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<document ID-DOI="10.11646/zootaxa.5277.1.4" ID-GBIF-Dataset="ad5ba4d1-c2ec-42e3-a76d-a5c9b395a6da" ID-ISSN="1175-5326" ID-Zenodo-Dep="7892052" ID-ZooBank="272D07DF-8471-4ADE-9689-DB8F588ACF35" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1682930765663" checkinUser="plazi" docAuthor="Short, Andrew E. Z., Torres-Gavosto, Juan Martin &amp; Hettinger, Ty" docDate="2023" docId="03BD87B6FFA1F42ECFDB7026FC28FA1F" docLanguage="en" docName="zootaxa.5277.1.4.pdf" docOrigin="Zootaxa 5277 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oocyclus morgani Garcia-Hernandez 2009" docType="treatment" docVersion="3" lastPageNumber="111" masterDocId="FF84FFCEFFB5F43ACF4C7204FFE1FFDE" masterDocTitle="A review of the Oocyclus Sharp of Ecuador with description of 12 new species (Coleoptera: Hydrophilidae: Laccobiini)" masterLastPageNumber="112" masterPageNumber="91" pageNumber="111" updateTime="1683122599468" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>A review of the Oocyclus Sharp of Ecuador with description of 12 new species (Coleoptera: Hydrophilidae: Laccobiini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Short, Andrew E. Z.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7467-7116</mods:nameIdentifier>
<mods:affiliation>Division of Entomology, Biodiversity Institute &amp; Department of Ecology and Evolutionary Biology, University of Kansas, 1501 Crestline Drive, Suite 140, Lawrence, KS 66045, U. S. A. &amp; aezshort @ gmail. com; https: // orcid. org / 0000 - 0002 - 7467 - 7116</mods:affiliation>
<mods:nameIdentifier type="email">aezshort@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Torres-Gavosto, Juan Martin</mods:namePart>
<mods:nameIdentifier type="ORCID">0009-0007-9274-2923</mods:nameIdentifier>
<mods:affiliation>Division of Entomology, Biodiversity Institute &amp; Department of Ecology and Evolutionary Biology, University of Kansas, 1501 Crestline Drive, Suite 140, Lawrence, KS 66045, U. S. A. &amp; https: // orcid. org / 0009 - 0007 - 9274 - 2923</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hettinger, Ty</mods:namePart>
<mods:affiliation>Division of Entomology, Biodiversity Institute &amp; Department of Ecology and Evolutionary Biology, University of Kansas, 1501 Crestline Drive, Suite 140, Lawrence, KS 66045, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5277</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>91</mods:start>
<mods:end>112</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5277.1.4</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5277.1.4</mods:identifier>
<mods:identifier type="GBIF-Dataset">ad5ba4d1-c2ec-42e3-a76d-a5c9b395a6da</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7892052</mods:identifier>
<mods:identifier type="ZooBank">272D07DF-8471-4ADE-9689-DB8F588ACF35</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7892100" ID-Zenodo-Dep="7892100" LSID="urn:lsid:plazi:treatment:03BD87B6FFA1F42ECFDB7026FC28FA1F" httpUri="http://treatment.plazi.org/id/03BD87B6FFA1F42ECFDB7026FC28FA1F" lastPageNumber="111" pageId="20" pageNumber="111">
<subSubSection box="[151,545,546,573]" pageId="20" pageNumber="111" type="nomenclature">
<paragraph blockId="20.[151,545,546,609]" box="[151,545,546,573]" pageId="20" pageNumber="111">
<heading bold="true" box="[151,545,546,573]" fontSize="11" level="1" pageId="20" pageNumber="111" reason="1">
<emphasis bold="true" box="[151,545,546,573]" pageId="20" pageNumber="111">
<taxonomicName ID-CoL="49RWB" authorityName="Garcia-Hernandez" authorityYear="2009" box="[151,464,546,573]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="111" phylum="Arthropoda" rank="species" species="morgani">
