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<document id="ACC06896CA2FC8F63009055154321A6F" ID-CLB-Dataset="28990" ID-DOI="10.11646/zootaxa.4461.1.7" ID-GBIF-Dataset="1fbcced0-ec5a-4afe-bcb1-496990e850c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="1459872" ID-ZooBank="3FACE488-20DC-46A4-A511-026B7F2DBEA2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539336504258" checkinUser="plazi" docAuthor="Evsyukov, Aleksandr, Golovatch, Sergei &amp; Reip, Hans S." docDate="2018" docId="610E87FB9319FF8AFF0DF58DFEF319DD" docLanguage="en" docName="zootaxa.4461.1.7.pdf" docOrigin="Zootaxa 4461 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Julus lignaui Verhoeff 1910" docType="treatment" docVersion="6" lastPageNumber="105" masterDocId="9D37FF839316FF9AFF9AF75AFFFE1E2B" masterDocTitle="The millipede genus Julus Linnaeus, 1758 in the Caucasus (Diplopoda: Julida: Julidae)" masterLastPageNumber="117" masterPageNumber="89" pageNumber="104" updateTime="1698579428612" updateUser="ExternalLinkService">
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<mods:title id="23084304AD8A51CD0478118FD4DA7646">The millipede genus Julus Linnaeus, 1758 in the Caucasus (Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="799A4338F8534D34B8B34DFFCEE79D51">Reip, Hans S.</mods:namePart>
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<taxonomicName id="2EA74D6E9319FF95FF0DF58DFE0A1CD9" ID-CoL="3QSYG" authority="Verhoeff, 1910" authorityName="Verhoeff" authorityYear="1910" box="[151,500,727,754]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="15" pageNumber="104" phylum="Arthropoda" rank="species" species="lignaui">
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<emphasis id="DBD3EAFF9319FF95FF0DF58DFEC91CDA" bold="true" box="[151,311,727,753]" italics="true" pageId="15" pageNumber="104">Julus lignaui</emphasis>
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<figureCitation id="719C2A689319FF95FF0DF5A0FF111D38" box="[151,239,762,787]" captionStart="FIGURE 1" captionStartId="4.[151,249,1820,1842]" captionTargetBox="[344,1236,854,1775]" captionTargetId="figure@4.[340,1248,841,1798]" captionTargetPageId="4" captionText="FIGURE 1. Antennae in Caucasian species of Julus, lateral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. dagestanus sp. n. H: J. khostensis sp. n. Scale bar: 0.3 mm." httpUri="https://zenodo.org/record/1459874/files/figure.png" pageId="15" pageNumber="104">Figs 1E</figureCitation>
,
<figureCitation id="719C2A689319FF95FF61F5A1FEE31D38" box="[251,285,763,787]" captionStart="FIGURE 2" captionStartId="5.[151,249,1601,1623]" captionTargetBox="[340,1248,985,1580]" captionTargetId="figure@5.[340,1248,985,1580]" captionTargetPageId="5" captionText="FIGURE 2. Gnathochilaria in Caucasian species of Julus, ventral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459876/files/figure.png" pageId="15" pageNumber="104">2E</figureCitation>
,
<figureCitation id="719C2A689319FF95FEB2F5A1FE951D38" box="[296,363,763,787]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (B–D), J. jedryczkowskii Golovatch, 1981 (E–F), J. kubanus Verhoeff, 1921 (G–I), J. lignaui Verhoeff, 1910 (J–M), J. lindholmi Lohmander, 1936 (N– P), J. subalpinus Lohmander, 1936 (Q–R), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views B–O; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (A–C, E, G, H, J–O, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="15" pageNumber="104">3J–M</figureCitation>
,
<figureCitation id="719C2A689319FF95FEECF5A0FE431D38" box="[374,445,762,787]" captionStart="FIGURE 6" captionStartId="10.[151,250,1294,1316]" captionTargetBox="[340,1248,199,1261]" captionTargetId="figure@10.[340,1248,193,1273]" captionTargetPageId="10" captionText="FIGURE 6. Male legs 2 in Julus lignaui Verhoeff, 1910 (A–D) and J. lindholmi Lohmander, 1936 (E–G), caudal views. A: from Krasnaya Polyana, Krasnodar Province. B: from Teberda, Karachaevo-Cherkessia. C: from Mestia, Georgia. D: from Teberda, Karachaevo-Cherkessia. E: from Svanetia, Georgia. F: from near Teberda, Karachaevo-Cherkessia. G: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (A–F), not to scale (G)." httpUri="https://zenodo.org/record/1459884/files/figure.png" pageId="15" pageNumber="104">6A–D</figureCitation>
,
<figureCitation id="719C2A689319FF95FE52F5A1FE1B1D38" box="[456,485,763,787]" captionStart="FIGURE 11" captionStartId="16.[151,250,1572,1594]" captionTargetBox="[350,1231,514,1553]" captionTargetId="figure@16.[340,1248,507,1563]" captionTargetPageId="16" captionText="FIGURE 11. Male gonopods in Julus lignaui Verhoeff, 1910 (A–B from Krasnaya Polyana, Krasnodar Province; C–D from Mestia, Georgia; E–H from Teberda, Karachaevo-Cherkessia). A: promere with flagella, caudal view. B, D, H: left gonopods, mesal view. C, E, G: promere, caudal view. F: right gonopod, mesal view. Scale bar: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459894/files/figure.png" pageId="15" pageNumber="104">11</figureCitation>
,
<figureCitation id="719C2A689319FF95FE6BF5A1FDDF1D38" box="[497,545,763,787]" captionStart="FIGURE 14" captionStartId="20.[151,250,1014,1036]" captionTargetBox="[340,1248,373,993]" captionTargetId="figure@20.[340,1248,373,993]" captionTargetPageId="20" captionText="FIGURE 14. Mesomere and opisthomere in Caucasian species of Julus, oral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936 G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. Scale bars: 0.3 mm. Abbreviations: fl, flagellum; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere." httpUri="https://zenodo.org/record/1459900/files/figure.png" pageId="15" pageNumber="104">14E</figureCitation>
,
<figureCitation id="719C2A689319FF95FDB7F5A0FDA21D38" box="[557,604,762,787]" captionStart="FIGURE 15" captionStartId="20.[151,249,1687,1709]" captionTargetBox="[340,1248,1178,1666]" captionTargetId="figure@20.[340,1248,1178,1666]" captionTargetPageId="20" captionText="FIGURE 15. Ventral edge of male segments 7 in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936 C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459902/files/figure.png" pageId="15" pageNumber="104">15E</figureCitation>
,
<figureCitation id="719C2A689319FF95FDF2F5A0FD691D38" box="[616,663,762,787]" captionStart="FIGURE 16" captionStartId="21.[151,249,1286,1308]" captionTargetBox="[340,1247,583,1264]" captionTargetId="figure@21.[340,1247,583,1264]" captionTargetPageId="21" captionText="FIGURE 16. Striation on metazonites in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459904/files/figure.png" pageId="15" pageNumber="104">16E</figureCitation>
,
<figureCitation id="719C2A689319FF95FD39F5A1FD2B1D38" box="[675,725,763,787]" captionStart="FIGURE 18" captionStartId="22.[151,249,1951,1973]" captionTargetBox="[340,1248,1166,1930]" captionTargetId="figure@22.[340,1248,1166,1930]" captionTargetPageId="22" captionText="FIGURE 18. Female vulvae and legs 2 in Caucasian species of Julus. A: J. lignaui Verhoeff, 1910. B: J. lindholmi Lohmander, 1936. C: J. subalpinus Lohmander, 1936. D: J. dagestanus sp. n. Abbreviations: vu, vulva; op, operculum; bu, bursa; am1 and am2, ampulla 1 and ampulla 2; l2, coxa and prefemur of leg 2. Scale bar: 0.3 mm." httpUri="https://zenodo.org/record/1459908/files/figure.png" pageId="15" pageNumber="104">18A</figureCitation>
, Map 1
</paragraph>
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<taxonomicName id="2EA74D6E9319FF95FF0DF418FED11D73" ID-CoL="3QT6K" authorityName="Linnaeus" authorityYear="1758" box="[151,303,834,856]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="15" pageNumber="104" phylum="Arthropoda" rank="species" species="terrestris">
<emphasis id="DBD3EAFF9319FF95FF0DF418FED11D73" box="[151,303,834,856]" italics="true" pageId="15" pageNumber="104">Julus terrestris</emphasis>
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—
<treatmentCitation id="680610FC9319FF95FED3F418FDFF1D73" author="" box="[329,513,834,856]" page="113" pageId="15" pageNumber="104">
<bibRefCitation id="8D364B1C9319FF95FED3F418FE2C1D73" box="[329,466,834,856]" pageId="15" pageNumber="104" refId="ref19444" refString="Lignau, N. (1903) [Myriapods of the Black Sea coast of the Caucasus]. Die Myriopoden am Kaukasischen Schwarzmeerufer. Memoires de la Societe Imperiale des Naturalistes de la Nouvelle-Russie. [Zapiski Imperatorskogo Novorossiyskogo obshchestva estestvoispytateley. Odessa], 25 (1), 82 - 149. [in Russian and German]" type="journal article">Lignau, 1903</bibRefCitation>
: 113
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(R);
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<bibRefCitation id="8D364B1C9319FF95FDACF418FD0C1D73" box="[566,754,834,856]" pageId="15" pageNumber="104" refId="ref19887" refString="Muralewicz, W. S. (1913) [Arthropods. - Arthropoda. Chapter VII. Class of myriapods. - Myriapoda]. Proceedings of the Society for the Study of the Black Sea Coast. [Trudy Obshchestva izucheniya Chernomorskogo poberezhya], 2, 215 - 221. [Sankt-Peterburg, in Russian]" type="book chapter">Muralewicz, 1913</bibRefCitation>
: 231
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(R).
