183 lines
22 KiB
XML
183 lines
22 KiB
XML
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<mods:title id="C470693B6C668BFE7AA4465CB674D820">A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="C467CE74107B14AE256FA842B95ECA54">Fabrizi, Silvia</mods:namePart>
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<mods:namePart id="9A313118C9A82556E0E273C908E70B75">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="9633FADAE97829249D073040C5632A22">Bai, Ming</mods:namePart>
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<mods:namePart id="1574258B52468448ABD4C126DDE9E0FE">Yang, Xing-Ke</mods:namePart>
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<mods:namePart id="9399F98403FAB0EEFF216C495244BA9F">Ahrens, Dirk</mods:namePart>
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<mods:title id="5F00E3B6BF993511C4052687CBE60E40">Zootaxa</mods:title>
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<mods:date id="2C22D16B4E9E8574BA6F90E48A3290D1">2021</mods:date>
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<mods:number id="BB4D79B9B1615D38EA47BC5C8581C9B9">2021-02-03</mods:number>
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<mods:number id="A064D9D7BA82FFFF7718CAD50DD3BA5A">4922</mods:number>
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<mods:classification id="9D19A77C20688274C8704683EA772022">journal article</mods:classification>
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<mods:identifier id="98A3BC2132DB221BBD1B30A943082403" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
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<treatment id="03BD87E66BD5FF70AF89FABCFD103CB9" ID-DOI="http://doi.org/10.5281/zenodo.4534656" ID-GBIF-Taxon="177323552" ID-Zenodo-Dep="4534656" LSID="urn:lsid:plazi:treatment:03BD87E66BD5FF70AF89FABCFD103CB9" httpUri="http://treatment.plazi.org/id/03BD87E66BD5FF70AF89FABCFD103CB9" lastPageId="181" lastPageNumber="182" pageId="180" pageNumber="181">
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<subSubSection id="C30E657B6BD5FF71AF89FABCFCC33FBD" box="[151,797,726,753]" pageId="180" pageNumber="181" type="nomenclature">
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<paragraph id="8BAB36F06BD5FF71AF89FABCFCC33FBD" blockId="180.[151,797,726,789]" box="[151,797,726,753]" pageId="180" pageNumber="181">
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<emphasis id="B960EAE26BD5FF71AF89FABCFCC33FBD" bold="true" box="[151,797,726,753]" pageId="180" pageNumber="181">
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<taxonomicName id="4C144D736BD5FF71AF89FABCFD103FBC" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" box="[151,718,726,753]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="mupingensis" status="sp. nov.">
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<emphasis id="B960EAE26BD5FF71AF89FABCFE733FBD" bold="true" box="[151,429,726,753]" italics="true" pageId="180" pageNumber="181">Maladera mupingensis</emphasis>
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Ahrens, Fabrizi & Liu
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25357996BD5FF71ADC5FABDFCC33FBD" box="[731,797,727,753]" pageId="180" pageNumber="181" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C30E657B6BD5FF71AF89FA91FEB13E59" box="[151,367,763,789]" pageId="180" pageNumber="181" type="description">
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<paragraph id="8BAB36F06BD5FF71AF89FA91FEB13E59" blockId="180.[151,797,726,789]" box="[151,367,763,789]" pageId="180" pageNumber="181">
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<figureCitation id="132F2A756BD5FF71AF89FA91FED13E59" box="[151,271,763,789]" captionStart="FIGURE 40" captionStartId="178.[151,250,1943,1967]" captionTargetBox="[196,1391,193,1911]" captionTargetId="figure-19@178.[182,1404,181,1920]" captionTargetPageId="178" captionText="FIGURE 40. A–D: Maladera lushanensis Ahrens, Fabrizi & Liu, sp. n. (holotype); E–H: M. maguanensis Ahrens, Fabrizi & Liu, sp. n. (holotype); I–L: M. mupingensis Ahrens, Fabrizi & Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496414" httpUri="https://zenodo.org/record/4496414/files/figure.png" pageId="180" pageNumber="181">Figures 40</figureCitation>
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I–L, 109
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30E657B6BD5FF71AF89FB29FE903ECD" pageId="180" pageNumber="181" type="materials_examined">
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<paragraph id="8BAB36F06BD5FF71AF89FB29FE903ECD" blockId="180.[151,1437,834,2013]" pageId="180" pageNumber="181">
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<emphasis id="B960EAE26BD5FF71AF89FB29FE1F3E10" bold="true" box="[151,449,834,861]" pageId="180" pageNumber="181">Type material examined.</emphasis>