<emphasis bold="true" box="[151,464,546,573]" italics="true" pageId="20" pageNumber="111">Oocyclus morgani/trujillo</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,330,583,609]" pageId="20" pageNumber="111" type="description">
<paragraph blockId="20.[151,545,546,609]" box="[151,330,583,609]" pageId="20" pageNumber="111">
(
<figureCitation box="[159,257,583,609]" captionStart="FIGURE 4" captionStartId="13.[151,250,1523,1547]" captionTargetBox="[151,1436,181,1505]" captionTargetId="figure-20@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 4. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) lepidus sp. n. (B) morgani/trujillo group" figureDoi="http://doi.org/10.5281/zenodo.7892068" httpUri="https://zenodo.org/record/7892068/files/figure.png" pageId="20" pageNumber="111">Figs. 4B</figureCitation>
,
<figureCitation box="[269,295,583,609]" captionStart="FIGURE 6" captionStartId="17.[151,250,1605,1629]" captionTargetBox="[157,1427,187,1586]" captionTargetId="figure-20@17.[151,1437,181,1587]" captionTargetPageId="17" captionText="FIGURE 6. Aedeagi of Oocyclus species in Ecuador with angulate or spinose posterolateral corners of the pronotum. (A) ustulatus sp. n. (B) radiatus sp. n. (C) pakcha sp. n. (D) sumak sp. n. (E) ankas sp. n. (F) lepidus sp. n. (G) bellus sp. n. (H) zamora sp. n. (I) specimen of the morgani/trujillo species group (J) specimen of the morgani/trujillo species group." figureDoi="http://doi.org/10.5281/zenodo.7892076" httpUri="https://zenodo.org/record/7892076/files/figure.png" pageId="20" pageNumber="111">6I</figureCitation>
,
<figureCitation box="[307,322,583,609]" captionStart="FIGURE 7" captionStartId="19.[151,250,1886,1910]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-20@19.[151,1436,181,1868]" captionTargetPageId="19" captionText="FIGURE 7. Distribution of Oocyclus spp. in Ecuador." figureDoi="http://doi.org/10.5281/zenodo.7892078" httpUri="https://zenodo.org/record/7892078/files/figure.png" pageId="20" pageNumber="111">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="111" type="materials_examined">
<paragraph blockId="20.[151,1437,654,1473]" pageId="20" pageNumber="111">
<emphasis bold="true" box="[151,504,654,680]" pageId="20" pageNumber="111">
Material Examined (
<specimenCount box="[402,490,654,680]" pageId="20" pageNumber="111" type="generic">72 exs.</specimenCount>
):
</emphasis>
<materialsCitation collectingDate="1979-08-21" collectionCode="USNM" collectorName="J. Anderson" country="Ecuador" elevation="1510" location="San Rafael Falls" municipality="Just" pageId="20" pageNumber="111" specimenCount="3" stateProvince="Napo">
<emphasis bold="true" box="[513,860,654,681]" pageId="20" pageNumber="111">
<collectingCountry box="[513,659,654,680]" name="Ecuador" pageId="20" pageNumber="111">ECUADOR</collectingCountry>
:
<collectingRegion box="[672,855,654,681]" country="Ecuador" name="Napo" pageId="20" pageNumber="111">Napo Province</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[869,913,655,681]" pageId="20" pageNumber="111">Just</collectingMunicipality>
above
<location LSID="urn:lsid:plazi:treatment:03BD87B6FFA1F42ECFDB7026FC28FA1F:8ECB607BFFA1F42ECCA9708BFB42FD77" box="[997,1187,655,681]" country="Ecuador" municipality="Just" name="San Rafael Falls" pageId="20" pageNumber="111" stateProvince="Napo">San Rafael Falls</location>
,
<quantity box="[1200,1280,654,681]" metricMagnitude="3" metricUnit="m" metricValue="1.51" pageId="20" pageNumber="111" unit="m" value="1510.0">
<elevation box="[1200,1280,654,681]" metricMagnitude="3" metricUnit="m" metricValue="1.51" pageId="20" pageNumber="111" unit="m" value="1510.0">1510m</elevation>
</quantity>
,
<date box="[1294,1432,655,681]" pageId="20" pageNumber="111" value="1979-08-21">
<collectingDate box="[1294,1432,655,681]" pageId="20" pageNumber="111" value="1979-08-21">21.viii.1979</collectingDate>
</date>
, leg.