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</paragraph>
<paragraph id="E91836ED9319FF95FF0DF43BFDA71D5C" blockId="15.[151,1436,834,1040]" box="[151,601,865,887]" pageId="15" pageNumber="104">
<treatmentCitationGroup id="C9B711C39319FF95FF0DF43BFDA71D5C" box="[151,601,865,887]" pageId="15" pageNumber="104">
<taxonomicName id="2EA74D6E9319FF95FF0DF43BFEA91D5C" ID-CoL="3QT5D" authorityName="Latzel" authorityYear="1884" box="[151,343,865,887]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="15" pageNumber="104" phylum="Arthropoda" rank="species" species="scandinavius">
<emphasis id="DBD3EAFF9319FF95FF0DF43BFEA91D5C" box="[151,343,865,887]" italics="true" pageId="15" pageNumber="104">Julus scandinavius</emphasis>
</taxonomicName>
—
<treatmentCitation id="680610FC9319FF95FEEBF43BFDD41D5C" author="" box="[369,554,865,887]" page="138" pageId="15" pageNumber="104">
<bibRefCitation id="8D364B1C9319FF95FEEBF43BFE041D5C" box="[369,506,865,887]" pageId="15" pageNumber="104" refId="ref19444" refString="Lignau, N. (1903) [Myriapods of the Black Sea coast of the Caucasus]. Die Myriopoden am Kaukasischen Schwarzmeerufer. Memoires de la Societe Imperiale des Naturalistes de la Nouvelle-Russie. [Zapiski Imperatorskogo Novorossiyskogo obshchestva estestvoispytateley. Odessa], 25 (1), 82 - 149. [in Russian and German]" type="journal article">Lignau, 1903</bibRefCitation>
: 138
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(R).
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<treatmentCitationGroup id="C9B711C39319FF95FF0DF4DAFDEA1DBD" box="[151,532,896,918]" pageId="15" pageNumber="104">
<taxonomicName id="2EA74D6E9319FF95FF0DF4DAFEDB1DBD" ID-CoL="6NBQT" authorityName="Latzel" authorityYear="1884" box="[151,293,896,918]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="15" pageNumber="104" phylum="Arthropoda" rank="species" species="ligulifer">
<emphasis id="DBD3EAFF9319FF95FF0DF4DAFEDB1DBD" box="[151,293,896,918]" italics="true" pageId="15" pageNumber="104">Julus ligulifer</emphasis>
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—
<treatmentCitation id="680610FC9319FF95FEA5F4DAFE1E1DBD" author="" box="[319,480,896,918]" page="3" pageId="15" pageNumber="104">
<bibRefCitation id="8D364B1C9319FF95FEA5F4DAFE371DBD" box="[319,457,896,918]" pageId="15" pageNumber="104" refId="ref18080" refString="Attems, C. (1907) Myriopoden aus der Krim und dem Kaukasus von Dr. A. Stuxberg gesammelt. Arkiv for zoologi, 3 (25), 1 - 16." type="journal article">Attems, 1907</bibRefCitation>
: 3
</treatmentCitation>
(M).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1BD65669319FF95FF0DF4C4FEDF1A3B" pageId="15" pageNumber="104" type="reference_group">
<paragraph id="E91836ED9319FF95FF0DF4C4FDE51D9F" blockId="15.[151,1436,834,1040]" box="[151,539,926,948]" pageId="15" pageNumber="104">
<treatmentCitationGroup id="C9B711C39319FF95FF0DF4C4FDE51D9F" box="[151,539,926,948]" pageId="15" pageNumber="104">
<taxonomicName id="2EA74D6E9319FF95FF0DF4C4FE151D9F" ID-CoL="3QSYG" authority="Verhoeff, 1910: 353" authorityName="Verhoeff" authorityPageNumber="353" authorityYear="1910" box="[151,491,926,948]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="15" pageNumber="104" phylum="Arthropoda" rank="species" species="lignaui">
<emphasis id="DBD3EAFF9319FF95FF0DF4C4FEE51D9F" box="[151,283,926,948]" italics="true" pageId="15" pageNumber="104">Julus lignaui</emphasis>
<treatmentCitation id="680610FC9319FF95FEBBF4C4FE151D9F" author="" box="[289,491,926,948]" page="353" pageId="15" pageNumber="104">
<bibRefCitation id="8D364B1C9319FF95FEBBF4C4FE451D9F" box="[289,443,926,948]" pageId="15" pageNumber="104" refId="ref20298" refString="Verhoeff, K. W. (1910) Ueber Dipopoden. 19. (39.) Aufsatz. Juliden und Ascospermophora. Jahreshefte des Vereins fur vaterlandische Naturkunde im Wurttemberg, 66, 337 - 398." type="journal article">Verhoeff, 1910</bibRefCitation>
: 353
</treatmentCitation>
</taxonomicName>
(D).
</treatmentCitationGroup>
</paragraph>
<paragraph id="E91836ED9319FF95FF0DF4E7FD761DF8" blockId="15.[151,1436,834,1040]" box="[151,648,957,979]" pageId="15" pageNumber="104">
<treatmentCitationGroup id="C9B711C39319FF95FF0DF4E7FD761DF8" box="[151,648,957,979]" pageId="15" pageNumber="104">
<taxonomicName id="2EA74D6E9319FF95FF0DF4E7FE781DF8" ID-CoL="84B7Z" box="[151,390,957,979]" class="Diplopoda" family="Julidae" genus="Micropodoiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="15" pageNumber="104" phylum="Arthropoda" rank="species" species="lignaui">
<emphasis id="DBD3EAFF9319FF95FF0DF4E7FE781DF8" box="[151,390,957,979]" italics="true" pageId="15" pageNumber="104">Micropodoiulus lignaui</emphasis>
</taxonomicName>
—
<treatmentCitation id="680610FC9319FF95FE3AF4E7FDA71DF8" author="" box="[416,601,957,979]" page="380" pageId="15" pageNumber="104">
<bibRefCitation id="8D364B1C9319FF95FE3AF4E7FDD71DF8" box="[416,553,957,979]" pageId="15" pageNumber="104" refId="ref19505" refString="Lignau, N. (1915) Vielfussler aus Abchasien. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de Petrograd. [Ezhegodnik zoologicheskogo muzeya Imperatorskoy akademii nauk. Petrograd], 19, 349 - 401." type="journal article">Lignau, 1915</bibRefCitation>
: 380
</treatmentCitation>
(R).
</treatmentCitationGroup>
</paragraph>
<paragraph id="E91836ED9319FF95FF0DF486FEDF1A3B" blockId="15.[151,1436,834,1040]" pageId="15" pageNumber="104">
<treatmentCitationGroup id="C9B711C39319FF95FF0DF486FEDF1A3B" pageId="15" pageNumber="104">
<taxonomicName id="2EA74D6E9319FF95FF0DF486FEE21DD9" ID-CoL="3QSYG" authorityName="Verhoeff" authorityYear="1910" box="[151,284,988,1010]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="15" pageNumber="104" phylum="Arthropoda" rank="species" species="lignaui">
<emphasis id="DBD3EAFF9319FF95FF0DF486FEE21DD9" box="[151,284,988,1010]" italics="true" pageId="15" pageNumber="104">Julus lignaui</emphasis>
</taxonomicName>
—
<treatmentCitation id="680610FC9319FF95FEACF487FE141DD9" author="" box="[310,490,988,1010]" page="1791" pageId="15" pageNumber="104">
<bibRefCitation id="8D364B1C9319FF95FEACF487FE531DD9" box="[310,429,988,1010]" pageId="15" pageNumber="104" refId="ref19345" refString="Lang, J. (1959) [To the knowledge of millipedes (Diplopoda) of the USSR territory]. Zoologicheskii zhurnal, 38 (12), 1790 - 1796. [in Russian, a summary in English]" type="journal article">Lang, 1959</bibRefCitation>
: 1791
</treatmentCitation>
(M);
<treatmentCitation id="680610FC9319FF95FDBDF486FCE81DD9" author="" box="[551,790,988,1010]" page="393" pageId="15" pageNumber="104">
<bibRefCitation id="8D364B1C9319FF95FDBDF486FD1B1DD9" box="[551,741,988,1010]" pageId="15" pageNumber="104" refId="ref19184" refString="Kobakhidze, D. N. (1965) [A list of millipedes (Diplopoda) of SSR Georgia]. Fragmenta Faunistica, 11 (21), 390 - 398. [in Russian, summaries in Polish and German]" type="journal article">Kobakhidze, 1965</bibRefCitation>
: 393
</treatmentCitation>
(M);
<treatmentCitation id="680610FC9319FF95FCC9F486FB481DD9" author="" box="[851,1206,988,1010]" page="386" pageId="15" pageNumber="104">
<bibRefCitation id="8D364B1C9319FF95FCC9F486FB7B1DD9" box="[851,1157,988,1010]" pageId="15" pageNumber="104" refId="ref19604" refString="Loksina, I. E. &amp; Golovatch, S. I. (1979) Diplopoda of the USSR fauna. Pedobiologia, 19, 381 - 389." type="journal article">Lokšina and Golovatch, 1979</bibRefCitation>
: 386
</treatmentCitation>
(M);
<treatmentCitation id="680610FC9319FF95FB68F486FF131A3B" author="" page="143" pageId="15" pageNumber="104">
<bibRefCitation id="8D364B1C9319FF95FB68F486FA661DD9" box="[1266,1432,988,1010]" pageId="15" pageNumber="104" refId="ref20114" refString="Talikadze, D. A. (1984) [On the millipede fauna (Diplopoda) of the Colchidan Province of the Caucasus]. Zoologicheskii zhurnal, 63 (1), 124 - 145 [in Russian, a summary in English]" type="journal article">Talikadze, 1984</bibRefCitation>
: 143
</treatmentCitation>
(M).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1BD65669319FF95FF0DF367FD301B95" pageId="15" pageNumber="104" type="materials_examined">
<paragraph id="E91836ED9319FF95FF0DF367FB561AE9" blockId="15.[151,1437,1085,2010]" pageId="15" pageNumber="104">
<materialsCitation id="59CF3CB09319FF95FF0DF367FA661A51" ID-GBIF-Occurrence="1919493205" collectingDate="2014-08-22" collectionCode="ZMUM" collectorName="Krasnaya Polyana, S &amp; K. Makarov" country="Russia" elevation="2000" latitude="43.774723" location="Mt Chugush" longLatPrecision="19" longitude="40.208336" municipality="Material" pageId="15" pageNumber="104" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Krasnodar">
<emphasis id="DBD3EAFF9319FF95FF0DF367FE7E1A7D" bold="true" box="[151,384,1085,1110]" pageId="15" pageNumber="104">
<collectingMunicipality id="097CAC979319FF95FF0DF367FF011A7D" box="[151,255,1085,1110]" pageId="15" pageNumber="104">Material</collectingMunicipality>
examined.