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||
<materialsCitation id="3B7C3CAD6BD5FF71AED7FB29FE903ECD" ID-GBIF-Occurrence="3034556526" collectionCode="USNM" country="China" location="China" pageId="180" pageNumber="181" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="54AF88526BD5FF71AED7FB29FDE83E11" box="[457,566,835,861]" pageId="180" pageNumber="181" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D12FD796BD5FF71AD5CFB29FD893E11" box="[578,599,835,861]" pageId="180" pageNumber="181" type="male">♂</specimenCount>
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“Muping Aug 25-28 ‘29/ Szechuen
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<collectingCountry id="F30376606BD5FF71ACEBFB29FBE63E11" box="[1013,1080,835,861]" name="China" pageId="180" pageNumber="181">China</collectingCountry>
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DC Graham/ 884
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<taxonomicName id="4C144D736BD5FF71AA15FB29FABC3E11" box="[1291,1378,835,861]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="180" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
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Asia spec.” (
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<collectionCode id="ED05AE356BD5FF71AFF5FB0DFE9C3ECD" box="[235,322,871,897]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="180" pageNumber="181" type="Museum">USNM</collectionCode>
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||
).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30E657B6BD5FF70AFD9FBE0FCB53C0D" lastPageId="181" lastPageNumber="182" pageId="180" pageNumber="181" type="description">
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||
<paragraph id="8BAB36F06BD5FF71AFD9FBE0FB153E85" blockId="180.[151,1437,834,2013]" pageId="180" pageNumber="181">
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||
<emphasis id="B960EAE26BD5FF71AFD9FBE0FE873EE8" bold="true" box="[199,345,906,932]" pageId="180" pageNumber="181">Description.</emphasis>
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Length:
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<quantity id="4CEC9B156BD5FF71AEA0FBE1FDFA3EE9" box="[446,548,907,933]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="180" pageNumber="181" unit="mm" value="10.3">10.3 mm</quantity>
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, length of elytra:
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<quantity id="4CEC9B156BD5FF71ADF5FBE1FC9D3EE9" box="[747,835,907,933]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="180" pageNumber="181" unit="mm" value="7.7">7.7 mm</quantity>
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, width:
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<quantity id="4CEC9B156BD5FF71AC80FBE0FC293EE9" box="[926,1015,906,933]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="180" pageNumber="181" unit="mm" value="5.5">5.5 mm</quantity>
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. Body wide, oval, reddish brown, antenna yellowish brown, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous.
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</paragraph>
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<paragraph id="8BAB36F06BD5FF71AFD9FBB9FC273841" blockId="180.[151,1437,834,2013]" pageId="180" pageNumber="181">
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Labroclypeus wide, lateral margins straight and moderately convergent, lateral margins produce with ocular canthus an indistinct angle, anterior angles convex, anterior margin shallowly emarginate medially, margins weakly reflexed; surface moderately shiny, at base widely dull, surface weakly convex, finely and densely punctate, distance between punctures smaller their diameter, with numerous long erect setae; smooth area in front of eyes three times as wide as long; frontoclypeal suture finely incised, bluntly angled medially; ocular canthus moderately wide and short (1/3 of ocular diameter), finely and densely punctate, short terminal seta absent. Eyes moderately large, ratio diameter/ interocular width: 0.66. Frons with fine and moderately dense punctures, with a few single setae beside eyes. Antenna with ten antennomeres, club with three antennomeres and straight, distinctly shorter than remaining antennomeres combined.
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<collectingRegion id="49D0F8126BD5FF71AEA8FC99FDCB3841" box="[438,533,1267,1293]" country="Myanmar" name="Chin" pageId="180" pageNumber="181">Mentum</collectingRegion>
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convexly elevated and flattened anteriorly.
