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<document ID-DOI="http://doi.org/10.5281/zenodo.3739808" ID-GBIF-Dataset="b7e93c79-44a6-4d9c-8b78-dcb62033fb0d" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="3739808" approvalRequired="13" approvalRequired_for_treatments="13" checkinTime="1585778501182" checkinUser="plazi" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="03BD87A2C665A217F87DF82FF6FE5327" docLanguage="en" docName="hbmw_9_Hipposideridae_210.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Hipposideros coxi Shelford 1901" docType="treatment" docVersion="16" lastPageNumber="246" masterDocId="FF84FFDAC676A204FFF8FF9AFFEF4346" masterDocTitle="Hipposideridae" masterLastPageNumber="258" masterPageNumber="227" pageNumber="246" updateTime="1658320025573" updateUser="diego">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Hipposideridae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>227</mods:start>
<mods:end>258</mods:end>
</mods:extent>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3739808</mods:identifier>
<mods:identifier type="GBIF-Dataset">b7e93c79-44a6-4d9c-8b78-dcb62033fb0d</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">3739808</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3810859" ID-GBIF-Taxon="163433788" ID-Zenodo-Dep="3810859" LSID="urn:lsid:plazi:treatment:03BD87A2C665A217F87DF82FF6FE5327" httpUri="http://treatment.plazi.org/id/03BD87A2C665A217F87DF82FF6FE5327" lastPageNumber="246" pageId="19" pageNumber="246">
<heading pageId="19" pageNumber="246">
<subSubSection box="[1925,2005,1973,2034]" pageId="19" pageNumber="246" type="multiple">
<paragraph blockId="19.[1923,3137,1973,2130]" box="[1925,2005,1973,2034]" pageId="19" pageNumber="246">
<figureCitation box="[1925,2005,1973,2034]" captionStart="Plate 18: Hipposideridae" captionStartId="18.[172,215,4473,4502]" captionTargetBox="[0,3699,0,4866]" captionTargetPageId="17" captionText="47. Short-tailed Leaf-nosed Bat (Hipposideros curtus), 49. Aellens Leaf-nosed Bat (Hipposideros marisae), 50. Schneiders Leaf-nosed Bat (Hipposideros speoris), 51. Spurred Leaf-nosed Bat (Hipposideros calcaratus), 52. Fawn-colored Leaf-nosed Bat (Hipposideros ceruinus), 53. Coxs Leaf-nosed Bat (Hipposideros coxi), 54. Cantors Leaf-nosed Bat (Hipposideros galeritus), 55. Aba Leaf-nosed Bat (Hipposideros abae), 56. Benito Leaf-nosed Bat (Hipposideros beatus), 57. Sundevalls Leaf-nosed Bat (Hipposideros caffer), 58. Sooty Leaf-nosed (Bat Hipposiderosfuliginosus), 59. Lamottes Leaf-nosed Bat (Hipposideros lamottei), 60. Noacks Leaf-nosed Bat (Hipposideros ruber), 61. Maghreb Leaf-nosed Bat (Hipposideros tephrus), 62. Dusky Leaf-nosed Bat (Hipposideros ater), 63. Bicolored Leaf-nosed Bat (Hipposideros bicolor), 64. Short-headed Leaf-nosed Bat (Hipposideros breviceps), 65. Least Leaf-nosed Bat (Hipposideros cineraceus), 66. Large Mindanao Leaf-nosed Bat (Hipposideros coronatus)" figureDoi="http://doi.org/10.5281/zenodo.3739814" httpUri="https://zenodo.org/record/3739814/files/figure.png" pageId="19" pageNumber="246">53.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[2030,2620,1973,2034]" pageId="19" pageNumber="246" type="vernacular_names">
<paragraph blockId="19.[1923,3137,1973,2130]" box="[2030,2620,1973,2034]" pageId="19" pageNumber="246">
<vernacularName box="[2030,2620,1973,2034]" pageId="19" pageNumber="246">Coxs Leaf-nosed Bat</vernacularName>
</paragraph>
</subSubSection>
<subSubSection box="[2684,3124,1973,2034]" pageId="19" pageNumber="246" type="nomenclature">
<paragraph blockId="19.[1923,3137,1973,2130]" box="[2684,3124,1973,2034]" pageId="19" pageNumber="246">
<taxonomicName authorityName="Shelford" authorityYear="1901" box="[2684,3124,1973,2034]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="246" phylum="Chordata" rank="species" species="coxi">
<emphasis box="[2684,3124,1973,2034]" italics="true" pageId="19" pageNumber="246">Hipposideros coxi</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="246" type="vernacular_names">
<paragraph blockId="19.[1923,3137,1973,2130]" box="[1925,3136,2050,2077]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[1925,2027,2052,2077]" pageId="19" pageNumber="246">French:</emphasis>
<vernacularName box="[2039,2291,2050,2077]" pageId="19" pageNumber="246">Phyllorhine de Cox</vernacularName>
/
<emphasis bold="true" box="[2317,2441,2052,2077]" pageId="19" pageNumber="246">German:</emphasis>