<collectorName box="[201,337,691,717]" pageId="20" pageNumber="111">J. Anderson</collectorName>
, cascade-seep (
<specimenCount box="[513,581,691,717]" pageId="20" pageNumber="111" type="generic">3 exs.</specimenCount>
,
<collectionCode box="[594,680,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="111" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation collectingDate="1976-01-25" collectionCode="USNM" collectorName="P. Spangler" country="Ecuador" location="18 km E Banos" pageId="20" pageNumber="111" specimenCount="39" stateProvince="Tungurahua">
<emphasis bold="true" box="[700,971,690,717]" pageId="20" pageNumber="111">
<collectingRegion box="[700,966,690,717]" country="Ecuador" name="Tungurahua" pageId="20" pageNumber="111">Tungurahua Province</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:03BD87B6FFA1F42ECFDB7026FC28FA1F:8ECB607BFFA1F42ECC9F70B7FB64FD13" box="[979,1157,691,717]" country="Ecuador" name="18 km E Banos" pageId="20" pageNumber="111" stateProvince="Tungurahua">
<quantity box="[979,1051,691,717]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="20" pageNumber="111" unit="km" value="18.0">18 km</quantity>
E Baños
</location>
, 5130′,
<date box="[1246,1354,690,717]" pageId="20" pageNumber="111" value="1976-01-25">
<collectingDate box="[1246,1354,690,717]" pageId="20" pageNumber="111" value="1976-01-25">25.i.1976</collectingDate>
</date>
, leg.
<collectorName pageId="20" pageNumber="111">P. Spangler</collectorName>
<emphasis box="[257,314,727,753]" italics="true" pageId="20" pageNumber="111">et al.</emphasis>
, seepage over rock outcrops in roadcuts, (
<specimenCount box="[784,867,727,753]" pageId="20" pageNumber="111" type="generic">39 exs.</specimenCount>
,
<collectionCode box="[878,964,727,753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="111" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1977-05-04" collectionCode="USNM" collectorName="P. Spangler &amp; D. Givens" country="Ecuador" location="Banos" pageId="20" pageNumber="111" specimenCount="3" stateProvince="Tungurahua">
<quantity box="[983,1040,726,753]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="111" unit="km" value="5.0">5 km</quantity>
S
<location LSID="urn:lsid:plazi:treatment:03BD87B6FFA1F42ECFDB7026FC28FA1F:8ECB607BFFA1F42ECB6170D3FB97FD2F" box="[1069,1142,727,753]" country="Ecuador" name="Banos" pageId="20" pageNumber="111" stateProvince="Tungurahua">Baños</location>
,
<date box="[1152,1251,727,753]" pageId="20" pageNumber="111" value="1977-05-04">
<collectingDate box="[1152,1251,727,753]" pageId="20" pageNumber="111" value="1977-05-04">4.v.1977</collectingDate>
</date>
, leg.
<collectorName box="[1310,1437,727,753]" pageId="20" pageNumber="111">P. Spangler</collectorName>
&amp;
<collectorName box="[183,298,763,789]" pageId="20" pageNumber="111">D. Givens</collectorName>
(
<specimenCount box="[317,388,763,789]" pageId="20" pageNumber="111" type="generic">3 exs.</specimenCount>
,
<collectionCode box="[402,488,763,789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="111" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1976-01-28" collectionCode="USNM" collectorName="P. Spangler" country="Ecuador" location="Banos" pageId="20" pageNumber="111" specimenCount="6" stateProvince="Tungurahua">
<quantity box="[509,583,763,789]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="111" unit="km" value="20.0">20 km</quantity>
E
<location LSID="urn:lsid:plazi:treatment:03BD87B6FFA1F42ECFDB7026FC28FA1F:8ECB607BFFA1F42ECD2770FFFD55FCCB" box="[619,692,763,789]" country="Ecuador" name="Banos" pageId="20" pageNumber="111" stateProvince="Tungurahua">Baños</location>
,
<date box="[706,814,763,789]" pageId="20" pageNumber="111" value="1976-01-28">
<collectingDate box="[706,814,763,789]" pageId="20" pageNumber="111" value="1976-01-28">28.i.1976</collectingDate>
</date>
, leg.