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</paragraph>
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<paragraph id="8BAB36F06BD5FF71AFD9FD7DFAEC38D1" blockId="180.[151,1437,834,2013]" pageId="180" pageNumber="181">Pronotum widest at base, lateral margins evenly convex and convergent anteriorly, anterior angles sharp and distinctly produced; anterior margin convex, with complete marginal line; basal margin without marginal line; surface finely and densely punctate, punctures with microscopic setae only, setae of lateral and anterior margin sparse; hypomeron carinate. Scutellum wide, triangular, punctation as in pronotum, impunctate on basal midline.</paragraph>
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<paragraph id="8BAB36F06BD5FF71AFD9FDCDFD233B61" blockId="180.[151,1437,834,2013]" pageId="180" pageNumber="181">Elytra convex, widest at posterior third, striae finely impressed, finely punctate, intervals weakly convex, finely and densely punctate, punctures concentrated along striae, with minute setae in punctures, odd intervals with a few erect short setae; epipleura ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes.</paragraph>
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<paragraph id="8BAB36F06BD5FF71AFD9FE5DFE9E3BAD" blockId="180.[151,1437,834,2013]" pageId="180" pageNumber="181">Ventral face coarsely and densely punctate, nearly glabrous, only metasternal plate and lateral metacoxa with a few robust setae. Distance between mesocoxae little less than width of mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.67. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta. Pygidium moderately convex, coarsely and densely punctate, on apex with a few long setae.</paragraph>
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<paragraph id="8BAB36F06BD5FF70AFD9FE81FEAE3C51" blockId="180.[151,1437,834,2013]" lastBlockId="181.[151,1437,151,501]" lastPageId="181" lastPageNumber="182" pageId="180" pageNumber="181">Legs moderately long and wide; femora superficially and sparsely punctate, with two longitudinal rows of setae. Metafemur shiny, anterior margin acute, without adjacent serrated line, anterior longitudinal row of setae absent; posterior ventral margin almost straight, not widened in apical half, neither ventrally nor dorsally serrated but smooth, glabrous. Metatibia moderately long and wide, dorsal and ventral margin subparallel, ratio width/length: 1/3.3, sharply carinate dorsally, with two groups of spines, basal one at basal third, apical one at three quarters of metatibial length, basally with a few robust and short setae; lateral face weakly longitudinally convex, with fine sparse punctures; medial face impunctate, apex shallowly and concavely emarginate interiorly near tarsal articulation. Tarsomeres sparsely finely punctate dorsally, with dense, fine setae ventrally; metatarsomeres ventrally glabrous, with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere as long as following two tarsomeres combined, little longer than dorsal tibial spur. Protibia short, bidentate.Anterior claws symmetrical.</paragraph>
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<paragraph id="8BAB36F06BD4FF70AFD9F94DFCB53C0D" blockId="181.[151,1437,151,501]" box="[199,875,295,321]" pageId="181" pageNumber="182">
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||
Aedeagus:
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||
<figureCitation id="132F2A756BD4FF70AE5AF94DFE4B3C0D" box="[324,405,295,321]" captionStart="FIGURE 40" captionStartId="178.[151,250,1943,1967]" captionTargetBox="[196,1391,193,1911]" captionTargetId="figure-19@178.[182,1404,181,1920]" captionTargetPageId="178" captionText="FIGURE 40. A–D: Maladera lushanensis Ahrens, Fabrizi & Liu, sp. n. (holotype); E–H: M. maguanensis Ahrens, Fabrizi & Liu, sp. n. (holotype); I–L: M. mupingensis Ahrens, Fabrizi & Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496414" httpUri="https://zenodo.org/record/4496414/files/figure.png" pageId="181" pageNumber="182">Fig. 40</figureCitation>
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I–K. Habitus:
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<figureCitation id="132F2A756BD4FF70AD2DF94DFD493C0D" box="[563,663,295,321]" captionStart="FIGURE 40" captionStartId="178.[151,250,1943,1967]" captionTargetBox="[196,1391,193,1911]" captionTargetId="figure-19@178.[182,1404,181,1920]" captionTargetPageId="178" captionText="FIGURE 40. A–D: Maladera lushanensis Ahrens, Fabrizi & Liu, sp. n. (holotype); E–H: M. maguanensis Ahrens, Fabrizi & Liu, sp. n. (holotype); I–L: M. mupingensis Ahrens, Fabrizi & Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496414" httpUri="https://zenodo.org/record/4496414/files/figure.png" pageId="181" pageNumber="182">
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Fig.