</emphasis>
<specimenCount id="FFA1FD649319FF95FE1CF364FE2A1A7D" box="[390,468,1085,1110]" pageId="15" pageNumber="104" type="male">1 male</specimenCount>
,
<specimenCount id="FFA1FD649319FF95FE44F364FDB41A7D" box="[478,586,1085,1110]" pageId="15" pageNumber="104" type="female">3 females</specimenCount>
(
<collectionCode id="8FB6AE289319FF95FDC2F364FD4B1A7D" box="[600,693,1086,1110]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="15" pageNumber="104">ZMUM</collectionCode>
),
<emphasis id="DBD3EAFF9319FF95FD52F367FC0D1A7D" bold="true" box="[712,1011,1085,1110]" pageId="15" pageNumber="104">
<collectingCountry id="91B0767D9319FF95FD52F367FCE71A7D" box="[712,793,1085,1110]" name="Russia" pageId="15" pageNumber="104">Russia</collectingCountry>
,
<collectingRegion id="2B63F80F9319FF95FCBEF367FC591A7D" box="[804,935,1085,1110]" country="Russia" name="Krasnodar" pageId="15" pageNumber="104">Krasnodar</collectingRegion>
Prov.,
</emphasis>
<locationDeviation id="F8AA2FCF9319FF95FC63F364FAB51A7D" box="[1017,1355,1085,1110]" pageId="15" pageNumber="104">
<quantity id="2E5F9B089319FF95FC63F364FBC11A7D" box="[1017,1087,1085,1110]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="104" unit="km" value="11.0">11 km</quantity>
N of
<collectorName id="4452533B9319FF95FBE7F364FAB51A7D" box="[1149,1355,1085,1110]" pageId="15" pageNumber="104">Krasnaya Polyana</collectorName>
</locationDeviation>
, S spur of
<location id="EC7860369319FF95FF2FF338FEBC1A51" LSID="urn:lsid:plazi:treatment:610E87FB9319FF8AFF0DF58DFEF319DD:EC7860369319FF95FF2FF338FEBC1A51" box="[181,322,1121,1146]" country="Russia" latitude="43.774723" longLatPrecision="19" longitude="40.208336" municipality="Material" name="Mt Chugush" pageId="15" pageNumber="104" stateProvince="Krasnodar">Mt Chugush</location>
, subalpine meadow,
<geoCoordinate id="8C93502A9319FF95FDB1F33BFD491A51" box="[555,695,1121,1146]" direction="north" orientation="latitude" pageId="15" pageNumber="104" precision="15" value="43.774723">43°46’29”N</geoCoordinate>
,
<geoCoordinate id="8C93502A9319FF95FD58F338FCB41A51" box="[706,842,1122,1146]" direction="east" orientation="longitude" pageId="15" pageNumber="104" precision="15" value="40.208336">40°12’30”E</geoCoordinate>
,
<elevation id="628AD1DE9319FF95FCCCF338FC191A51" box="[854,999,1121,1146]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="104" unit="m" value="2000.0">
<quantity id="2E5F9B089319FF95FCCCF338FC551A52" box="[854,939,1122,1146]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="104" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="8D5DE9C59319FF95FC69F338FB771A51" box="[1011,1161,1122,1146]" pageId="15" pageNumber="104" value="2014-08-22">
<date id="9D19102D9319FF95FC69F338FB771A51" box="[1011,1161,1122,1146]" pageId="15" pageNumber="104" value="2014-08-22">22.VIII.2014</date>
</collectingDate>
, leg.
<collectorName id="4452533B9319FF95FB5EF338FAB41A51" box="[1220,1354,1121,1146]" pageId="15" pageNumber="104">K. Makarov</collectorName>
and A
</materialsCitation>
.
<materialsCitation id="59CF3CB09319FF95FF0DF3DFFB561AE9" ID-GBIF-Occurrence="1919493204" collectingDate="2012-08-19" collectionCode="SMNG" collectorName="F. Walther." country="Russia" county="Matalin" elevation="1830" latitude="43.8908" location="Gora Armovka" longLatPrecision="6" longitude="40.6631" municipality="Yatyrgvarta Ridge" pageId="15" pageNumber="104" specimenCount="1" specimenCount-male="1" stateProvince="Krasnodar">
<collectingCounty id="00794E619319FF95FF0DF3DFFF0C1AB5" box="[151,242,1157,1182]" pageId="15" pageNumber="104">Matalin</collectingCounty>
;
<specimenCount id="FFA1FD649319FF95FE9BF3DCFEAF1AB5" box="[257,337,1157,1182]" pageId="15" pageNumber="104" type="male">1 male</specimenCount>
(
<collectionCode id="8FB6AE289319FF95FEFEF3DCFE451AB5" box="[356,443,1158,1182]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="15" pageNumber="104">SMNG</collectionCode>
),
<collectingMunicipality id="097CAC979319FF95FE48F3DCFD581AB5" box="[466,678,1157,1182]" pageId="15" pageNumber="104">Yatyrgvarta Ridge</collectingMunicipality>
, hut N of
<location id="EC7860369319FF95FCBDF3DCFC291AB5" LSID="urn:lsid:plazi:treatment:610E87FB9319FF8AFF0DF58DFEF319DD:EC7860369319FF95FCBDF3DCFC291AB5" box="[807,983,1157,1182]" country="Russia" county="Matalin" latitude="43.8908" longLatPrecision="6" longitude="40.6631" municipality="Yatyrgvarta Ridge" name="Gora Armovka" pageId="15" pageNumber="104" stateProvince="Krasnodar">Gora Armovka</location>
,
<geoCoordinate id="8C93502A9319FF95FC7CF3DCFB9A1AB5" box="[998,1124,1158,1182]" direction="north" orientation="latitude" pageId="15" pageNumber="104" precision="5" value="43.8908">43.8908°N</geoCoordinate>
,
<geoCoordinate id="8C93502A9319FF95FBE9F3DFFB131AB5" box="[1139,1261,1157,1182]" direction="east" orientation="longitude" pageId="15" pageNumber="104" precision="5" value="40.6631">40.6631°E</geoCoordinate>
,
<elevation id="628AD1DE9319FF95FB66F3DCFA691AB5" box="[1276,1431,1157,1182]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="15" pageNumber="104" unit="m" value="1830.0">
<quantity id="2E5F9B089319FF95FB66F3DCFAA81AB6" box="[1276,1366,1158,1182]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="15" pageNumber="104" unit="m" value="1830.0">1830 m</quantity>
a.s.l.
</elevation>
, sparse deciduous forest (
<taxonomicName id="2EA74D6E9319FF95FE36F3F3FDAF1AE9" authority=", Sorbus" authorityName="Sorbus" box="[428,593,1193,1218]" class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="104" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9319FF95FE36F3F3FDAF1AE9" box="[428,593,1193,1218]" italics="true" pageId="15" pageNumber="104">Betula, Sorbus</emphasis>
</taxonomicName>
) with hight vegetation,
<collectingDate id="8D5DE9C59319FF95FCC4F3F0FC0C1AE9" box="[862,1010,1194,1218]" pageId="15" pageNumber="104" value="2012-08-19">
<date id="9D19102D9319FF95FCC4F3F0FC0C1AE9" box="[862,1010,1194,1218]" pageId="15" pageNumber="104" value="2012-08-19">19.VIII.2012</date>
</collectingDate>
, leg.
<collectorName id="4452533B9319FF95FBB4F3F0FB561AE9" box="[1070,1192,1193,1218]" pageId="15" pageNumber="104">F. Walther.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="E91836ED9319FF95FF5DF394FB971B21" blockId="15.[151,1437,1085,2010]" pageId="15" pageNumber="104">
<materialsCitation id="59CF3CB09319FF95FF5DF394FB971B21" ID-GBIF-Occurrence="1919493221" collectingDate="1981-06" collectingDateMax="1981-07-31" collectingDateMin="1981-06" collectionCode="ZMUM" collectorName="V. Onipchenko." country="Russia" county="Teberda" elevation="2750" location="Karachaevo-Cherkessia" municipality="Mt Malaya Khatipara" pageId="15" pageNumber="104" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<specimenCount id="FFA1FD649319FF95FF5DF394FEEA1ACD" box="[199,276,1229,1254]" pageId="15" pageNumber="104" type="male">1 male</specimenCount>
,
<specimenCount id="FFA1FD649319FF95FE85F394FE751ACD" box="[287,395,1229,1254]" pageId="15" pageNumber="104" type="female">3 females</specimenCount>
(
<collectionCode id="8FB6AE289319FF95FE03F394FE081ACD" box="[409,502,1230,1254]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="15" pageNumber="104">ZMUM</collectionCode>
),
<emphasis id="DBD3EAFF9319FF95FD93F397FC741ACD" bold="true" box="[521,906,1229,1254]" pageId="15" pageNumber="104">
<collectingCountry id="91B0767D9319FF95FD93F397FDA41ACD" box="[521,602,1229,1254]" name="Russia" pageId="15" pageNumber="104">Russia</collectingCountry>
,
<location id="EC7860369319FF95FDFFF397FC7B1ACD" LSID="urn:lsid:plazi:treatment:610E87FB9319FF8AFF0DF58DFEF319DD:EC7860369319FF95FDFFF397FC7B1ACD" box="[613,901,1229,1254]" country="Russia" county="Teberda" municipality="Mt Malaya Khatipara" name="Karachaevo-Cherkessia" pageId="15" pageNumber="104">Karachaevo-Cherkessia</location>
,
</emphasis>
<location id="EC7860369319FF95FC0BF397FB5F1ACE" LSID="urn:lsid:plazi:treatment:610E87FB9319FF8AFF0DF58DFEF319DD:EC7860369319FF95FC0BF397FB5F1ACE" box="[913,1185,1229,1254]" country="Russia" county="Teberda" municipality="Mt Malaya Khatipara" name="Teberda Nature Reserve" pageId="15" pageNumber="104">Teberda Nature Reserve</location>
,
<collectingMunicipality id="097CAC979319FF95FB36F394FA651ACD" box="[1196,1435,1229,1254]" pageId="15" pageNumber="104">Mt Malaya Khatipara</collectingMunicipality>
above
<collectingCounty id="00794E619319FF95FF7BF3ABFEC01B21" box="[225,318,1265,1290]" pageId="15" pageNumber="104">Teberda</collectingCounty>
, alpine meadow,
<elevation id="628AD1DE9319FF95FE65F3A8FD291B21" box="[511,727,1265,1290]" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="15" pageNumber="104" unit="m" value="2750.0" valueMax="2800.0" valueMin="2700.0">
<quantity id="2E5F9B089319FF95FE65F3A8FD641B22" box="[511,666,1266,1290]" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="15" pageNumber="104" unit="m" value="2750.0" valueMax="2800.0" valueMin="2700.0">2700–2800 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="8D5DE9C59319FF95FD79F3A8FC801B21" box="[739,894,1266,1290]" pageId="15" pageNumber="104" value="1981-06" valueMax="1981-07-31" valueMin="1981-06-00">
<date id="9D19102D9319FF95FD79F3A8FC801B21" box="[739,894,1266,1290]" pageId="15" pageNumber="104" value="1981-06" valueMax="1981-07-31" valueMin="1981-06-00">VI.–VII.1981</date>
</collectingDate>
, leg.