<collectorName box="[879,1008,763,789]" pageId="20" pageNumber="111">P. Spangler</collectorName>
<emphasis box="[1017,1075,763,789]" italics="true" pageId="20" pageNumber="111">et al.</emphasis>
, seepage over rock outcrops in roadcuts (
<specimenCount box="[261,330,799,825]" pageId="20" pageNumber="111" type="generic">6 exs.</specimenCount>
,
<collectionCode box="[341,427,799,825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="111" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1976-01-27" collectionCode="USNM" collectorName="P. Spangler" country="Ecuador" location="Banos" pageId="20" pageNumber="111" specimenCount="20" stateProvince="Tungurahua">
<location LSID="urn:lsid:plazi:treatment:03BD87B6FFA1F42ECFDB7026FC28FA1F:8ECB607BFFA1F42ECEF2711BFDE6FCE7" box="[446,519,799,825]" country="Ecuador" name="Banos" pageId="20" pageNumber="111" stateProvince="Tungurahua">Baños</location>
,
<date box="[529,638,799,825]" pageId="20" pageNumber="111" value="1976-01-27">
<collectingDate box="[529,638,799,825]" pageId="20" pageNumber="111" value="1976-01-27">27.i.1976</collectingDate>
</date>
, leg.
<collectorName box="[697,823,799,825]" pageId="20" pageNumber="111">P. Spangler</collectorName>
<emphasis box="[830,886,799,825]" italics="true" pageId="20" pageNumber="111">et al.</emphasis>
, seepage over rock outcrops in roadcuts (
<specimenCount box="[1349,1431,799,825]" pageId="20" pageNumber="111" type="generic">20 exs.</specimenCount>
,
<collectionCode box="[151,237,835,861]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="111" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation box="[256,943,835,861]" collectingDate="1977-05-20" collectionCode="USNM" collectorName="W. Steiner" country="Ecuador" location="Rio Negro" pageId="20" pageNumber="111" specimenCount="1" stateProvince="Tungurahua">
<quantity box="[256,313,835,861]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="111" unit="km" value="6.0">6 km</quantity>
E
<location LSID="urn:lsid:plazi:treatment:03BD87B6FFA1F42ECFDB7026FC28FA1F:8ECB607BFFA1F42ECE147147FE30FC83" box="[344,465,835,861]" country="Ecuador" name="Rio Negro" pageId="20" pageNumber="111" stateProvince="Tungurahua">Rio Negro</location>
,
<date box="[476,588,835,861]" pageId="20" pageNumber="111" value="1977-05-20">
<collectingDate box="[476,588,835,861]" pageId="20" pageNumber="111" value="1977-05-20">20.v.1977</collectingDate>
</date>
, leg.
<collectorName box="[647,764,835,861]" pageId="20" pageNumber="111">W. Steiner</collectorName>
(
<specimenCount box="[783,837,835,861]" pageId="20" pageNumber="111" type="generic">1 ex.</specimenCount>
,
<collectionCode box="[849,935,835,861]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="111" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="111" type="diagnosis">
<paragraph blockId="20.[151,1437,654,1473]" pageId="20" pageNumber="111">
<emphasis bold="true" box="[199,322,870,896]" pageId="20" pageNumber="111">Diagnosis.</emphasis>
Smaller sized species (Length =
<quantity box="[679,819,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="4.1" metricValueMin="3.1" pageId="20" pageNumber="111" unit="mm" value="3.5999999999999996" valueMax="4.1" valueMin="3.1">3.1–4.1 mm</quantity>
.). Posterolateral corners of pronotum spinose. Prosternal carinae set with a pair of spines anteriorly. Elytra with first row of systematic punctures bearing short, linear clumps of dense erect setae, giving it a “tufted” appearance. Elytral margins not explanate, without a fringe of long setae. Abdominal ventrites brown.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="111" type="discussion">
<paragraph blockId="20.[151,1437,654,1473]" pageId="20" pageNumber="111">
<emphasis bold="true" box="[199,315,1015,1041]" pageId="20" pageNumber="111">Remarks.</emphasis>
There are several series of specimens in the material we examined (mostly from the Baños area of central
<collectingCountry box="[234,329,1051,1077]" name="Ecuador" pageId="20" pageNumber="111">Ecuador</collectingCountry>
) that represent at least one additional species. Indeed, we had originally thought there might be two closely related species among this batch of material. There is no question they are different than any of the other 13 Ecuadorian species enumerated in this manuscript. However, they compare well with the descriptions of