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<specimenCount id="9D12FD796BD4FF70AD79F94DFD493C0D" box="[615,663,295,321]" pageId="181" pageNumber="182" type="generic">40L</specimenCount>
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</figureCitation>
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. Female unknown.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C30E657B6BD4FF70AFD9F920FCF93CE1" pageId="181" pageNumber="182" type="diagnosis">
|
||
<paragraph id="8BAB36F06BD4FF70AFD9F920FCF93CE1" blockId="181.[151,1437,151,501]" pageId="181" pageNumber="182">
|
||
<emphasis id="B960EAE26BD4FF70AFD9F920FE9C3C28" bold="true" box="[199,322,330,356]" pageId="181" pageNumber="182">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C144D736BD4FF70AE56F921FC903C29" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" box="[328,846,331,357]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="mupingensis" status="sp. nov.">
|
||
<emphasis id="B960EAE26BD4FF70AE56F921FD943C28" box="[328,586,331,357]" italics="true" pageId="181" pageNumber="182">Maladera mupingensis</emphasis>
|
||
Ahrens, Fabrizi & Liu
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25357996BD4FF70AC47F921FC4B3C29" box="[857,917,331,357]" pageId="181" pageNumber="182" rank="species">sp. n.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C144D736BD4FF70AB37F921FEEA3CC5" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="guanxianensis" status="sp. nov.">
|
||
<emphasis id="B960EAE26BD4FF70AB37F921FAE13C28" box="[1065,1343,331,357]" italics="true" pageId="181" pageNumber="182">Maladera guanxianensis</emphasis>
|
||
Ahrens, Fabrizi & Liu
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25357996BD4FF70AE21F905FEA53CC5" box="[319,379,367,393]" pageId="181" pageNumber="182" rank="species">sp. n.</taxonomicNameLabel>
|
||
and all other species possessing a dorsal apophysis on the phallobase in having the right paramere distinctly divided into a long dorsal and ventral lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6BD4FF70AFD9F9DDFA583C9E" box="[199,1414,439,466]" pageId="181" pageNumber="182" type="etymology">
|
||
<paragraph id="8BAB36F06BD4FF70AFD9F9DDFA583C9E" blockId="181.[151,1437,151,501]" box="[199,1414,439,466]" pageId="181" pageNumber="182">
|
||
<emphasis id="B960EAE26BD4FF70AFD9F9DDFE903C9D" bold="true" box="[199,334,439,465]" pageId="181" pageNumber="182">Etymology.</emphasis>
|
||
The new species is named after the
|
||
<typeStatus id="54AF88526BD4FF70ADFDF9DDFCCD3C9D" box="[739,787,439,465]" pageId="181" pageNumber="182">type</typeStatus>
|
||
locality, Muping (adjective in the nominative singular).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6BD4FF70AFD9F9B0FD103CB9" box="[199,718,474,501]" pageId="181" pageNumber="182" type="distribution">
|
||
<paragraph id="8BAB36F06BD4FF70AFD9F9B0FD103CB9" blockId="181.[151,1437,151,501]" box="[199,718,474,501]" pageId="181" pageNumber="182">
|
||
<emphasis id="B960EAE26BD4FF70AFD9F9B0FEBF3CB8" bold="true" box="[199,353,474,500]" pageId="181" pageNumber="182">Distribution.</emphasis>
|
||
See map (
|
||
<figureCitation id="132F2A756BD4FF70AEC8F9B1FDE83CB9" box="[470,566,475,501]" captionStart="FIGURE 109" captionStartId="356.[151,250,1044,1068]" captionTargetBox="[203,1383,189,1012]" captionTargetId="figure-19@356.[196,1391,181,1020]" captionTargetPageId="356" captionText="FIGURE 109. Distribution of the Maladera granuligera group: Maladera maguanensis sp. n., M. mupingensis sp. n., M. shiruguanensis sp. n., M. siniaevi Ahrens, 2004." figureDoi="http://doi.org/10.5281/zenodo.4496614" httpUri="https://zenodo.org/record/4496614/files/figure.png" pageId="181" pageNumber="182">Fig. 109</figureCitation>
|
||
) and
|
||
<tableCitation id="C696034B6BD4FF70AD6AF9B1FD163CB9" box="[628,712,475,501]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant & Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="181" pageNumber="182">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |