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<mods:title id="D3C68D29612D91AAE747C0FB395C4FFA">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="1E7262F7A4132E0C98DCF103555500C5">Liu, Wan-Gang</mods:namePart>
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<subSubSection id="C30E657B6B09FFADAF89FF64FDFC3A64" box="[151,546,1806,1833]" pageId="104" pageNumber="105" type="nomenclature">
<paragraph id="8BAB36F06B09FFADAF89FF64FDFC3A64" blockId="104.[151,546,1806,1869]" box="[151,546,1806,1833]" pageId="104" pageNumber="105">
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<taxonomicName id="4C144D736B09FFADAF89FF64FDFC3A64" authority="(Arrow, 1946)" baseAuthorityName="Arrow" baseAuthorityYear="1946" box="[151,546,1806,1833]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="egregia">
<emphasis id="B960EAE26B09FFADAF89FF64FDFC3A64" bold="true" box="[151,546,1806,1833]" pageId="104" pageNumber="105">
<emphasis id="B960EAE26B09FFADAF89FF64FEB03A65" bold="true" box="[151,366,1806,1833]" italics="true" pageId="104" pageNumber="105">Maladera egregia</emphasis>
(Arrow, 1946)
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B09FFADAF89FF59FEB13A01" box="[151,367,1843,1869]" pageId="104" pageNumber="105" type="description">
<paragraph id="8BAB36F06B09FFADAF89FF59FEB13A01" blockId="104.[151,546,1806,1869]" box="[151,367,1843,1869]" pageId="104" pageNumber="105">
<figureCitation id="132F2A756B09FFADAF89FF59FED03A01" box="[151,270,1843,1869]" captionStart="FIGURE 22" captionStartId="105.[151,250,1900,1924]" captionTargetBox="[213,1369,283,1871]" captionTargetId="figure-75@105.[202,1385,273,1876]" captionTargetPageId="105" captionText="FIGURE 22. AD: Maladera egregia (Arrow, 1946) (syntype: Myanmar: Kambaiti); EI: M. levis (Frey, 1972) (holotype); JM: M. hmong Ahrens, 2004 (Paratype: Vietnam: Sa Pa). A, E, J: aedeagus, left side lateral view; C, H, L: aedeagus, right side lateral view; B, F, K: parameres, dorsal view; G: parameres, ventral view; D, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496362" httpUri="https://zenodo.org/record/4496362/files/figure.png" pageId="104" pageNumber="105">Figures 22</figureCitation>
AD, 97
</paragraph>
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<paragraph id="8BAB36F06B09FFADAF89FF1DFE263AC3" blockId="104.[151,546,1911,1966]" box="[151,504,1911,1935]" pageId="104" pageNumber="105">
<treatmentCitationGroup id="AB0411DE6B09FFADAF89FF1DFE263AC3" box="[151,504,1911,1935]" pageId="104" pageNumber="105">
<treatmentCitation id="0AB510E16B09FFADAF89FF1DFE2A3AC3" author="Arrow, G. J." box="[151,500,1911,1935]" page="14" pageId="104" pageNumber="105" year="1946">
<taxonomicName id="4C144D736B09FFADAF89FF1DFE2A3AC3" authority="Arrow, 1946 b: 14" authorityName="Arrow" authorityPageNumber="14" authorityYear="1946" box="[151,500,1911,1935]" class="Insecta" family="Melolonthidae" genus="Aserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="egregia">
<emphasis id="B960EAE26B09FFADAF89FF1DFEE93AC3" box="[151,311,1911,1935]" italics="true" pageId="104" pageNumber="105">Aserica egregia</emphasis>
<bibRefCitation id="EF854B016B09FFADAE20FF1DFE2A3AC3" author="Arrow, G. J." box="[318,500,1911,1935]" pageId="104" pageNumber="105" pagination="1 - 33" refId="ref331251" refString="Arrow, G. J. (1946 b) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Melolonthidae. Arkiv for Zoologie, 38 A (9), 1 - 33." type="journal article" year="1946">Arrow, 1946b: 14</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BAB36F06B09FFADAF89FFFCFDFC3AE2" blockId="104.[151,546,1911,1966]" box="[151,546,1942,1966]" pageId="104" pageNumber="105">