<collectorName id="4452533B9319FF95FC20F3A8FB971B21" box="[954,1129,1265,1290]" pageId="15" pageNumber="104">V. Onipchenko.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="E91836ED9319FF95FF5DF24CFEF01B5D" blockId="15.[151,1437,1085,2010]" pageId="15" pageNumber="104">
<materialsCitation id="59CF3CB09319FF95FF5DF24CFEE01B79" ID-GBIF-Occurrence="1919493226" collectingDate="2014-08-28" collectionCode="ZMUM" collectorName="I. Turbanov" county="Arabika Massif" location="Trekhusa Grotto" municipality="Kamennyi Klad Mt. Range" pageId="15" pageNumber="104" specimenCount="1" specimenCount-male="1" stateProvince="Abkhazia">
<specimenCount id="FFA1FD649319FF95FF5DF24CFEED1B05" box="[199,275,1301,1326]" pageId="15" pageNumber="104" type="male">1 male</specimenCount>
(
<collectionCode id="8FB6AE289319FF95FEBBF24CFE801B05" box="[289,382,1302,1326]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="15" pageNumber="104">ZMUM</collectionCode>
),
<emphasis id="DBD3EAFF9319FF95FE08F24FFDF21B05" bold="true" box="[402,524,1301,1326]" pageId="15" pageNumber="104">
<collectingRegion id="2B63F80F9319FF95FE08F24FFDF91B05" box="[402,519,1301,1326]" country="Georgia" name="Abkhazia" pageId="15" pageNumber="104">Abkhazia</collectingRegion>
,
</emphasis>
<collectingCounty id="00794E619319FF95FD89F24FFD3D1B05" box="[531,707,1301,1326]" pageId="15" pageNumber="104">Arabika Massif</collectingCounty>
,
<collectingMunicipality id="097CAC979319FF95FD55F24FFBFF1B05" box="[719,1025,1301,1326]" pageId="15" pageNumber="104">Kamennyi Klad Mt. Range</collectingMunicipality>
,
<location id="EC7860369319FF95FB97F24FFB391B05" LSID="urn:lsid:plazi:treatment:610E87FB9319FF8AFF0DF58DFEF319DD:EC7860369319FF95FB97F24FFB391B05" box="[1037,1223,1301,1326]" county="Arabika Massif" municipality="Kamennyi Klad Mt. Range" name="Trekhusa Grotto" pageId="15" pageNumber="104" stateProvince="Abkhazia">Trekhusa Grotto</location>
,
<collectingDate id="8D5DE9C59319FF95FB48F24CFA991B05" box="[1234,1383,1302,1326]" pageId="15" pageNumber="104" value="2014-08-28">
<date id="9D19102D9319FF95FB48F24CFA991B05" box="[1234,1383,1302,1326]" pageId="15" pageNumber="104" value="2014-08-28">28.VIII.2014</date>
</collectingDate>
, leg.
<collectorName id="4452533B9319FF95FF0DF260FEE01B79" box="[151,286,1337,1362]" pageId="15" pageNumber="104">I. Turbanov</collectorName>
</materialsCitation>
;
<materialsCitation id="59CF3CB09319FF95FEB1F260FEF01B5D" ID-GBIF-Occurrence="1919493224" collectingDate="1984-08" collectionCode="ZMUM" collectorName="E. Kvavadze." elevation="1750" location="Marukh Pass" municipality="Adante Valley" pageId="15" pageNumber="104" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Abkhazia">
<specimenCount id="FFA1FD649319FF95FEB1F260FE871B79" box="[299,377,1337,1362]" pageId="15" pageNumber="104" type="male">1 male</specimenCount>
,
<specimenCount id="FFA1FD649319FF95FE1FF260FE161B79" box="[389,488,1337,1362]" pageId="15" pageNumber="104" type="female">1 female</specimenCount>
(
<collectionCode id="8FB6AE289319FF95FE6DF260FDAA1B79" box="[503,596,1338,1362]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="15" pageNumber="104">ZMUM</collectionCode>
),
<collectingMunicipality id="097CAC979319FF95FDF2F263FCF41B79" box="[616,778,1337,1362]" pageId="15" pageNumber="104">Adante Valley</collectingMunicipality>
, near
<location id="EC7860369319FF95FCD7F263FC1C1B7A" LSID="urn:lsid:plazi:treatment:610E87FB9319FF8AFF0DF58DFEF319DD:EC7860369319FF95FCD7F263FC1C1B7A" box="[845,994,1337,1362]" municipality="Adante Valley" name="Marukh Pass" pageId="15" pageNumber="104" stateProvince="Abkhazia">Marukh Pass</location>
,
<elevation id="628AD1DE9319FF95FC75F260FB361B79" box="[1007,1224,1337,1362]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="15" pageNumber="104" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">
<quantity id="2E5F9B089319FF95FC75F260FB741B7A" box="[1007,1162,1338,1362]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="15" pageNumber="104" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">1700–1800 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="8D5DE9C59319FF95FB4FF260FAB81B79" box="[1237,1350,1338,1362]" pageId="15" pageNumber="104" value="1984-08">
<date id="9D19102D9319FF95FB4FF260FAB81B79" box="[1237,1350,1338,1362]" pageId="15" pageNumber="104" value="1984-08">VIII.1984</date>
</collectingDate>
, leg.
<collectorName id="4452533B9319FF95FA1EF260FEF01B5D" pageId="15" pageNumber="104">E. Kvavadze.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="E91836ED9319FF95FF5DF2D8FD301B95" blockId="15.[151,1437,1085,2010]" pageId="15" pageNumber="104">
3 males, 5 females (ZMUM),
<emphasis id="DBD3EAFF9319FF95FD8DF2DBFD801BB1" bold="true" box="[535,638,1409,1434]" pageId="15" pageNumber="104">Georgia,</emphasis>
Mestia,
<quantity id="2E5F9B089319FF95FD7AF2DBFCCB1BB2" box="[736,821,1409,1434]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="104" unit="m" value="1500.0">1500 m</quantity>
a.s.l.,
<taxonomicName id="2EA74D6E9319FF95FCE4F2DBFC391BB1" box="[894,967,1409,1434]" class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="104" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9319FF95FCE4F2DBFC391BB1" box="[894,967,1409,1434]" italics="true" pageId="15" pageNumber="104">Betula</emphasis>
</taxonomicName>
and
<taxonomicName id="2EA74D6E9319FF95FC64F2DBFB5D1BB1" box="[1022,1187,1409,1434]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="15" pageNumber="104" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9319FF95FC64F2DBFB5D1BB1" box="[1022,1187,1409,1434]" italics="true" pageId="15" pageNumber="104">Rhododendron</emphasis>
</taxonomicName>
on moraine, litter and under stones,
<date id="9D19102D9319FF95FEA8F2FFFE061B95" box="[306,504,1445,1470]" pageId="15" pageNumber="104" value="1986-09-05" valueMax="1986-09-16" valueMin="1986-09-05">5 and 16.IX.1986</date>
, leg. S. Golovatch.
