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<document id="4207F54C86321DA3AEBE84DE9051C240" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66BD6FF7DAF89FBE0FD103E59" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera guangzhaishanica Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="185" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="184" updateTime="1698926631736" updateUser="ExternalLinkService">
<mods:mods id="8D50139B06562E41EA7B901BD03DBC6E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6759A7D195F0A9CF2591F6E3626E3C67">
<mods:title id="03499D925775864D5F7835E10566CF45">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
</mods:titleInfo>
<mods:name id="D5D4E03A4143059B0C03D5B2162A8C7C" type="personal">
<mods:role id="CF1B380828AE71819BF06D6A5F4669A5">
<mods:roleTerm id="EDC3562D4411D76FB888D76B8F58B4A5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9DAE3543E82C3A4D3F295444C710FDA7">Fabrizi, Silvia</mods:namePart>
<mods:nameIdentifier id="93C761DBB67B68E2FCEC6960F3F1B690" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="BFB7B544CB5327B645CB63BF5A3411E8" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:name id="7B8D8AA7BBEBA571C5FD890CC5A7376A" type="personal">
<mods:role id="58A07D1E5D9D410F575F2BB7565DB8A2">
<mods:roleTerm id="C8344B6E61058181382808F3C38C4C5C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="65AF9472C4D0BE7EBB4591C03BFD3BC6">Liu, Wan-Gang</mods:namePart>
<mods:nameIdentifier id="D769BD79055C62A66E0A025BC0CA9883" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:nameIdentifier id="125A7A10B77FDF41BA2475B8F475471B" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="4C3A1B0AEE0962293DF574F2DD732577" type="personal">
<mods:role id="0ED300B762EA1EBA70C5DCBD08CA1965">
<mods:roleTerm id="AE984A97DE0A82D4AB6C5556CE5E9F4C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DDC42504BD2C013E596A038235F48589">Bai, Ming</mods:namePart>
<mods:nameIdentifier id="3490E3A0D1823305E98C8B7F2124DD20" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="4E446D7900BBF1A51090F912EBB8CCBA" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="8A375E3138BDB402136FA20DA924DF22" type="personal">
<mods:role id="96FAA5AC4E42E8A21A31A2ED83239AA8">
<mods:roleTerm id="AD4A5DFF50193F90FF84277C82A4AB2C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C77C05A2B1C9D6E0AC6F4B8D398DACB6">Yang, Xing-Ke</mods:namePart>
<mods:nameIdentifier id="31854A3371B704D0D2D6023C5901B0D3" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="84D7163F08360ADCC4B71C523D35ED2C" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="82A7A2D37E035561394638ABDCF59E00" type="personal">
<mods:role id="4659E3E6821F0B8C0C5A23CB58720B7A">
<mods:roleTerm id="F931C4CBB8119BC367C4EDF497F3CD12">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="877F503AC0F1A0C05198C96C087ADA70">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="D6D80DA8B2D3F4004D6E94D18991B57B" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="A42793FFCB6C02539A22CF9D0CB1FD89" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="7FA48FE62C147F9B98AB99363C0C07E8">text</mods:typeOfResource>
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<mods:titleInfo id="84CCC8EEF25BCDBC753A01D73CE15EEB">
<mods:title id="F9AB5CF41F196C836A1B0768AA6BA1F9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="564509B43291F0A62831CA9B2670AD27">
