae present also on imbricate disc; anterior margin broadly inwardly arcuate medially, a tuft of four longer setae present near median notch.</paragraph>
<paragraph id="1CB3CE63AB4A93397A0A4EEBEC617ACF" pageId="169" pageNumber="170">Clypeus (Fig. 607) faintly concave, sloping down laterally, microscopically punctate; frontal and supraorbital striae complete; frontal disc (Fig. 604) punctate, punctures separated by about their own to twice their diameter; eyes convex, well visible from above.</paragraph>
<paragraph id="DD7AE4BA5201A7876CCBFB61325EBE44" lastPageId="170" lastPageNumber="171" pageId="169" pageNumber="170">
Pronotal sides (Fig. 603) on basal half gently narrowing anteriorly, on apical half strongly narrowing, apical angles prominent, pronotal depressions present, shallow, an
<pageBreakToken id="2DCE2A6546847C558BDF3A47D042D0C6" pageId="170" pageNumber="171" start="start">terior</pageBreakToken>
incision for head moderate; marginal pronotal stria carinate, visible along its entire length from dorsal view, ending short of pronotal base; pronotal disc laterally with a depressed band of deep dense coarse elongate punctures originating approximately in pronotal depressions, reaching basal angles of pronotum, between it and pronotal margin a broad glabrous band present; rest of the pronotal disc with only scattered microscopic punctation (or even glabrous in some specimens); several rows of punctures present along pronotal base, ante-scutellar area impunctate; pronotal hypomeron in dense amber setae well visible also from dorsal view; scutellum small, visible.
</paragraph>
<paragraph id="2A978410CA727FEE89819706C390062B" pageId="170" pageNumber="171">Elytral epipleura almost impunctate; marginal epipleural stria complete; marginal elytral stria well impressed and carinate, continued as weakened, but complete apical elytral stria that is connected to incomplete sutural elytral stria. Humeral elytral stria well impressed on basal third, sometimes connected to rather long median fragment of inner subhumeral stria creating thus a complementary dorsal elytral stria parallel to first; elytra usually with four thin impunctate dorsal elytral striae 1-4; striae 1-3 originating at elytral base and slightly surpassing elytral half apically, fourth dorsal elytral stria occasionally slightly shortened basally, can be variously long apically, even intermittent; in first elytral interval often elongate strioles present; fourth elytral stria basally curved toward shortened sutural elytral stria, but never connected with it; sutural elytral stria abbreviated on basal fourth to fifth, well-impressed, apically connected with apical elytral stria; elytral disc on apical half (roughly) punctate, punctures very fine, sparse, separated by several times their diameter, slightly entering elytral intervals and reaching beyond elytral mid-length basally between fourth elytral and sutural stria; elytral flanks impunctate; punctures reach elytral apex.</paragraph>
<paragraph id="D709148640EF76C832F8FE344F09E470" pageId="170" pageNumber="171">Propygidium (Fig. 608) densely punctate laterally, punctures becoming sparser medially; pygidium (Fig. 608) with more even, but sparser punctation.</paragraph>
<paragraph id="8B1E46EF31A278B22F21533FFCDAC072" pageId="170" pageNumber="171">Anterior margin of median portion of prosternum (Fig. 609) straight; marginal prosternal stria present laterally and also as medial fragment; prosternal process on prosternal keel convex, glabrous; carinal prosternal striae absent; lateral prosternal striae carinate, rather short, surface around them with long amber setae.</paragraph>
<paragraph id="32713A273E4EDB4FB5E2039CC3E92C60" pageId="170" pageNumber="171">Discal marginal mesoventral stria (Fig. 610) weakly impressed, complete, anteriorly faintly inwardly arcuate, at times interrupted medially; disc slightly convex, glabrous; meso-metaventral stria absent; meso-metaventral suture visible; intercoxal disc of metaventrite glabrous, only in post-metacoxal area with several tiny punctures, in males with faint median longitudinal depression, in females slightly convex; lateral metaventral stria (Fig. 611) well impressed, carinate, straight, shortened; lateral disc of metaventrite (Fig. 611) slightly concave, with sparse shallow round setigerous punctures separated by less than their to about their own diameter, between large punctures another type of much finer sparse punctures present; metepisternum (Fig. 611) similar, but punctures somewhat deeper, on fused metepimeron punctures disappear; metepisternal stria present, deeply impressed, often complete. Intercoxal disc of first abdominal ventrite completely striate laterally, glabrous.</paragraph>
<paragraph id="3C7ED605FC15E2C741740077614E30F9" lastPageId="172" lastPageNumber="173" pageId="170" pageNumber="171">
Protibia slightly dilated, outer margin with two large triangular teeth topped by triangular denticle, followed by one more lower tooth topped by denticle and another
<pageBreakToken id="5F4913B1E16351C87AA4C7E836E48A86" pageId="171" pageNumber="172" start="start">two</pageBreakToken>
tiny denticles diminishing in size proximally; setae of outer row regular, short; protarsal groove shallow; anterior protibial stria present on basal half, next obliterated; setae of median row approximately as long as those of outer row, but sparser; two tarsal denticles present near tarsal insertion; protibial spur bent, large and stout, growing out from apical margin of protibia; apical margin of protibia ventrally with three tiny denticles; outer part of posterior surface obscurely variolate, punctate, separated from
<pageBreakToken id="8D6841FDB97231C7E0E866AD400C62A3" pageId="172" pageNumber="173" start="start">glabrous</pageBreakToken>
and narrow median part of posterior surface; posterior protibial stria complete, bearing along its length a dense row of long, well-sclerotized setae; inner row of setae long, brush-like.