</paragraph>
</subSubSection>
<subSubSection id="A1BD65669319FF8AFF5DF293FCAE1935" lastPageId="16" lastPageNumber="105" pageId="15" pageNumber="104" type="diagnosis">
<paragraph id="E91836ED9319FF95FF5DF293FBEA1891" blockId="15.[151,1437,1085,2010]" pageId="15" pageNumber="104">
<emphasis id="DBD3EAFF9319FF95FF5DF293FEBF1BC9" bold="true" box="[199,321,1481,1506]" pageId="15" pageNumber="104">Diagnosis.</emphasis>
Differs from other species of the genus by the following combination of characters. Main process (
<emphasis id="DBD3EAFF9319FF95FF3AF2B7FF39182D" bold="true" box="[160,199,1517,1542]" pageId="15" pageNumber="104">mp</emphasis>
) of male coxa 2 long, with or without a small apical bulb, but always with a small caudal tooth or uncus at apex (
<figureCitation id="719C2A689319FF95FF44F14BFE9C1801" box="[222,354,1553,1578]" captionStart="FIGURE 6" captionStartId="10.[151,250,1294,1316]" captionTargetBox="[340,1248,199,1261]" captionTargetId="figure@10.[340,1248,193,1273]" captionTargetPageId="10" captionText="FIGURE 6. Male legs 2 in Julus lignaui Verhoeff, 1910 (A–D) and J. lindholmi Lohmander, 1936 (E–G), caudal views. A: from Krasnaya Polyana, Krasnodar Province. B: from Teberda, Karachaevo-Cherkessia. C: from Mestia, Georgia. D: from Teberda, Karachaevo-Cherkessia. E: from Svanetia, Georgia. F: from near Teberda, Karachaevo-Cherkessia. G: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (A–F), not to scale (G)." httpUri="https://zenodo.org/record/1459884/files/figure.png" pageId="15" pageNumber="104">Figs 6A–D</figureCitation>
). Gonopodal promere (
<emphasis id="DBD3EAFF9319FF95FDF5F14BFD751801" bold="true" box="[623,651,1553,1578]" pageId="15" pageNumber="104">pr</emphasis>
) narrow, round at apex, with an elongated lateral lamella (
<emphasis id="DBD3EAFF9319FF95FAA3F14BFAB11801" bold="true" box="[1337,1359,1553,1578]" pageId="15" pageNumber="104">la</emphasis>
) (
<figureCitation id="719C2A689319FF95FAF0F14BFE781865" captionStart="FIGURE 11" captionStartId="16.[151,250,1572,1594]" captionTargetBox="[350,1231,514,1553]" captionTargetId="figure@16.[340,1248,507,1563]" captionTargetPageId="16" captionText="FIGURE 11. Male gonopods in Julus lignaui Verhoeff, 1910 (A–B from Krasnaya Polyana, Krasnodar Province; C–D from Mestia, Georgia; E–H from Teberda, Karachaevo-Cherkessia). A: promere with flagella, caudal view. B, D, H: left gonopods, mesal view. C, E, G: promere, caudal view. F: right gonopod, mesal view. Scale bar: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459894/files/figure.png" pageId="15" pageNumber="104">Figs 11A, 11C, 11E, 11G</figureCitation>
). Flagella (
<emphasis id="DBD3EAFF9319FF95FD90F16FFDE51865" bold="true" box="[522,539,1589,1614]" pageId="15" pageNumber="104">fl</emphasis>
) wide basally, gradually attenuating towards apex (
<figureCitation id="719C2A689319FF95FBE8F16FFF341859" captionStart="FIGURE 11" captionStartId="16.[151,250,1572,1594]" captionTargetBox="[350,1231,514,1553]" captionTargetId="figure@16.[340,1248,507,1563]" captionTargetPageId="16" captionText="FIGURE 11. Male gonopods in Julus lignaui Verhoeff, 1910 (A–B from Krasnaya Polyana, Krasnodar Province; C–D from Mestia, Georgia; E–H from Teberda, Karachaevo-Cherkessia). A: promere with flagella, caudal view. B, D, H: left gonopods, mesal view. C, E, G: promere, caudal view. F: right gonopod, mesal view. Scale bar: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459894/files/figure.png" pageId="15" pageNumber="104">Figs 11A, 11B, 11D, 11F, 11H</figureCitation>
). Mesomere (
<emphasis id="DBD3EAFF9319FF95FEF3F103FE751859" bold="true" box="[361,395,1625,1650]" pageId="15" pageNumber="104">ms</emphasis>
) rounded at apex (
<figureCitation id="719C2A689319FF95FDFDF103FC721859" box="[615,908,1625,1650]" captionStart="FIGURE 11" captionStartId="16.[151,250,1572,1594]" captionTargetBox="[350,1231,514,1553]" captionTargetId="figure@16.[340,1248,507,1563]" captionTargetPageId="16" captionText="FIGURE 11. Male gonopods in Julus lignaui Verhoeff, 1910 (A–B from Krasnaya Polyana, Krasnodar Province; C–D from Mestia, Georgia; E–H from Teberda, Karachaevo-Cherkessia). A: promere with flagella, caudal view. B, D, H: left gonopods, mesal view. C, E, G: promere, caudal view. F: right gonopod, mesal view. Scale bar: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459894/files/figure.png" pageId="15" pageNumber="104">Figs 11B, 11D, 11F, 11H</figureCitation>
). Opisthomere (
<emphasis id="DBD3EAFF9319FF95FBDFF103FB9C1859" bold="true" box="[1093,1122,1625,1650]" pageId="15" pageNumber="104">op</emphasis>
) and lateral outgrowth (
<emphasis id="DBD3EAFF9319FF95FAE7F103FA6D1859" bold="true" box="[1405,1427,1625,1650]" pageId="15" pageNumber="104">lo</emphasis>
) sharpened on top (
<figureCitation id="719C2A689319FF95FEE3F127FDD218BD" box="[377,556,1661,1686]" captionStart="FIGURE 11" captionStartId="16.[151,250,1572,1594]" captionTargetBox="[350,1231,514,1553]" captionTargetId="figure@16.[340,1248,507,1563]" captionTargetPageId="16" captionText="FIGURE 11. Male gonopods in Julus lignaui Verhoeff, 1910 (A–B from Krasnaya Polyana, Krasnodar Province; C–D from Mestia, Georgia; E–H from Teberda, Karachaevo-Cherkessia). A: promere with flagella, caudal view. B, D, H: left gonopods, mesal view. C, E, G: promere, caudal view. F: right gonopod, mesal view. Scale bar: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459894/files/figure.png" pageId="15" pageNumber="104">Figs 11B, 11D</figureCitation>
,
<figureCitation id="719C2A689319FF95FDA4F124FD8E18BD" box="[574,624,1662,1686]" captionStart="FIGURE 14" captionStartId="20.[151,250,1014,1036]" captionTargetBox="[340,1248,373,993]" captionTargetId="figure@20.[340,1248,373,993]" captionTargetPageId="20" captionText="FIGURE 14. Mesomere and opisthomere in Caucasian species of Julus, oral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936 G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. Scale bars: 0.3 mm. Abbreviations: fl, flagellum; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere." httpUri="https://zenodo.org/record/1459900/files/figure.png" pageId="15" pageNumber="104">14E</figureCitation>
). Striations on metazonae regular and rough (
<figureCitation id="719C2A689319FF95FB01F127FAF618BD" box="[1179,1288,1661,1686]" captionStart="FIGURE 16" captionStartId="21.[151,249,1286,1308]" captionTargetBox="[340,1247,583,1264]" captionTargetId="figure@21.[340,1247,583,1264]" captionTargetPageId="21" captionText="FIGURE 16. Striation on metazonites in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459904/files/figure.png" pageId="15" pageNumber="104">Fig. 16E</figureCitation>
). Shape and arrangement of ampullae of female vulvae much like in
<taxonomicName id="2EA74D6E9319FF95FC96F1F8FC6E1891" authorityName="Lohmander" authorityYear="1936" box="[780,912,1697,1722]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="15" pageNumber="104" phylum="Arthropoda" rank="species" species="colchicus">
<emphasis id="DBD3EAFF9319FF95FC96F1F8FC6E1891" box="[780,912,1697,1722]" italics="true" pageId="15" pageNumber="104">J. colchicus</emphasis>
</taxonomicName>
(
<figureCitation id="719C2A689319FF95FC05F1FBFBF61891" box="[927,1032,1697,1722]" captionStart="FIGURE 18" captionStartId="22.[151,249,1951,1973]" captionTargetBox="[340,1248,1166,1930]" captionTargetId="figure@22.[340,1248,1166,1930]" captionTargetPageId="22" captionText="FIGURE 18. Female vulvae and legs 2 in Caucasian species of Julus. A: J. lignaui Verhoeff, 1910. B: J. lindholmi Lohmander, 1936. C: J. subalpinus Lohmander, 1936. D: J. dagestanus sp. n. Abbreviations: vu, vulva; op, operculum; bu, bursa; am1 and am2, ampulla 1 and ampulla 2; l2, coxa and prefemur of leg 2. Scale bar: 0.3 mm." httpUri="https://zenodo.org/record/1459908/files/figure.png" pageId="15" pageNumber="104">Fig. 18A</figureCitation>
).
</paragraph>
<paragraph id="E91836ED9319FF95FF5DF19FFB56190D" blockId="15.[151,1437,1085,2010]" pageId="15" pageNumber="104">
<emphasis id="DBD3EAFF9319FF95FF5DF19FFE6018F5" bold="true" box="[199,414,1733,1758]" pageId="15" pageNumber="104">Descriptive notes.</emphasis>
Length
<quantity id="2E5F9B089319FF95FD99F19CFD7C18F6" box="[515,642,1734,1758]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="15" pageNumber="104" unit="mm" value="25.0" valueMax="27.0" valueMin="23.0">23–27 mm</quantity>
, width
<quantity id="2E5F9B089319FF95FD47F19FFC9418F6" box="[733,874,1733,1758]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.5" metricValueMin="1.3" pageId="15" pageNumber="104" unit="mm" value="1.9" valueMax="2.5" valueMin="1.3">1.3–2.5 mm</quantity>
. Number of segments from 44+1+T to 50+1+T. Body grey with a thin, black, dorsal, axial line, venter lighter; legs light brown. Antennae and anal valves brown. Eye patches composed of
<emphasis id="DBD3EAFF9319FF95FE21F054FE28190E" box="[443,470,1806,1829]" italics="true" pageId="15" pageNumber="104">ca</emphasis>
45 ocelli. Striations on metazonae regular and rough (
<figureCitation id="719C2A689319FF95FBACF057FB65190D" box="[1078,1179,1805,1830]" captionStart="FIGURE 16" captionStartId="21.[151,249,1286,1308]" captionTargetBox="[340,1247,583,1264]" captionTargetId="figure@21.[340,1247,583,1264]" captionTargetPageId="21" captionText="FIGURE 16. Striation on metazonites in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459904/files/figure.png" pageId="15" pageNumber="104">Fig. 16E</figureCitation>
).