<mods:date id="3C694E57534A103240E2B4DF73EA5B20">2021</mods:date>
<mods:detail id="AFFC82F9CA841974EEF3F8EDA7BED202" type="pubDate">
<mods:number id="9E2BD45845633418E0C93389AC526BA8">2021-02-03</mods:number>
</mods:detail>
<mods:detail id="17CD262D7D3DD30EC61BAAF33748FDDA" type="volume">
<mods:number id="063932B0E869E68052F8298C4E6B74D1">4922</mods:number>
</mods:detail>
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<mods:number id="73793A8CD178E6D375840AB72F66EB69">1</mods:number>
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<mods:extent id="6B87518F480BAA2DBD380DE8C6C34EBA" unit="page">
<mods:start id="4C06F963BB892019CC1B9B56463DEA31">1</mods:start>
<mods:end id="10459D2B94E61B33E3FF8C3581733C75">400</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="FEFD2003CA8E124912260963D3AC9B3B">journal article</mods:classification>
<mods:identifier id="75A0BBE04B87A4DFDE4CB39F230B479C" type="CLB-Dataset">8257</mods:identifier>
<mods:identifier id="ED7772B30ADF6355D356FD21B7BF0156" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
<mods:identifier id="7B0D8882880B244860D2955CB35E299B" type="GBIF-Dataset">188411de-569c-4408-a016-1a04b858c142</mods:identifier>
<mods:identifier id="CFBFD383E9FE10FA8FB4421B81EC2A77" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="34708D73E42B3AD627F7035C9342C3E8" type="Zenodo-Dep">4496316</mods:identifier>
<mods:identifier id="0BB6B7B8A5AFFC5C3014E98008CE1429" type="ZooBank">D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C</mods:identifier>
</mods:mods>
<treatment id="03BD87E66BD6FF7DAF89FBE0FD103E59" ID-DOI="http://doi.org/10.5281/zenodo.4534662" ID-GBIF-Taxon="177323553" ID-Zenodo-Dep="4534662" LSID="urn:lsid:plazi:treatment:03BD87E66BD6FF7DAF89FBE0FD103E59" httpUri="http://treatment.plazi.org/id/03BD87E66BD6FF7DAF89FBE0FD103E59" lastPageId="184" lastPageNumber="185" pageId="183" pageNumber="184">
<subSubSection id="C30E657B6BD6FF72AF89FBE0FCBC3EE9" box="[151,866,906,933]" pageId="183" pageNumber="184" type="nomenclature">
<paragraph id="8BAB36F06BD6FF72AF89FBE0FCBC3EE9" blockId="183.[151,866,906,969]" box="[151,866,906,933]" pageId="183" pageNumber="184">
<heading id="D0E3819C6BD6FF72AF89FBE0FCBC3EE9" bold="true" box="[151,866,906,933]" fontSize="11" level="2" pageId="183" pageNumber="184" reason="4">
<emphasis id="B960EAE26BD6FF72AF89FBE0FCBC3EE9" bold="true" box="[151,866,906,933]" pageId="183" pageNumber="184">
<taxonomicName id="4C144D736BD6FF72AF89FBE0FCCD3EE8" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,787,906,933]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="guangzhaishanica" status="sp. nov.">
<emphasis id="B960EAE26BD6FF72AF89FBE0FE2C3EE9" bold="true" box="[151,498,906,933]" italics="true" pageId="183" pageNumber="184">Maladera guangzhaishanica</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BD6FF72AC01FBE1FCBC3EE9" box="[799,866,907,933]" pageId="183" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD6FF72AF89FBC5FEA43E85" box="[151,378,943,969]" pageId="183" pageNumber="184" type="description">
<paragraph id="8BAB36F06BD6FF72AF89FBC5FEA43E85" blockId="183.[151,866,906,969]" box="[151,378,943,969]" pageId="183" pageNumber="184">