<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2010" box="[1324,1436,1123,1149]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="111" phylum="Arthropoda" rank="species" species="trujillo">
<emphasis box="[1324,1436,1123,1149]" italics="true" pageId="20" pageNumber="111">O. trujillo</emphasis>
</taxonomicName>
from
<collectingCountry box="[212,327,1159,1185]" name="Venezuela" pageId="20" pageNumber="111">Venezuela</collectingCountry>
and
<taxonomicName authorityName="Garcia-Hernandez" authorityYear="2009" box="[381,509,1159,1185]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="111" phylum="Arthropoda" rank="species" species="morgani">
<emphasis box="[381,509,1159,1185]" italics="true" pageId="20" pageNumber="111">O. morgani</emphasis>
</taxonomicName>
from
<collectingCountry box="[576,690,1159,1185]" name="Colombia" pageId="20" pageNumber="111">Colombia</collectingCountry>
. It should be noted that we have examined specimens of
<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2010" box="[1324,1436,1159,1185]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="111" phylum="Arthropoda" rank="species" species="trujillo">
<emphasis box="[1324,1436,1159,1185]" italics="true" pageId="20" pageNumber="111">O. trujillo</emphasis>
</taxonomicName>
but only seen images of the dorsum and side of the pronotum of
<taxonomicName authorityName="Garcia-Hernandez" authorityYear="2009" box="[872,1000,1195,1221]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="111" phylum="Arthropoda" rank="species" species="morgani">
<emphasis box="[872,1000,1195,1221]" italics="true" pageId="20" pageNumber="111">O. morgani</emphasis>
</taxonomicName>
. Both these two named species as well as the series from
<collectingCountry box="[354,447,1231,1257]" name="Ecuador" pageId="20" pageNumber="111">Ecuador</collectingCountry>
are of similar size and share many of the same characters: spinose posterolateral corners of the pronotum, pale anterolateral corners of the pronotum, similar conditions of the pro and mesoventrites, and most distinctively, having the elytral series of punctures appearing as a series of short linear tufts of erect setae. There are some observable difference in the precise distribution of these tufts, but other species have shown broad conspecific variation in this feature (e.g.
<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2010" box="[616,730,1375,1401]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="111" phylum="Arthropoda" rank="species" species="floccus">
<emphasis box="[616,730,1375,1401]" italics="true" pageId="20" pageNumber="111">O. floccus</emphasis>
</taxonomicName>
, see
<bibRefCitation author="Santana, L. &amp; Short, A. E. Z. &amp; Benetti, C. J. &amp; Hamada, N." box="[789,1012,1375,1401]" pageId="20" pageNumber="111" pagination="587 - 594" refId="ref16821" refString="Santana, L., Short, A. E. Z., Benetti, C. J. &amp; Hamada, N. (2023) New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera: Hydrophilidae) from the Guiana Shield. Zootaxa, 5230 (5), 587 - 594. https: // doi. org / 10.11646 / zootaxa. 5230.5.6" type="journal article" year="2023">
Santana
<emphasis box="[886,943,1375,1401]" italics="true" pageId="20" pageNumber="111">et al.</emphasis>
2023
</bibRefCitation>
). While we believe it is very possible that these Ecuadorian specimens represent one or more new species, we here take a conservative approach until the differences between
<taxonomicName authorityName="Garcia-Hernandez" authorityYear="2009" box="[382,511,1447,1473]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="111" phylum="Arthropoda" rank="species" species="morgani">
<emphasis box="[382,511,1447,1473]" italics="true" pageId="20" pageNumber="111">O. morgani</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2010" box="[565,677,1447,1473]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="111" phylum="Arthropoda" rank="species" species="trujillo">
<emphasis box="[565,677,1447,1473]" italics="true" pageId="20" pageNumber="111">O. trujillo</emphasis>
</taxonomicName>
can be better illuminated.
</paragraph>
</subSubSection>
</treatment>
</document>
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