<treatmentCitationGroup id="AB0411DE6B09FFADAF89FFFCFDFC3AE2" box="[151,546,1942,1966]" pageId="104" pageNumber="105">
<taxonomicName id="4C144D736B09FFADAF89FFFCFDC03AE2" authority=": Ahrens 2004 a: 324" authorityName="Ahrens" authorityPageNumber="324" authorityYear="2004" box="[151,542,1942,1966]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="egregia">
<emphasis id="B960EAE26B09FFADAF89FFFCFE933AE1" box="[151,333,1942,1966]" italics="true" pageId="104" pageNumber="105">Maladera egregia</emphasis>
:
<treatmentCitation id="0AB510E16B09FFADAE44FFFCFDC03AE2" author="Ahrens, D." box="[346,542,1942,1966]" page="324" pageId="104" pageNumber="105" year="2004">
<bibRefCitation id="EF854B016B09FFADAE44FFFCFDC03AE2" author="Ahrens, D." box="[346,542,1942,1966]" pageId="104" pageNumber="105" refId="ref329734" refString="Ahrens, D. (2004 a) Monographie der Sericini des Himalaya (Coleoptera, Scarabaeidae). Dissertation. de-Verlag im Internet GmbH, Berlin, 534 pp." type="book" year="2004">Ahrens 2004a: 324</bibRefCitation>
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.
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<emphasis id="B960EAE26B09FFADAF89FFB6FE1D3ABA" bold="true" box="[151,451,2012,2038]" pageId="104" pageNumber="105">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6B09FFADAED2FFB6FCF53ABB" ID-GBIF-Occurrence="3034556712" box="[460,811,2012,2039]" country="Myanmar" location="Burma" pageId="104" pageNumber="105" specimenCount="3" specimenCount-male="3" typeStatus="syntype">
<typeStatus id="54AF88526B09FFADAED2FFB6FDE93ABA" box="[460,567,2012,2038]" pageId="104" pageNumber="105" type="syntype">Syntypes</typeStatus>
:
<specimenCount id="9D12FD796B09FFADAD5DFFB6FD5A3ABB" box="[579,644,2012,2039]" pageId="104" pageNumber="105" type="male">3 ♂♂</specimenCount>
“N. E.
<collectingCountry id="F30376606B09FFADADC1FFB7FCF53ABB" box="[735,811,2013,2039]" name="Myanmar" pageId="104" pageNumber="105">Burma</collectingCountry>
</materialsCitation>
<materialsCitation id="3B7C3CAD6B09FFACAC2BFFB6FC2D3DFD" ID-GBIF-Occurrence="3034555771" collectingDate="1934-05-28" collectionCode="BMNH" collectorName="R. Malaise" country="Myanmar" county="Cotype" elevation="2000" lastPageId="105" lastPageNumber="106" location="Malaise" municipality="Malaise" pageId="104" pageNumber="105" specimenCount="1" specimenCount-male="2" typeStatus="syntype">
Kambaiti
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<elevation id="0039D1C36B09FFADACB8FFB6FC233ABB" box="[934,1021,2012,2039]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="104" pageNumber="105" unit="m" value="2000.0">2000 m</elevation>
</quantity>
<date id="FFAA10306B09FFADAB18FFB6FBAB3ABB" box="[1030,1141,2012,2039]" pageId="104" pageNumber="105" value="1934-05-28">
<collectingDate id="EFEEE9D86B09FFADAB18FFB6FBAB3ABB" box="[1030,1141,2012,2039]" pageId="104" pageNumber="105" value="1934-05-28">28.5.1934</collectingDate>
</date>
<location id="8ECB602B6B09FFADAB61FFB6FB093ABA" LSID="urn:lsid:plazi:treatment:03BD87E66B09FFAFAF89FF64FD1E387D:8ECB602B6B09FFADAB61FFB6FB093ABA" box="[1151,1239,2012,2038]" country="Myanmar" name="Malaise" pageId="104" pageNumber="105">Malaise</location>
/ N. E.