<figureCitation id="132F2A756BD6FF72AF89FBC5FED33E85" box="[151,269,943,969]" captionStart="FIGURE 41" captionStartId="182.[151,250,1936,1960]" captionTargetBox="[258,1331,322,1901]" captionTargetId="figure-72@182.[244,1343,309,1913]" captionTargetPageId="182" captionText="FIGURE 41. A–D: Maladera guanxianensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. guangzhaishanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. shiruguanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496416" httpUri="https://zenodo.org/record/4496416/files/figure.png" pageId="183" pageNumber="184">Figures 41</figureCitation>
E–H, 108
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD6FF72AF89FB9DFEF53931" pageId="183" pageNumber="184" type="materials_examined">
<paragraph id="8BAB36F06BD6FF72AF89FB9DFEF53931" blockId="183.[151,1437,1014,2013]" pageId="183" pageNumber="184">
<emphasis id="B960EAE26BD6FF72AF89FB9DFE1D395C" bold="true" box="[151,451,1014,1041]" pageId="183" pageNumber="184">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6BD6FF72AED3FB9DFC693979" ID-GBIF-Occurrence="3034556732" collectionCode="ZFMK" country="China" elevation="465" latitude="25.7" location="Jaroslav Turna" longLatPrecision="1245" longitude="116.78333" municipality="Guang Zhai Shan" pageId="183" pageNumber="184" specimenCount="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="holotype">
<typeStatus id="54AF88526BD6FF72AED3FB9DFDE4395D" box="[461,570,1015,1041]" pageId="183" pageNumber="184" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D12FD796BD6FF72AD58FB9DFD85395D" box="[582,603,1015,1041]" pageId="183" pageNumber="184" type="male">♂</specimenCount>
“
<collectingCountry id="F30376606BD6FF72AD70FB9DFD68395D" box="[622,694,1015,1041]" name="China" pageId="183" pageNumber="184">China</collectingCountry>
, SW
<collectingRegion id="49D0F8126BD6FF72ADE8FB9DFC9F395D" box="[758,833,1015,1041]" country="China" name="Fujian" pageId="183" pageNumber="184">Fujian</collectingRegion>
, 28.iv.-24.VII.
<collectingMunicipality id="6BCFAC8A6BD6FF72ACF1FB9DFB69395D" box="[1007,1207,1015,1041]" pageId="183" pageNumber="184">Guang Zhai Shan</collectingMunicipality>
<geoCoordinate id="EE2050376BD6FF72ABDFFB9CFAFF395D" box="[1217,1313,1014,1041]" degrees="25" direction="north" minutes="42" orientation="latitude" pageId="183" pageNumber="184" precision="925" value="25.7">25°42’N</geoCoordinate>
<geoCoordinate id="EE2050376BD6FF72AA35FB9DFA49395D" box="[1323,1431,1015,1041]" degrees="116" direction="east" minutes="47" orientation="longitude" pageId="183" pageNumber="184" precision="925" value="116.78333">116°47’E</geoCoordinate>
,
<quantity id="4CEC9B156BD6FF72AF89FC70FED43979" box="[151,266,1050,1077]" metricMagnitude="2" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="183" pageNumber="184" unit="m" value="465.0" valueMax="480.0" valueMin="450.0">
<elevation id="0039D1C36BD6FF72AF89FC70FED43979" box="[151,266,1050,1077]" metricMagnitude="2" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="183" pageNumber="184" unit="m" value="465.0" valueMax="480.0" valueMin="450.0">450-480m</elevation>
</quantity>
<location id="8ECB602B6BD6FF72AE0CFC71FE683979" LSID="urn:lsid:plazi:treatment:03BD87E66BD6FF7DAF89FBE0FD103E59:8ECB602B6BD6FF72AE0CFC71FE683979" box="[274,438,1051,1077]" country="China" latitude="25.7" longLatPrecision="1245" longitude="116.78333" municipality="Guang Zhai Shan" name="Jaroslav Turna" pageId="183" pageNumber="184" stateProvince="Fujian">Jaroslav Turna</location>