</paragraph>
<paragraph id="E91836ED9319FF95FF5DF06BFCE219B9" blockId="15.[151,1437,1085,2010]" pageId="15" pageNumber="104">
All other characters as in
<taxonomicName id="2EA74D6E9319FF95FE72F068FD7F1961" authorityName="Evsyukov" authorityYear="2018" box="[488,641,1841,1866]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="15" pageNumber="104" phylum="Arthropoda" rank="species" species="alexandrae">
<emphasis id="DBD3EAFF9319FF95FE72F068FD7F1961" box="[488,641,1841,1866]" italics="true" pageId="15" pageNumber="104">J. alexandrae</emphasis>
</taxonomicName>
, except as follows. Antennae
<emphasis id="DBD3EAFF9319FF95FC4EF068FBE41962" box="[980,1050,1842,1865]" italics="true" pageId="15" pageNumber="104">in situ</emphasis>
reaching back to body segment 3. Antennomeres 5 and 6 each with a corolla of bacilliform sensilla (
<figureCitation id="719C2A689319FF95FC1EF00FFC241945" box="[900,986,1877,1902]" captionStart="FIGURE 1" captionStartId="4.[151,249,1820,1842]" captionTargetBox="[344,1236,854,1775]" captionTargetId="figure@4.[340,1248,841,1798]" captionTargetPageId="4" captionText="FIGURE 1. Antennae in Caucasian species of Julus, lateral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. dagestanus sp. n. H: J. khostensis sp. n. Scale bar: 0.3 mm." httpUri="https://zenodo.org/record/1459874/files/figure.png" pageId="15" pageNumber="104">Fig. 1E</figureCitation>
). Each stipes of gnathochilarium with a large group of setae parabasally below mentum (
<figureCitation id="719C2A689319FF95FD22F023FCF119B9" box="[696,783,1913,1938]" captionStart="FIGURE 2" captionStartId="5.[151,249,1601,1623]" captionTargetBox="[340,1248,985,1580]" captionTargetId="figure@5.[340,1248,985,1580]" captionTargetPageId="5" captionText="FIGURE 2. Gnathochilaria in Caucasian species of Julus, ventral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459876/files/figure.png" pageId="15" pageNumber="104">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="E91836ED9319FF8AFF5DF0C7FEC71F6B" blockId="15.[151,1437,1085,2010]" lastBlockId="16.[151,1436,151,464]" lastPageId="16" lastPageNumber="105" pageId="15" pageNumber="104">
Male leg-pair 1 (
<figureCitation id="719C2A689319FF95FE0CF0C7FDE8199D" box="[406,534,1949,1974]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (B–D), J. jedryczkowskii Golovatch, 1981 (E–F), J. kubanus Verhoeff, 1921 (G–I), J. lignaui Verhoeff, 1910 (J–M), J. lindholmi Lohmander, 1936 (N– P), J. subalpinus Lohmander, 1936 (Q–R), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views B–O; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (A–C, E, G, H, J–O, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="15" pageNumber="104">Figs 3J–M</figureCitation>
) usually with a well-developed suture between coxa (
<emphasis id="DBD3EAFF9319FF95FB3FF0C7FB41199D" bold="true" box="[1189,1215,1949,1974]" pageId="15" pageNumber="104">cx</emphasis>
) and 1-segmented telopodite remnant (
<emphasis id="DBD3EAFF9319FF95FEE6F09BFE6C19F1" bold="true" box="[380,402,1985,2010]" pageId="15" pageNumber="104">te</emphasis>
), but sometimes the suture is absent (
<figureCitation id="719C2A689319FF95FCDBF09BFC3819F1" box="[833,966,1985,2010]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (B–D), J. jedryczkowskii Golovatch, 1981 (E–F), J. kubanus Verhoeff, 1921 (G–I), J. lignaui Verhoeff, 1910 (J–M), J. lindholmi Lohmander, 1936 (N– P), J. subalpinus Lohmander, 1936 (Q–R), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views B–O; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (A–C, E, G, H, J–O, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="15" pageNumber="104">Figs 3J, 3L</figureCitation>
);
<emphasis id="DBD3EAFF9319FF95FC46F09BFC0919F1" bold="true" box="[988,1015,1985,2010]" pageId="15" pageNumber="104">cx</emphasis>
with (
<figureCitation id="719C2A689319FF95FBDAF09BFB4D19F1" box="[1088,1203,1985,2010]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (B–D), J. jedryczkowskii Golovatch, 1981 (E–F), J. kubanus Verhoeff, 1921 (G–I), J. lignaui Verhoeff, 1910 (J–M), J. lindholmi Lohmander, 1936 (N– P), J. subalpinus Lohmander, 1936 (Q–R), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views B–O; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (A–C, E, G, H, J–O, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="15" pageNumber="104">Figs 3J–L</figureCitation>
) or without a lateral process (
<figureCitation id="719C2A689306FF8AFF66F7CDFEA21E9B" box="[252,348,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (B–D), J. jedryczkowskii Golovatch, 1981 (E–F), J. kubanus Verhoeff, 1921 (G–I), J. lignaui Verhoeff, 1910 (J–M), J. lindholmi Lohmander, 1936 (N– P), J. subalpinus Lohmander, 1936 (Q–R), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views B–O; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (A–C, E, G, H, J–O, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="16" pageNumber="105">Fig. 3M</figureCitation>
);
<emphasis id="DBD3EAFF9306FF8AFEE9F7CDFE771E9B" bold="true" box="[371,393,151,176]" pageId="16" pageNumber="105">te</emphasis>
round, with a group of thin setae. Male leg-pair 2 with three coxal processes (
<figureCitation id="719C2A689306FF8AFA97F7CDFA6E1E9B" box="[1293,1424,151,176]" captionStart="FIGURE 6" captionStartId="10.[151,250,1294,1316]" captionTargetBox="[340,1248,199,1261]" captionTargetId="figure@10.[340,1248,193,1273]" captionTargetPageId="10" captionText="FIGURE 6. Male legs 2 in Julus lignaui Verhoeff, 1910 (A–D) and J. lindholmi Lohmander, 1936 (E–G), caudal views. A: from Krasnaya Polyana, Krasnodar Province. B: from Teberda, Karachaevo-Cherkessia. C: from Mestia, Georgia. D: from Teberda, Karachaevo-Cherkessia. E: from Svanetia, Georgia. F: from near Teberda, Karachaevo-Cherkessia. G: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (A–F), not to scale (G)." httpUri="https://zenodo.org/record/1459884/files/figure.png" pageId="16" pageNumber="105">Figs 6A–D</figureCitation>
). Anterolateral process (
<emphasis id="DBD3EAFF9306FF8AFE0CF7E6FE451EFE" bold="true" box="[406,443,188,213]" pageId="16" pageNumber="105">alp</emphasis>
) short, rounded at apex. Main process (
<emphasis id="DBD3EAFF9306FF8AFCE2F7E6FC611EFE" bold="true" box="[888,927,188,213]" pageId="16" pageNumber="105">mp</emphasis>
) long, usually with a small bulb and a caudal dentiform outgrowth at apex (
<figureCitation id="719C2A689306FF8AFE68F785FD7B1ED3" box="[498,645,223,248]" captionStart="FIGURE 6" captionStartId="10.[151,250,1294,1316]" captionTargetBox="[340,1248,199,1261]" captionTargetId="figure@10.[340,1248,193,1273]" captionTargetPageId="10" captionText="FIGURE 6. Male legs 2 in Julus lignaui Verhoeff, 1910 (A–D) and J. lindholmi Lohmander, 1936 (E–G), caudal views. A: from Krasnaya Polyana, Krasnodar Province. B: from Teberda, Karachaevo-Cherkessia. C: from Mestia, Georgia. D: from Teberda, Karachaevo-Cherkessia. E: from Svanetia, Georgia. F: from near Teberda, Karachaevo-Cherkessia. G: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (A–F), not to scale (G)." httpUri="https://zenodo.org/record/1459884/files/figure.png" pageId="16" pageNumber="105">Figs 6A, 6C</figureCitation>
), but sometimes slender throughout and with a small, apical, caudal uncus (
<figureCitation id="719C2A689306FF8AFF70F65EFEBD1F36" box="[234,323,260,285]" captionStart="FIGURE 6" captionStartId="10.[151,250,1294,1316]" captionTargetBox="[340,1248,199,1261]" captionTargetId="figure@10.[340,1248,193,1273]" captionTargetPageId="10" captionText="FIGURE 6. Male legs 2 in Julus lignaui Verhoeff, 1910 (A–D) and J. lindholmi Lohmander, 1936 (E–G), caudal views. A: from Krasnaya Polyana, Krasnodar Province. B: from Teberda, Karachaevo-Cherkessia. C: from Mestia, Georgia. D: from Teberda, Karachaevo-Cherkessia. E: from Svanetia, Georgia. F: from near Teberda, Karachaevo-Cherkessia. G: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (A–F), not to scale (G)." httpUri="https://zenodo.org/record/1459884/files/figure.png" pageId="16" pageNumber="105">Fig. 6B</figureCitation>
). Posterior process (
<emphasis id="DBD3EAFF9306FF8AFDB4F65EFDB31F36" bold="true" box="[558,589,260,285]" pageId="16" pageNumber="105">pp</emphasis>
) typically with three setae (
<figureCitation id="719C2A689306FF8AFC0AF65EFC151F36" box="[912,1003,260,285]" captionStart="FIGURE 6" captionStartId="10.[151,250,1294,1316]" captionTargetBox="[340,1248,199,1261]" captionTargetId="figure@10.[340,1248,193,1273]" captionTargetPageId="10" captionText="FIGURE 6. Male legs 2 in Julus lignaui Verhoeff, 1910 (A–D) and J. lindholmi Lohmander, 1936 (E–G), caudal views. A: from Krasnaya Polyana, Krasnodar Province. B: from Teberda, Karachaevo-Cherkessia. C: from Mestia, Georgia. D: from Teberda, Karachaevo-Cherkessia. E: from Svanetia, Georgia. F: from near Teberda, Karachaevo-Cherkessia. G: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (A–F), not to scale (G)." httpUri="https://zenodo.org/record/1459884/files/figure.png" pageId="16" pageNumber="105">Fig. 6C</figureCitation>
). Penes (
<emphasis id="DBD3EAFF9306FF8AFBCEF65EFB9D1F36" bold="true" box="[1108,1123,260,285]" pageId="16" pageNumber="105">p</emphasis>
) short, emarginate apically (
<figureCitation id="719C2A689306FF8AFF05F67DFED21F6B" box="[159,300,295,320]" captionStart="FIGURE 6" captionStartId="10.[151,250,1294,1316]" captionTargetBox="[340,1248,199,1261]" captionTargetId="figure@10.[340,1248,193,1273]" captionTargetPageId="10" captionText="FIGURE 6. Male legs 2 in Julus lignaui Verhoeff, 1910 (A–D) and J. lindholmi Lohmander, 1936 (E–G), caudal views. A: from Krasnaya Polyana, Krasnodar Province. B: from Teberda, Karachaevo-Cherkessia. C: from Mestia, Georgia. D: from Teberda, Karachaevo-Cherkessia. E: from Svanetia, Georgia. F: from near Teberda, Karachaevo-Cherkessia. G: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (A–F), not to scale (G)." httpUri="https://zenodo.org/record/1459884/files/figure.png" pageId="16" pageNumber="105">Figs 6A, 6C</figureCitation>
).