<collectingCountry id="F30376606B09FFADAA33FFB7FAA73ABB" box="[1325,1401,2013,2039]" name="Myanmar" pageId="104" pageNumber="105">Burma</collectingCountry>
R. Malaise B. M. 1945-71/ Cotype/
<taxonomicName id="4C144D736B08FFACAD0CF8FDFCCA3DFD" authority="Arrow" authorityName="Arrow" authorityYear="1946" box="[530,788,151,177]" class="Insecta" family="Melolonthidae" genus="Aserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="egregia">
<emphasis id="B960EAE26B08FFACAD0CF8FDFD1B3DFC" box="[530,709,151,177]" italics="true" pageId="105" pageNumber="106">Aserica egregia</emphasis>
Arrow
</taxonomicName>
co-type” (
<collectionCode id="ED05AE356B08FFACAC93F8FDFC393DFD" box="[909,999,151,177]" pageId="105" pageNumber="106">BMNH</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B7C3CAD6B08FFACACE2F8FDFBFA3D99" ID-GBIF-Occurrence="3034556660" collectionCode="BMNH" collectorName="R. Malaise" country="Myanmar" county="Cotype" elevation="2000" location="Kambaiti" municipality="Malaise" pageId="105" pageNumber="106" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D12FD796B08FFACACE2F8FDFBF93DFD" box="[1020,1063,151,177]" pageId="105" pageNumber="106" type="male">1 ♂</specimenCount>
“N. E.
<collectingCountry id="F30376606B08FFACAB9FF8FDFB133DFD" box="[1153,1229,151,177]" name="Myanmar" pageId="105" pageNumber="106">Burma</collectingCountry>
Kambaiti
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<elevation id="0039D1C36B08FFACAA58F8FDFA423DFD" box="[1350,1436,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="105" pageNumber="106" unit="m" value="2000.0">2000 m</elevation>
</quantity>
20/6.1934
<collectorName id="26E153266B08FFACAE10F8D1FE5B3D99" box="[270,389,187,213]" pageId="105" pageNumber="106">R. Malaise</collectorName>
/ N. E.
<collectingCountry id="F30376606B08FFACAECDF8D1FDC13D99" box="[467,543,187,213]" name="Myanmar" pageId="105" pageNumber="106">Burma</collectingCountry>
<collectorName id="26E153266B08FFACAD3BF8D1FD433D99" box="[549,669,187,213]" pageId="105" pageNumber="106">R. Malaise</collectorName>
B. M. 1945-71/ Cotype” (
<collectionCode id="ED05AE356B08FFACACA0F8D1FBC63D99" box="[958,1048,187,213]" pageId="105" pageNumber="106">BMNH</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B7C3CAD6B08FFACAB34F8D1FA593DB5" ID-GBIF-Occurrence="3034556810" collectionCode="BMNH" collectorName="R. Malaise" country="Myanmar" county="Cotype" elevation="2000" location="Kambaiti" municipality="Malaise" pageId="105" pageNumber="106" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D12FD796B08FFACAB34F8D1FB8D3D99" box="[1066,1107,187,213]" pageId="105" pageNumber="106" type="male">1 ♂</specimenCount>
“N. E.
<collectingCountry id="F30376606B08FFACABBAF8D1FB2E3D99" box="[1188,1264,187,213]" name="Myanmar" pageId="105" pageNumber="106">Burma</collectingCountry>
<location id="8ECB602B6B08FFACABE8F8D1FA803D99" LSID="urn:lsid:plazi:treatment:03BD87E66B09FFAFAF89FF64FD1E387D:8ECB602B6B08FFACABE8F8D1FA803D99" box="[1270,1374,187,213]" country="Myanmar" county="Cotype" municipality="Malaise" name="Kambaiti" pageId="105" pageNumber="106">Kambaiti</location>
<quantity id="4CEC9B156B08FFACAA7AF8D1FF733DB5" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="105" pageNumber="106" unit="m" value="2000.0">
<elevation id="0039D1C36B08FFACAA7AF8D1FF733DB5" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="105" pageNumber="106" unit="m" value="2000.0">2000 m</elevation>
</quantity>
4/6.1934
<collectingMunicipality id="6BCFAC8A6B08FFACAE00F8B5FEA83DB5" box="[286,374,223,249]" pageId="105" pageNumber="106">Malaise</collectingMunicipality>
/ N. E.