</paragraph>
<paragraph id="C06D4EC14E7C6C81EF5F9B69ED893FBE" pageId="172" pageNumber="173">Mesotibia on outer margin with about five widely separated denticles, fourth and fifth denticles situated on prominent teeth; median tooth of mesotibia bears two denticles, best observable from ventral view; setae of outer row strongly sclerotized, sparse, almost as long as denticles; setae of median row shorter and finer than those of outer row; posterior mesotibial stria shortened on apical half; mesotibial spur rather long and stout; on anterior face of mesotibia a row of about four widely-spaced denticles present; anterior face of mesotibia with scattered fine punctation; anterior mesotibial stria incomplete; inner anterior denticles weakly developed, usually only one or two present; apical margin of mesotibia with a dense row of about four stout denticles; setae of inner row rather long, sparse. Metatibia longer and more slender than mesotibia; teeth on its outer margin lower than those of mesotibia; denticles of both rows sparser.</paragraph>
<paragraph id="78BA5E363B8B0FC6BFB68463F9852369" pageId="172" pageNumber="173">Male genitalia. Eighth sternite (Figs 612-613) about as broad as long, strongly sclerotized, fused medially, with sparse pores and pseudopores, apex laterally with two short setae, vela present, short, laterally with two tufts of short setae; eighth tergite apically faintly inwardly arcuate, basally strongly inwardly arcuate; eighth tergite and eighth sternite fused laterally (Fig. 614). Ninth tergite (Figs 615-616) laterally strongly sclerotized, medially with a strongly sclerotized part resembling a fusion seen also in other congeners, with pores and pseudopores, well visible, especially laterally; ninth tergite basally faintly inwardly arcuate; tenth tergite basally inwardly arcuate, apices strongly sclerotized; spiculum gastrale (Fig. 615) dilated on both ends, otherwise almost parallel-sided, apical end strongly sclerotized, basal end slightly inwardly arcuate, cordate. Aedeagus (Figs 617-618) on basal half almost parallel-sided, on apical half dilated and thickened; parameres fused along basal two-thirds; aedeagus laterally slightly curved from lateral view; basal piece of aedeagus short, ratio of its length: length of parameres approximately 1: 6; apex of aedeagus slightly dilated, ventrally with a tuft of microscopic setae.</paragraph>
<caption id="13B6395F16EF54876BEE80CF84BB1062" pageId="172" pageNumber="173">
<paragraph id="59605C6F665AC223B0062E940DBE716B" pageId="172" pageNumber="173">
Figures 612-618. 612
<taxonomicName id="E257AE310D9C06FE8C8BAD9C137019DD" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) viridipennis" order="Coleoptera" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="viridipennis" subGenus="Saprinus">Saprinus (Saprinus) viridipennis</taxonomicName>
Lewis, 1901 male terminalia: 8th sternite + 8th tergite, ventral view 613 ditto, dorsal view 614 ditto, lateral view 615 male terminalia: 9th + 10th tergites, dorsal view; spiculum gastrale, ventral view 616 male terminalia: 9th + 10th tergites; spiculum gastrale, lateral view 617 male terminalia: aedeagus, dorsal view 618 ditto, lateral view.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>