leg., 2006/ 783
<taxonomicName id="4C144D736BD6FF72AD70FC71FD1B3979" box="[622,709,1051,1077]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
Asia spec.” (
<collectionCode id="ED05AE356BD6FF72AC45FC71FC713979" box="[859,943,1051,1077]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="183" pageNumber="184" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BD6FF72ACDCFC71FD0F3915" ID-GBIF-Occurrence="3034555589" collectingDate="2003-10-01" collectionCode="ZFMK" country="China" location="Baimianshan" pageId="183" pageNumber="184" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54AF88526BD6FF72ACDCFC71FBF63979" box="[962,1064,1051,1077]" pageId="183" pageNumber="184" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D12FD796BD6FF72AB2DFC71FB833979" box="[1075,1117,1051,1077]" pageId="183" pageNumber="184" type="male">1 ♂</specimenCount>
“[
<collectingCountry id="F30376606BD6FF72AB68FC70FB1E3979" box="[1142,1216,1050,1077]" name="China" pageId="183" pageNumber="184">China</collectingCountry>
]
<location id="8ECB602B6BD6FF72ABD0FC71FABE3979" LSID="urn:lsid:plazi:treatment:03BD87E66BD6FF7DAF89FBE0FD103E59:8ECB602B6BD6FF72ABD0FC71FABE3979" box="[1230,1376,1051,1077]" country="China" name="Baimianshan" pageId="183" pageNumber="184">Baimianshan</location>
Yanling, F-Henugu,
<date id="FFAA10306BD6FF72AE4FFC55FE653915" box="[337,443,1087,1113]" pageId="183" pageNumber="184" value="2003-10-01">
<collectingDate id="EFEEE9D86BD6FF72AE4FFC55FE653915" box="[337,443,1087,1113]" pageId="183" pageNumber="184" value="2003-10-01">1.X.2003</collectingDate>
</date>
, Leg. Li
<emphasis id="B960EAE26BD6FF72AD3CFC55FD843915" box="[546,602,1087,1113]" italics="true" pageId="183" pageNumber="184">et al.</emphasis>
” (
<collectionCode id="ED05AE356BD6FF72AD6BFC55FD173915" box="[629,713,1087,1113]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="183" pageNumber="184" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BD6FF72ADC2FC55FEF93931" ID-GBIF-Occurrence="3034556719" collectingDate="1977-05-10" collectionCode="IZAS" collectorName="Zhang" country="China" location="Mt. Lushan" pageId="183" pageNumber="184" specimenCount="1" specimenCount-male="1" stateProvince="Jiangxi" typeStatus="paratype">
<specimenCount id="9D12FD796BD6FF72ADC2FC55FCD93915" box="[732,775,1087,1113]" pageId="183" pageNumber="184" type="male">1 ♂</specimenCount>
“[
<collectingCountry id="F30376606BD6FF72AC01FC54FCB73915" box="[799,873,1086,1113]" name="China" pageId="183" pageNumber="184">China</collectingCountry>
]
<location id="8ECB602B6BD6FF72AC69FC55FC253915" LSID="urn:lsid:plazi:treatment:03BD87E66BD6FF7DAF89FBE0FD103E59:8ECB602B6BD6FF72AC69FC55FC253915" box="[887,1019,1087,1113]" country="China" name="Mt. Lushan" pageId="183" pageNumber="184" stateProvince="Jiangxi">Mt. Lushan</location>
,
<collectingRegion id="49D0F8126BD6FF72AB19FC55FB853915" box="[1031,1115,1087,1113]" country="China" name="Jiangxi" pageId="183" pageNumber="184">Jiangxi</collectingRegion>
,
<date id="FFAA10306BD6FF72AB78FC55FB023915" box="[1126,1244,1087,1113]" pageId="183" pageNumber="184" value="1977-05-10">
<collectingDate id="EFEEE9D86BD6FF72AB78FC55FB023915" box="[1126,1244,1087,1113]" pageId="183" pageNumber="184" value="1977-05-10">10.V.1977</collectingDate>
</date>
, leg.