<collectingCountry id="F30376606B08FFACAED9F8B5FDCD3DB5" box="[455,531,223,249]" name="Myanmar" pageId="105" pageNumber="106">Burma</collectingCountry>
<collectorName id="26E153266B08FFACAD04F8B5FD4D3DB5" box="[538,659,223,249]" pageId="105" pageNumber="106">R. Malaise</collectorName>
B. M. 1945-71/
<collectingCounty id="62CA4E7C6B08FFACAC50F8B5FC403DB5" box="[846,926,223,249]" pageId="105" pageNumber="106">Cotype</collectingCounty>
/
<taxonomicName id="4C144D736B08FFACACB3F8B5FB703DB5" authority="Arrow" authorityName="Arrow" authorityYear="1946" box="[941,1198,223,249]" class="Insecta" family="Melolonthidae" genus="Aserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="egregia">
<emphasis id="B960EAE26B08FFACACB3F8B5FB813DB4" box="[941,1119,223,249]" italics="true" pageId="105" pageNumber="106">Aserica egregia</emphasis>
Arrow
</taxonomicName>
co-type” (
<collectionCode id="ED05AE356B08FFACAA3BF8B5FAA13DB5" box="[1317,1407,223,249]" pageId="105" pageNumber="106">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6B66786B08FFACAF89FF06FF0F3ABC" ID-DOI="http://doi.org/10.5281/zenodo.4496362" ID-Zenodo-Dep="4496362" httpUri="https://zenodo.org/record/4496362/files/figure.png" pageId="105" pageNumber="106" startId="105.[151,250,1900,1924]" targetBox="[213,1369,283,1871]" targetPageId="105">
<paragraph id="8BAB36F06B08FFACAF89FF06FF0F3ABC" blockId="105.[151,1437,1900,2032]" pageId="105" pageNumber="106">
<emphasis id="B960EAE26B08FFACAF89FF06FEBF3AC8" bold="true" box="[151,353,1900,1924]" pageId="105" pageNumber="106">FIGURE 22. AD:</emphasis>
<taxonomicName id="4C144D736B08FFACAE74FF06FD633AC8" authority="(Arrow, 1946)" baseAuthorityName="Arrow" baseAuthorityYear="1946" box="[362,701,1900,1924]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="egregia">
<emphasis id="B960EAE26B08FFACAE74FF06FDFC3AC8" box="[362,546,1900,1924]" italics="true" pageId="105" pageNumber="106">Maladera egregia</emphasis>
(Arrow, 1946)
</taxonomicName>
(syntype: Myanmar: Kambaiti);
<emphasis id="B960EAE26B08FFACAB0DFF06FB9C3AC8" bold="true" box="[1043,1090,1900,1924]" pageId="105" pageNumber="106">EI:</emphasis>
<taxonomicName id="4C144D736B08FFACAB55FF07FAFD3AC8" authority="(Frey, 1972)" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1099,1315,1900,1924]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="B960EAE26B08FFACAB55FF07FB423AC8" box="[1099,1180,1900,1924]" italics="true" pageId="105" pageNumber="106">M. levis</emphasis>
(Frey, 1972)
</taxonomicName>
(holotype);
<emphasis id="B960EAE26B08FFACAF89FFFAFF0F3AE4" bold="true" box="[151,209,1936,1960]" pageId="105" pageNumber="106">JM:</emphasis>
<taxonomicName id="4C144D736B08FFACAFC6FFFBFE0E3AE4" authority="Ahrens, 2004" authorityName="Ahrens" authorityYear="2004" box="[216,464,1936,1960]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="hmong">
<emphasis id="B960EAE26B08FFACAFC6FFFBFE9E3AE4" box="[216,320,1936,1960]" italics="true" pageId="105" pageNumber="106">M. hmong</emphasis>
Ahrens, 2004
</taxonomicName>
(Paratype: Vietnam: Sa Pa).