</paragraph>
<paragraph id="E91836ED9306FF8AFF5DF617FB461FA3" blockId="16.[151,1436,151,464]" pageId="16" pageNumber="105">
Promere (
<emphasis id="DBD3EAFF9306FF8AFEA0F616FEA81F4E" bold="true" box="[314,342,332,357]" pageId="16" pageNumber="105">pr</emphasis>
) narrow, rounded at apex, with an elongated lateral lamella (
<emphasis id="DBD3EAFF9306FF8AFBB7F616FBBD1F4E" bold="true" box="[1069,1091,332,357]" pageId="16" pageNumber="105">la</emphasis>
) (
<figureCitation id="719C2A689306FF8AFBC5F616FA6E1F4F" box="[1119,1424,332,357]" captionStart="FIGURE 11" captionStartId="16.[151,250,1572,1594]" captionTargetBox="[350,1231,514,1553]" captionTargetId="figure@16.[340,1248,507,1563]" captionTargetPageId="16" captionText="FIGURE 11. Male gonopods in Julus lignaui Verhoeff, 1910 (A–B from Krasnaya Polyana, Krasnodar Province; C–D from Mestia, Georgia; E–H from Teberda, Karachaevo-Cherkessia). A: promere with flagella, caudal view. B, D, H: left gonopods, mesal view. C, E, G: promere, caudal view. F: right gonopod, mesal view. Scale bar: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459894/files/figure.png" pageId="16" pageNumber="105">Figs 11A, 11C, 11E, 11G</figureCitation>
). Flagella (
<emphasis id="DBD3EAFF9306FF8AFE9BF635FEEC1FA3" bold="true" box="[257,274,367,392]" pageId="16" pageNumber="105">fl</emphasis>
) wide basally, gradually attenuating towards apex (
<figureCitation id="719C2A689306FF8AFCC9F635FB551FA3" box="[851,1195,367,392]" captionStart="FIGURE 11" captionStartId="16.[151,250,1572,1594]" captionTargetBox="[350,1231,514,1553]" captionTargetId="figure@16.[340,1248,507,1563]" captionTargetPageId="16" captionText="FIGURE 11. Male gonopods in Julus lignaui Verhoeff, 1910 (A–B from Krasnaya Polyana, Krasnodar Province; C–D from Mestia, Georgia; E–H from Teberda, Karachaevo-Cherkessia). A: promere with flagella, caudal view. B, D, H: left gonopods, mesal view. C, E, G: promere, caudal view. F: right gonopod, mesal view. Scale bar: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459894/files/figure.png" pageId="16" pageNumber="105">Figs 11A, 11B, 11D, 11F, 11H</figureCitation>
).
</paragraph>
<paragraph id="E91836ED9306FF8AFF5DF6CFFE471FFB" blockId="16.[151,1436,151,464]" pageId="16" pageNumber="105">
Mesomere (
<emphasis id="DBD3EAFF9306FF8AFED7F6CEFE911F86" bold="true" box="[333,367,404,429]" pageId="16" pageNumber="105">ms</emphasis>
) rounded at apex (
<figureCitation id="719C2A689306FF8AFDDAF6CEFCE11F87" box="[576,799,404,429]" captionStart="FIGURE 11" captionStartId="16.[151,250,1572,1594]" captionTargetBox="[350,1231,514,1553]" captionTargetId="figure@16.[340,1248,507,1563]" captionTargetPageId="16" captionText="FIGURE 11. Male gonopods in Julus lignaui Verhoeff, 1910 (A–B from Krasnaya Polyana, Krasnodar Province; C–D from Mestia, Georgia; E–H from Teberda, Karachaevo-Cherkessia). A: promere with flagella, caudal view. B, D, H: left gonopods, mesal view. C, E, G: promere, caudal view. F: right gonopod, mesal view. Scale bar: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459894/files/figure.png" pageId="16" pageNumber="105">Figs 11B, 11D, 11F</figureCitation>
). Opisthomere (
<emphasis id="DBD3EAFF9306FF8AFC49F6CEFC0E1F86" bold="true" box="[979,1008,404,429]" pageId="16" pageNumber="105">op</emphasis>
) and lateral outgrowth (
<emphasis id="DBD3EAFF9306FF8AFB65F6CEFAEB1F86" bold="true" box="[1279,1301,404,429]" pageId="16" pageNumber="105">lo</emphasis>
) pointed on top (
<figureCitation id="719C2A689306FF8AFF53F6EDFE911FFB" box="[201,367,439,464]" captionStart="FIGURE 11" captionStartId="16.[151,250,1572,1594]" captionTargetBox="[350,1231,514,1553]" captionTargetId="figure@16.[340,1248,507,1563]" captionTargetPageId="16" captionText="FIGURE 11. Male gonopods in Julus lignaui Verhoeff, 1910 (A–B from Krasnaya Polyana, Krasnodar Province; C–D from Mestia, Georgia; E–H from Teberda, Karachaevo-Cherkessia). A: promere with flagella, caudal view. B, D, H: left gonopods, mesal view. C, E, G: promere, caudal view. F: right gonopod, mesal view. Scale bar: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459894/files/figure.png" pageId="16" pageNumber="105">Figs 11B, 11D</figureCitation>
,
<figureCitation id="719C2A689306FF8AFEE1F6E2FE521FFB" box="[379,428,440,464]" captionStart="FIGURE 14" captionStartId="20.[151,250,1014,1036]" captionTargetBox="[340,1248,373,993]" captionTargetId="figure@20.[340,1248,373,993]" captionTargetPageId="20" captionText="FIGURE 14. Mesomere and opisthomere in Caucasian species of Julus, oral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936 G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. Scale bars: 0.3 mm. Abbreviations: fl, flagellum; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere." httpUri="https://zenodo.org/record/1459900/files/figure.png" pageId="16" pageNumber="105">14E</figureCitation>
).
</paragraph>
<caption id="BDD866659306FF8AFF0DF17EFB4B18BE" httpUri="https://zenodo.org/record/1459894/files/figure.png" pageId="16" pageNumber="105" startId="16.[151,250,1572,1594]" targetBox="[350,1231,514,1553]" targetPageId="16">
<paragraph id="E91836ED9306FF8AFF0DF17EFB4B18BE" blockId="16.[151,1436,1572,1685]" pageId="16" pageNumber="105">
<emphasis id="DBD3EAFF9306FF8AFF0DF17EFEE11811" bold="true" box="[151,287,1572,1594]" pageId="16" pageNumber="105">FIGURE 11.</emphasis>
Male gonopods in
<taxonomicName id="2EA74D6E9306FF8AFE73F17EFCF11811" authority="Verhoeff, 1910" authorityName="Verhoeff" authorityYear="1910" box="[489,783,1572,1594]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="16" pageNumber="105" phylum="Arthropoda" rank="species" species="lignaui">
<emphasis id="DBD3EAFF9306FF8AFE73F17EFD901811" box="[489,622,1572,1594]" italics="true" pageId="16" pageNumber="105">Julus lignaui</emphasis>
Verhoeff, 1910
</taxonomicName>
(
<emphasis id="DBD3EAFF9306FF8AFC85F17EFCB11811" bold="true" box="[799,847,1572,1594]" pageId="16" pageNumber="105">A–B</emphasis>
from Krasnaya Polyana, Krasnodar Province;
<emphasis id="DBD3EAFF9306FF8AFAA8F17EFA9D1811" bold="true" box="[1330,1379,1572,1594]" pageId="16" pageNumber="105">C–D</emphasis>
from Mestia, Georgia;
<emphasis id="DBD3EAFF9306FF8AFED2F118FE861873" bold="true" box="[328,376,1602,1624]" pageId="16" pageNumber="105">E–H</emphasis>
from Teberda, Karachaevo-Cherkessia).
<emphasis id="DBD3EAFF9306FF8AFC86F118FCC81873" bold="true" box="[796,822,1602,1624]" pageId="16" pageNumber="105">A:</emphasis>
promere with flagella, caudal view.
<emphasis id="DBD3EAFF9306FF8AFB30F118FAFD1873" bold="true" box="[1194,1283,1602,1624]" pageId="16" pageNumber="105">B, D, H:</emphasis>
left gonopods, mesal view.
<emphasis id="DBD3EAFF9306FF8AFE8EF13BFE95185C" bold="true" box="[276,363,1633,1655]" pageId="16" pageNumber="105">C, E, G:</emphasis>
promere, caudal view.
<emphasis id="DBD3EAFF9306FF8AFDCDF13BFD8E185C" bold="true" box="[599,624,1633,1655]" pageId="16" pageNumber="105">F:</emphasis>
right gonopod, mesal view. Scale bar: 0.3 mm.
<emphasis id="DBD3EAFF9306FF8AFBC8F13BFB15185C" bold="true" box="[1106,1259,1633,1655]" pageId="16" pageNumber="105">Abbreviations</emphasis>
:
<emphasis id="DBD3EAFF9306FF8AFB62F13BFAF0185C" bold="true" box="[1272,1294,1633,1655]" pageId="16" pageNumber="105">fl,</emphasis>
flagellum;
<emphasis id="DBD3EAFF9306FF8AFA18F13BFA62185C" bold="true" box="[1410,1436,1633,1655]" pageId="16" pageNumber="105">la,</emphasis>
lamella on promere;
<emphasis id="DBD3EAFF9306FF8AFEF3F125FE7D18BE" bold="true" box="[361,387,1663,1685]" pageId="16" pageNumber="105">lo,</emphasis>
lateral outgrowth of opisthomere;
<emphasis id="DBD3EAFF9306FF8AFD79F125FCF618BE" bold="true" box="[739,776,1663,1685]" pageId="16" pageNumber="105">ms,</emphasis>
mesomere;
<emphasis id="DBD3EAFF9306FF8AFC1EF125FC5B18BE" bold="true" box="[900,933,1663,1685]" pageId="16" pageNumber="105">op,</emphasis>
opisthomere;
<emphasis id="DBD3EAFF9306FF8AFBACF125FBAD18BE" bold="true" box="[1078,1107,1663,1685]" pageId="16" pageNumber="105">pr,</emphasis>
promere.
</paragraph>
</caption>
<paragraph id="E91836ED9306FF8AFF5DF1E7FAA318FD" blockId="16.[151,1436,1725,2038]" box="[199,1373,1725,1750]" pageId="16" pageNumber="105">
Ventral edge of male segment 7 with small curved lamellae bordering the gonopodal aperture (
<figureCitation id="719C2A689306FF8AFB71F1E7FAAE18FD" box="[1259,1360,1725,1750]" captionStart="FIGURE 15" captionStartId="20.[151,249,1687,1709]" captionTargetBox="[340,1248,1178,1666]" captionTargetId="figure@20.[340,1248,1178,1666]" captionTargetPageId="20" captionText="FIGURE 15. Ventral edge of male segments 7 in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936 C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459902/files/figure.png" pageId="16" pageNumber="105">Fig. 15E</figureCitation>
).
</paragraph>
<paragraph id="E91836ED9306FF8AFF5DF1BBFCAE1935" blockId="16.[151,1436,1725,2038]" pageId="16" pageNumber="105">
Female vulva (
<emphasis id="DBD3EAFF9306FF8AFEE9F1BBFE6E18D1" bold="true" box="[371,400,1761,1786]" pageId="16" pageNumber="105">vu</emphasis>
) narrow, apices of bursa (
<emphasis id="DBD3EAFF9306FF8AFD24F1BBFD2318D1" bold="true" box="[702,733,1761,1786]" pageId="16" pageNumber="105">bu</emphasis>
) rounded; ampullae (
<emphasis id="DBD3EAFF9306FF8AFC4FF1BBFBF718D1" bold="true" box="[981,1033,1761,1786]" pageId="16" pageNumber="105">am1</emphasis>
and
<emphasis id="DBD3EAFF9306FF8AFBD9F1BBFB8818D1" bold="true" box="[1091,1142,1761,1786]" pageId="16" pageNumber="105">am2</emphasis>
) elongated. Each coxa of female leg-pair 2 (
<emphasis id="DBD3EAFF9306FF8AFEFCF05FFE821935" bold="true" box="[358,380,1797,1822]" pageId="16" pageNumber="105">l2</emphasis>
) with a small pointed process (
<figureCitation id="719C2A689306FF8AFD41F05FFCBC1935" box="[731,834,1797,1822]" captionStart="FIGURE 18" captionStartId="22.[151,249,1951,1973]" captionTargetBox="[340,1248,1166,1930]" captionTargetId="figure@22.[340,1248,1166,1930]" captionTargetPageId="22" captionText="FIGURE 18. Female vulvae and legs 2 in Caucasian species of Julus. A: J. lignaui Verhoeff, 1910. B: J. lindholmi Lohmander, 1936. C: J. subalpinus Lohmander, 1936. D: J. dagestanus sp. n. Abbreviations: vu, vulva; op, operculum; bu, bursa; am1 and am2, ampulla 1 and ampulla 2; l2, coxa and prefemur of leg 2. Scale bar: 0.3 mm." httpUri="https://zenodo.org/record/1459908/files/figure.png" pageId="16" pageNumber="105">Fig. 18A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A1BD65669306FF8AFF5DF073FEF319DD" pageId="16" pageNumber="105" type="discussion">
<paragraph id="E91836ED9306FF8AFF5DF073FEF319DD" blockId="16.[151,1436,1725,2038]" pageId="16" pageNumber="105">
<emphasis id="DBD3EAFF9306FF8AFF5DF073FEC51969" bold="true" box="[199,315,1833,1858]" pageId="16" pageNumber="105">Remarks.</emphasis>
A fairly rare, probably high-montane species hitherto recorded only from the
<collectingRegion id="2B63F80F9306FF8AFB21F073FACE1969" box="[1211,1328,1833,1858]" country="Russia" name="Krasnodar" pageId="16" pageNumber="105">Krasnodar</collectingRegion>
Province and
<collectingRegion id="2B63F80F9306FF8AFF55F017FEC4194D" box="[207,314,1869,1894]" country="Georgia" name="Abkhazia" pageId="16" pageNumber="105">Abkhazia</collectingRegion>
(
<bibRefCitation id="8D364B1C9306FF8AFECBF017FE12194D" box="[337,492,1869,1894]" pageId="16" pageNumber="105" refId="ref19444" refString="Lignau, N. (1903) [Myriapods of the Black Sea coast of the Caucasus]. Die Myriopoden am Kaukasischen Schwarzmeerufer. Memoires de la Societe Imperiale des Naturalistes de la Nouvelle-Russie. [Zapiski Imperatorskogo Novorossiyskogo obshchestva estestvoispytateley. Odessa], 25 (1), 82 - 149. [in Russian and German]" type="journal article">Lignau 1903</bibRefCitation>
;
<bibRefCitation id="8D364B1C9306FF8AFD9BF017FD53194D" box="[513,685,1869,1894]" pageId="16" pageNumber="105" refId="ref20298" refString="Verhoeff, K. W. (1910) Ueber Dipopoden. 19. (39.) Aufsatz. Juliden und Ascospermophora. Jahreshefte des Vereins fur vaterlandische Naturkunde im Wurttemberg, 66, 337 - 398." type="journal article">Verhoeff 1910</bibRefCitation>
;
<bibRefCitation id="8D364B1C9306FF8AFD5BF014FCBB194D" box="[705,837,1869,1894]" pageId="16" pageNumber="105" refId="ref19345" refString="Lang, J. (1959) [To the knowledge of millipedes (Diplopoda) of the USSR territory]. Zoologicheskii zhurnal, 38 (12), 1790 - 1796. [in Russian, a summary in English]" type="journal article">Lang 1959</bibRefCitation>
;
<bibRefCitation id="8D364B1C9306FF8AFCC3F017FBD5194D" box="[857,1067,1869,1894]" pageId="16" pageNumber="105" refId="ref19184" refString="Kobakhidze, D. N. (1965) [A list of millipedes (Diplopoda) of SSR Georgia]. Fragmenta Faunistica, 11 (21), 390 - 398. [in Russian, summaries in Polish and German]" type="journal article">Kobakhidze 1965</bibRefCitation>
;
<bibRefCitation id="8D364B1C9306FF8AFBA5F017FB08194D" box="[1087,1270,1869,1894]" pageId="16" pageNumber="105" refId="ref20114" refString="Talikadze, D. A. (1984) [On the millipede fauna (Diplopoda) of the Colchidan Province of the Caucasus]. Zoologicheskii zhurnal, 63 (1), 124 - 145 [in Russian, a summary in English]" type="journal article">Talikadze 1984</bibRefCitation>
;
<bibRefCitation id="8D364B1C9306FF8AFA90F017FEAE19A2" pageId="16" pageNumber="105" refId="ref19604" refString="Loksina, I. E. &amp; Golovatch, S. I. (1979) Diplopoda of the USSR fauna. Pedobiologia, 19, 381 - 389." type="journal article">Lokšina and Golovatch 1979</bibRefCitation>
). Based on geographical evidence alone, we cannot exclude the possibility that Lohmander’s (in
<bibRefCitation id="8D364B1C9306FF8AFF0DF0CFFEA01985" box="[151,350,1941,1966]" pageId="16" pageNumber="105" refId="ref19152" refString="Kobakhidze, D. N. (1964) [Myriapoda]. Animal life of Georgia, 2 (Arthropoda), 186 - 195 [Tbilisi, in Georgian]" type="book chapter">Kobakhidze 1964</bibRefCitation>
) “
<taxonomicName id="2EA74D6E9306FF8AFEE0F0CCFDF71986" box="[378,521,1942,1965]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="16" pageNumber="105" phylum="Arthropoda" rank="species" species="svaneticus">
<emphasis id="DBD3EAFF9306FF8AFEE0F0CCFDF71986" box="[378,521,1942,1965]" italics="true" pageId="16" pageNumber="105">J. svaneticus</emphasis>
</taxonomicName>
”
<emphasis id="DBD3EAFF9306FF8AFD86F0CFFD371985" bold="true" box="[540,713,1941,1966]" pageId="16" pageNumber="105">nomen nudum</emphasis>
actually represented
<taxonomicName id="2EA74D6E9306FF8AFC22F0CCFBDC1985" authorityName="Verhoeff" authorityYear="1910" box="[952,1058,1941,1966]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="16" pageNumber="105" phylum="Arthropoda" rank="species" species="lignaui">
<emphasis id="DBD3EAFF9306FF8AFC22F0CCFBDC1985" box="[952,1058,1941,1966]" italics="true" pageId="16" pageNumber="105">J. lignaui</emphasis>
</taxonomicName>
, the only species that
<bibRefCitation id="8D364B1C9306FF8AFA81F0CFFF1F19F9" pageId="16" pageNumber="105" refId="ref19572" refString="Lohmander, H. (1936) Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles Handlingar, 5, 5 B (1), 1 - 196." type="journal article">Lohmander (1936)</bibRefCitation>
failed to revise because of the lack of material. It is for this reason that we illustrate
<taxonomicName id="2EA74D6E9306FF8AFB01F0E0FAFA19F9" authorityName="Verhoeff" authorityYear="1910" box="[1179,1284,1977,2002]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="16" pageNumber="105" phylum="Arthropoda" rank="species" species="lignaui">
<emphasis id="DBD3EAFF9306FF8AFB01F0E0FAFA19F9" box="[1179,1284,1977,2002]" italics="true" pageId="16" pageNumber="105">J. lignaui</emphasis>
</taxonomicName>
well to show variations.
</paragraph>
</subSubSection>
</treatment>
</document>
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