<collectorName id="26E153266BD6FF72AA06FC55FABE3915" box="[1304,1376,1087,1113]" pageId="183" pageNumber="184">Zhang</collectorName>
Youwei” (
<collectionCode id="ED05AE356BD6FF72AFC5FC09FEC13931" box="[219,287,1123,1149]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="183" pageNumber="184">IZAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD6FF7DAFD9FCECFCA83F55" lastPageId="184" lastPageNumber="185" pageId="183" pageNumber="184" type="description">
<paragraph id="8BAB36F06BD6FF72AFD9FCECFB153989" blockId="183.[151,1437,1014,2013]" pageId="183" pageNumber="184">
<emphasis id="B960EAE26BD6FF72AFD9FCECFE8739EC" bold="true" box="[199,345,1158,1184]" pageId="183" pageNumber="184">Description.</emphasis>
Length:
<quantity id="4CEC9B156BD6FF72AEA0FCEDFDFA39ED" box="[446,548,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="183" pageNumber="184" unit="mm" value="10.2">10.2 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156BD6FF72ADF5FCEDFC9D39ED" box="[747,835,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="183" pageNumber="184" unit="mm" value="7.9">7.9 mm</quantity>
, width:
<quantity id="4CEC9B156BD6FF72AC80FCEDFC2939ED" box="[926,1015,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="183" pageNumber="184" unit="mm" value="6.4">6.4 mm</quantity>
. Body wide, oval, reddish brown, antenna yellowish brown, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous.
</paragraph>
<paragraph id="8BAB36F06BD6FF72AFD9FCA5FC273B45" blockId="183.[151,1437,1014,2013]" pageId="183" pageNumber="184">
Labroclypeus wide, lateral margins weakly convex and moderately convergent, lateral margins produce with ocular canthus an indistinct angle, anterior angles convex, anterior margin shallowly emarginate medially, margins weakly reflexed; surface weakly shiny, widely dull at base, surface convex medially, finely and densely punctate, distance between punctures smaller their diameter, with a few long erect setae; smooth area in front of eyes three times as wide as long; frontoclypeal suture finely incised, bluntly angled medially; ocular canthus moderately wide and short (1/3 of ocular diameter), finely and densely punctate, without a short terminal seta. Eyes small, ratio diameter/ interocular width: 0.58. Frons with fine and moderately dense punctures, with a few single setae beside eyes. Antenna with ten antennomeres, club with three antennomeres and straight, distinctly shorter than remaining antennomeres combined.
<collectingRegion id="49D0F8126BD6FF72AEA8FD85FDCB3B45" box="[438,533,1519,1545]" country="Myanmar" name="Chin" pageId="183" pageNumber="184">Mentum</collectingRegion>
convexly elevated and flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06BD6FF72AFD9FE79FB1C3BD5" blockId="183.[151,1437,1014,2013]" pageId="183" pageNumber="184">Pronotum widest at base, lateral margins weakly convex and weakly convergent anteriorly, anterior angles sharp and distinctly produced; anterior margin convex, with widely interrupted marginal line; basal margin without marginal line; surface finely and densely punctate, punctures with microscopic setae only, setae of lateral and anterior margin sparse; hypomeron carinate. Scutellum wide, triangular, punctation as in pronotum.</paragraph>
<paragraph id="8BAB36F06BD6FF72AFD9FEC9FDA03A65" blockId="183.[151,1437,1014,2013]" pageId="183" pageNumber="184">
Elytra (left elytron slightly deformed in
<typeStatus id="54AF88526BD6FF72AD9AFEC9FD363BF2" box="[644,744,1699,1726]" pageId="183" pageNumber="184" type="holotype">holotype</typeStatus>
) widest at posterior third, striae finely impressed, finely punctate, intervals flat, finely and densely punctate, with minute setae in punctures, odd intervals with a few erect short setae; epipleura ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes.