<emphasis id="B960EAE26B08FFACADE4FFFAFC903AE4" bold="true" box="[762,846,1936,1960]" pageId="105" pageNumber="106">A, E, J:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B960EAE26B08FFACABBEFFFAFB243AE4" bold="true" box="[1184,1274,1936,1960]" pageId="105" pageNumber="106">C, H, L:</emphasis>
aedeagus, right side lateral view;
<emphasis id="B960EAE26B08FFACAE54FFDEFE433A80" bold="true" box="[330,413,1972,1996]" pageId="105" pageNumber="106">B, F, K:</emphasis>
parameres, dorsal view;
<emphasis id="B960EAE26B08FFACAD85FFDEFD693A80" bold="true" box="[667,695,1972,1996]" pageId="105" pageNumber="106">G:</emphasis>
parameres, ventral view;
<emphasis id="B960EAE26B08FFACACA3FFDEFBCD3A80" bold="true" box="[957,1043,1972,1996]" pageId="105" pageNumber="106">D, I, M:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<subSubSection id="C30E657B6B0BFFAFAFD9F8FCFD763989" pageId="106" pageNumber="107" type="description">
<paragraph id="8BAB36F06B0BFFAFAFD9F8FCFAD83D99" blockId="106.[151,1437,150,1329]" pageId="106" pageNumber="107">
<emphasis id="B960EAE26B0BFFAFAFD9F8FCFEAB3DFC" bold="true" box="[199,373,150,176]" pageId="106" pageNumber="107">Redescription.</emphasis>
Length:
<quantity id="4CEC9B156B0BFFAFAEC2F8FDFDEB3DFD" box="[476,565,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="106" pageNumber="107" unit="mm" value="8.6">8.6 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156B0BFFAFADE0F8FDFC893DFD" box="[766,855,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="106" pageNumber="107" unit="mm" value="6.4">6.4 mm</quantity>
, width:
<quantity id="4CEC9B156B0BFFAFACADF8FCFBD33DFD" box="[947,1037,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="106" pageNumber="107" unit="mm" value="5.6">5.6 mm</quantity>
. Body oblong-oval, reddish brown, antenna yellow, iridescent and shiny, frons and pronotum somewhat dull, dorsal face nearly glabrous.
</paragraph>
<paragraph id="8BAB36F06B0BFFAFAFD9F8B5FB973CB9" blockId="106.[151,1437,150,1329]" pageId="106" pageNumber="107">
Labroclypeus wide and subtrapezoidal, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles moderately rounded, anterior margin distinctly emarginate, margins weakly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex, finely, densely punctate, with a few long setae anteriorly; frontoclypeal suture finely incised, weakly curved; smooth area anterior to eye convex, 1.5 times as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely densely punctate, without terminal setae. Frons with sparse and fine punctures, with a few fine setae beside eyes. Eyes small, ratio diameter/ interocular width: 0.52. Antenna with ten antennomeres; club with three antennomeres and straight, as long as remaining antennomeres combined.
<collectingRegion id="49D0F8126B0BFFAFAD07F9B1FDA63CB9" box="[537,632,475,501]" country="Myanmar" name="Chin" pageId="106" pageNumber="107">Mentum</collectingRegion>
elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06B0BFFAFAFD9F995FCAB3FE5" blockId="106.[151,1437,150,1329]" pageId="106" pageNumber="107">Pronotum transverse, widest at base, lateral margins evenly convex and weakly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin convex, with fine marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; lateral margin sparsely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures, impunctate medially.</paragraph>
<paragraph id="8BAB36F06B0BFFAFAFD9FAD9FF0A3E12" blockId="106.[151,1437,150,1329]" pageId="106" pageNumber="107">Elytra widest in posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, moderately dense punctures concentrated along striae, with minute setae in punctures, penultimate lateral interval with a few fine setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification).</paragraph>
<paragraph id="8BAB36F06B0BFFAFAFD9FB0DFBE13E85" blockId="106.[151,1437,150,1329]" pageId="106" pageNumber="107">
Ventral surface dull, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.49. Abdomen in available
<typeStatus id="54AF88526B0BFFAFAC0FFBC5FCB63E85" box="[785,872,943,969]" pageId="106" pageNumber="107" type="syntype">syntype</typeStatus>
specimen missing.