</paragraph>
<paragraph id="8BAB36F06BD6FF72AFD9FF59FF383A91" blockId="183.[151,1437,1014,2013]" pageId="183" pageNumber="184">Ventral face coarsely and densely punctate, nearly glabrous, only metasternal plate and lateral metacoxa with a few robust setae. Mesosternum between mesocoxae wide, as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.71. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta. Pygidium strongly convex, coarsely and densely punctate, long setae on apical margin absent.</paragraph>
<paragraph id="8BAB36F06BD9FF7DAFD9F8FDFB2A3CB9" blockId="184.[151,1437,151,789]" pageId="184" pageNumber="185">Legs moderately long and wide; femora superficially and sparsely punctate, with two longitudinal rows of setae. Metafemur shiny, anterior margin acute, without adjacent serrated line, anterior longitudinal row of setae absent; posterior ventral margin almost straight, not widened in apical half, neither ventrally nor dorsally serrated but smooth, glabrous. Metatibia short and wide, flattened, ratio width/length: 1/2.9, sharply carinate dorsally, with two groups of spines, basal one at basal third, apical one at three quarters of metatibial length, basally with a few robust and short setae; lateral face weakly longitudinally convex, with a very superficial fine punctures; medial face impunctate, apex shallowly and concavely emarginate interiorly near tarsal articulation. Tarsomeres sparsely finely punctate dorsally, with dense, fine setae ventrally; metatarsomeres ventrally glabrous, with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere as long as following two tarsomeres combined, little longer than dorsal tibial spur. Protibia short, bidentate. Anterior claws symmetrical.</paragraph>
<paragraph id="8BAB36F06BD9FF7DAFD9F995FCA83F55" blockId="184.[151,1437,151,789]" box="[199,886,511,537]" pageId="184" pageNumber="185">
Aedeagus:
<figureCitation id="132F2A756BD9FF7DAE5AF995FE4D3F55" box="[324,403,511,537]" captionStart="FIGURE 41" captionStartId="182.[151,250,1936,1960]" captionTargetBox="[258,1331,322,1901]" captionTargetId="figure-72@182.[244,1343,309,1913]" captionTargetPageId="182" captionText="FIGURE 41. A–D: Maladera guanxianensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. guangzhaishanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. shiruguanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496416" httpUri="https://zenodo.org/record/4496416/files/figure.png" pageId="184" pageNumber="185">Fig. 41</figureCitation>
E–G. Habitus:
<figureCitation id="132F2A756BD9FF7DAD25F995FD7F3F55" box="[571,673,511,537]" captionStart="FIGURE 41" captionStartId="182.[151,250,1936,1960]" captionTargetBox="[258,1331,322,1901]" captionTargetId="figure-72@182.[244,1343,309,1913]" captionTargetPageId="182" captionText="FIGURE 41. A–D: Maladera guanxianensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. guangzhaishanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. shiruguanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496416" httpUri="https://zenodo.org/record/4496416/files/figure.png" pageId="184" pageNumber="185">Fig. 41H</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD9FF7DAFD9FA48FA433FCA" pageId="184" pageNumber="185" type="diagnosis">
<paragraph id="8BAB36F06BD9FF7DAFD9FA48FA433FCA" blockId="184.[151,1437,151,789]" pageId="184" pageNumber="185">
<emphasis id="B960EAE26BD9FF7DAFD9FA48FE9C3F70" bold="true" box="[199,322,546,572]" pageId="184" pageNumber="185">Diagnosis.</emphasis>
<taxonomicName id="4C144D736BD9FF7DAE55FA49FC423F71" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[331,924,547,573]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="guangzhaishanica" status="sp. nov.">
<emphasis id="B960EAE26BD9FF7DAE55FA49FD513F71" box="[331,655,547,573]" italics="true" pageId="184" pageNumber="185">Maladera guangzhaishanica</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BD9FF7DACB4FA49FC363F71" box="[938,1000,547,573]" pageId="184" pageNumber="185" rank="species">sp. n.</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C144D736BD9FF7DABDFFA49FE353F2D" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="guanxiamensis" status="sp. nov.">
<emphasis id="B960EAE26BD9FF7DABDFFA49FF3D3F2C" italics="true" pageId="184" pageNumber="185">Maladera guanxiamensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BD9FF7DAEE6FA2DFDEB3F2D" box="[504,565,583,609]" pageId="184" pageNumber="185" rank="species">sp. n.</taxonomicNameLabel>
in habitus and shape of the aedeagus. The former is distinguished in having the left paramere wider (lateral view) and the right paramere shorter and strongly pointed (both in lateral view).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD9FF7DAFD9FAE5FE593F82" pageId="184" pageNumber="185" type="etymology">
<paragraph id="8BAB36F06BD9FF7DAFD9FAE5FE593F82" blockId="184.[151,1437,151,789]" pageId="184" pageNumber="185">
<emphasis id="B960EAE26BD9FF7DAFD9FAE5FE903FE5" bold="true" box="[199,334,655,681]" pageId="184" pageNumber="185">Etymology.</emphasis>
The name of the new species is derived from its
<typeStatus id="54AF88526BD9FF7DAC60FAE5FC703FE5" box="[894,942,655,681]" pageId="184" pageNumber="185">type</typeStatus>
locality, Guang Zhai
<collectingRegion id="49D0F8126BD9FF7DABB8FAE5FB003FE5" box="[1190,1246,655,681]" country="Myanmar" name="Shan" pageId="184" pageNumber="185">Shan</collectingRegion>
(adjective in the nominative singular).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD9FF7DAFD9FABCFBAC3FBD" box="[199,1138,726,753]" pageId="184" pageNumber="185" type="description">
<paragraph id="8BAB36F06BD9FF7DAFD9FABCFBAC3FBD" blockId="184.[151,1437,151,789]" box="[199,1138,726,753]" pageId="184" pageNumber="185">
<emphasis id="B960EAE26BD9FF7DAFD9FABCFEE13FBC" bold="true" box="[199,319,726,752]" pageId="184" pageNumber="185">Variation.</emphasis>
Length:
<quantity id="4CEC9B156BD9FF7DAEBAFABDFDE53FBD" box="[420,571,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="1.005" metricValueMax="1.02" metricValueMin="0.99" pageId="184" pageNumber="185" unit="mm" value="10.05" valueMax="10.2" valueMin="9.9">9.9–10.2 mm</quantity>
, length of elytra: 7.0–
<quantity id="4CEC9B156BD9FF7DAC2CFABDFC573FBD" box="[818,905,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="184" pageNumber="185" unit="mm" value="7.9">7.9 mm</quantity>
, width:
<quantity id="4CEC9B156BD9FF7DACFDFABCFBB33FBD" box="[995,1133,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.4" metricValueMin="5.6" pageId="184" pageNumber="185" unit="mm" value="6.0" valueMax="6.4" valueMin="5.6">5.6–6.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BD9FF7DAFD9FA90FD103E59" box="[199,718,762,789]" pageId="184" pageNumber="185" type="distribution">
<paragraph id="8BAB36F06BD9FF7DAFD9FA90FD103E59" blockId="184.[151,1437,151,789]" box="[199,718,762,789]" pageId="184" pageNumber="185">
<emphasis id="B960EAE26BD9FF7DAFD9FA90FEBF3E58" bold="true" box="[199,353,762,788]" pageId="184" pageNumber="185">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756BD9FF7DAEC8FA91FDE83E59" box="[470,566,763,789]" captionStart="FIGURE 108" captionStartId="355.[151,250,1963,1987]" captionTargetBox="[214,1370,1124,1931]" captionTargetId="figure-60@355.[208,1379,1118,1940]" captionTargetPageId="355" captionText="FIGURE 108. Distribution of the Maladera granuligera group: Maladera baoxingensis sp. n., M. guangzhaishanica sp. n., M. guanxianensis sp. n., M. lushanensis sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496610" httpUri="https://zenodo.org/record/4496610/files/figure.png" pageId="184" pageNumber="185">Fig. 108</figureCitation>
) and
<tableCitation id="C696034B6BD9FF7DAD6AFA91FD163E59" box="[628,712,763,789]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="184" pageNumber="185">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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