</paragraph>
<paragraph id="8BAB36F06B0BFFAFAFD9FBB9FE7839ED" blockId="106.[151,1437,150,1329]" pageId="106" pageNumber="107">Legs moderately long and narrow, shiny; femora with two longitudinal rows of setae, coarsely and moderately densely punctate. Metafemur dull, anterior margin acute, with adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
<paragraph id="8BAB36F06B0BFFAFAFD9FCC1FD763989" blockId="106.[151,1437,150,1329]" box="[199,680,1195,1221]" pageId="106" pageNumber="107">
Aedeagus:
<figureCitation id="132F2A756B0BFFAFAE5AFCC1FE4B3989" box="[324,405,1195,1221]" captionStart="FIGURE 22" captionStartId="105.[151,250,1900,1924]" captionTargetBox="[213,1369,283,1871]" captionTargetId="figure-75@105.[202,1385,273,1876]" captionTargetPageId="105" captionText="FIGURE 22. AD: Maladera egregia (Arrow, 1946) (syntype: Myanmar: Kambaiti); EI: M. levis (Frey, 1972) (holotype); JM: M. hmong Ahrens, 2004 (Paratype: Vietnam: Sa Pa). A, E, J: aedeagus, left side lateral view; C, H, L: aedeagus, right side lateral view; B, F, K: parameres, dorsal view; G: parameres, ventral view; D, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496362" httpUri="https://zenodo.org/record/4496362/files/figure.png" pageId="106" pageNumber="107">Fig. 22</figureCitation>
AC. Habitus:
<figureCitation id="132F2A756B0BFFAFAD23FCC1FD7D3989" box="[573,675,1195,1221]" captionStart="FIGURE 22" captionStartId="105.[151,250,1900,1924]" captionTargetBox="[213,1369,283,1871]" captionTargetId="figure-75@105.[202,1385,273,1876]" captionTargetPageId="105" captionText="FIGURE 22. AD: Maladera egregia (Arrow, 1946) (syntype: Myanmar: Kambaiti); EI: M. levis (Frey, 1972) (holotype); JM: M. hmong Ahrens, 2004 (Paratype: Vietnam: Sa Pa). A, E, J: aedeagus, left side lateral view; C, H, L: aedeagus, right side lateral view; B, F, K: parameres, dorsal view; G: parameres, ventral view; D, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496362" httpUri="https://zenodo.org/record/4496362/files/figure.png" pageId="106" pageNumber="107">Fig. 22D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B0BFFAFAFD9FCA5FEC53841" pageId="106" pageNumber="107" type="discussion">
<paragraph id="8BAB36F06B0BFFAFAFD9FCA5FEC53841" blockId="106.[151,1437,150,1329]" pageId="106" pageNumber="107">
<emphasis id="B960EAE26B0BFFAFAFD9FCA5FEE539A5" bold="true" box="[199,315,1231,1257]" pageId="106" pageNumber="107">Remarks.</emphasis>
The metatibiae were lacking in the
<typeStatus id="54AF88526B0BFFAFADCBFCA5FCF239A5" box="[725,812,1231,1257]" pageId="106" pageNumber="107" type="syntype">syntype</typeStatus>
specimen we examined and are thus omitted in the redescription.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B0BFFAFAFD9FD7CFD1E387D" box="[199,704,1302,1329]" pageId="106" pageNumber="107" type="distribution">
<paragraph id="8BAB36F06B0BFFAFAFD9FD7CFD1E387D" blockId="106.[151,1437,150,1329]" box="[199,704,1302,1329]" pageId="106" pageNumber="107">
<emphasis id="B960EAE26B0BFFAFAFD9FD7CFEBF387C" bold="true" box="[199,353,1302,1328]" pageId="106" pageNumber="107">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756B0BFFAFAEC8FD7DFDF6387D" box="[470,552,1303,1329]" captionStart="FIGURE 97" captionStartId="350.[151,250,1034,1058]" captionTargetBox="[208,1378,187,1003]" captionTargetId="figure-19@350.[202,1386,181,1010]" captionTargetPageId="350" captionText="FIGURE 97. Distribution of the Maladera egregia group: Maladera diaolinensis sp. n., M. egregia (Arrow, 1946), M. gusakovi sp. n., M. hmong Ahrens, 2004, M. hubeiensis sp. n., M. laocaiensis sp. n., M. levis (Frey, 1972)." figureDoi="http://doi.org/10.5281/zenodo.4496578" httpUri="https://zenodo.org/record/4496578/files/figure.png" pageId="106" pageNumber="107">Fig. 97</figureCitation>
) and
<tableCitation id="C696034B6B0BFFAFAD78FD7DFD64387D" box="[614,698,1303,1329]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x records from material examined herein, o records from literature, with no material of this study)." pageId="106" pageNumber="107">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>