<vernacularName box="[2450,2701,2050,2077]" pageId="19" pageNumber="246">Cox-Rundblattnase</vernacularName>
/
<emphasis bold="true" box="[2728,2852,2052,2077]" pageId="19" pageNumber="246">Spanish:</emphasis>
<vernacularName box="[2864,3136,2050,2077]" pageId="19" pageNumber="246">Hiposidérido de Cox</vernacularName>
</paragraph>
<paragraph blockId="19.[1923,3137,1973,2130]" box="[1924,2540,2103,2130]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[1924,2258,2105,2130]" pageId="19" pageNumber="246">Other common names:</emphasis>
<vernacularName box="[2269,2540,2103,2130]" pageId="19" pageNumber="246">Cox's Roundleaf Bat</vernacularName>
</paragraph>
</subSubSection>
</heading>
<subSubSection pageId="19" pageNumber="246" type="reference_group">
<paragraph blockId="19.[2738,3546,2202,2771]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[2747,2958,2202,2241]" pageId="19" pageNumber="246">Taxonomy.</emphasis>
<taxonomicName authority="Shelford, 1901" authorityName="Shelford" authorityYear="1901" class="Mammalia" family="Hipposideridae" genus="Hipposideros" isUncertain="true" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="246" phylum="Chordata" rank="species" species="coxi">Hipposiderus [sic] coxi Shelford, 1901</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="246" type="materials_examined">
<paragraph blockId="19.[2738,3546,2202,2771]" pageId="19" pageNumber="246">
<materialsCitation ID-GBIF-Occurrence="3753598364" pageId="19" pageNumber="246">
“Mount Penrisen,
<collectingCountry box="[3381,3538,2255,2294]" name="Malaysia" pageId="19" pageNumber="246">Sarawak</collectingCountry>
, Borneo [
<collectingCountry box="[2917,3092,2308,2347]" name="Malaysia" pageId="19" pageNumber="246">Malaysia</collectingCountry>
],
<quantity box="[3122,3296,2308,2347]" metricMagnitude="3" metricUnit="m" metricValue="1.28016" pageId="19" pageNumber="246" unit="ft" value="4200.0">4200 feet</quantity>
[=
<quantity box="[3362,3509,2308,2347]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="19" pageNumber="246" unit="m" value="1280.0">1280 m</quantity>
].”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="246" type="discussion">
<paragraph blockId="19.[2738,3546,2202,2771]" pageId="19" pageNumber="246">
<taxonomicName authorityName="Shelford" authorityYear="1901" box="[2748,3065,2362,2401]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="246" phylum="Chordata" rank="species" species="coxi">Hipposideros coxi</taxonomicName>
was formerly included in the
<taxonomicName authorityName="Temminck" authorityYear="1834" box="[2894,3008,2414,2453]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="246" phylum="Chordata" rank="species" species="bicolor">bicolor</taxonomicName>
species group, but is now placed in the
<taxonomicName authorityName="Dobson" authorityYear="1877" box="[3057,3235,2467,2506]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="246" phylum="Chordata" rank="species" species="calcaratus">calcaratus</taxonomicName>
species group. Monotypic.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="246" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.3749141" ID-Zenodo-Dep="3749141" httpUri="https://zenodo.org/record/3749141/files/figure.png" inLine="true" pageId="19" pageNumber="246" startId="19.[1923,1995,2785,2824]" targetBox="[1921,2716,2207,2764]" targetPageId="19">
<paragraph blockId="19.[2738,3546,2202,2771]" lastBlockId="19.[1920,3544,2785,4193]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[2746,2984,2574,2613]" pageId="19" pageNumber="246">Distribution.</emphasis>
Known only from a few localities in W Borneo (N &amp; SW
<collectingCountry box="[3360,3522,2626,2665]" name="Malaysia" pageId="19" pageNumber="246">Sarawak</collectingCountry>
), including Gunung Mulu, Bako, and Tanjung Datu national parks, Mt Penrissen, and Bau Limestone area; it has been reported in Central Kalimantan based on acoustic surveys, but this requires confirmation.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="19" pageNumber="246" type="description">
<paragraph blockId="19.[1920,3544,2785,4193]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[1923,2263,2891,2930]" pageId="19" pageNumber="246">Descriptive notes.</emphasis>
Forearm
<quantity box="[2452,2656,2891,2930]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="19" pageNumber="246" unit="mm" value="54.0" valueMax="55.0" valueMin="53.0">53—55 mm</quantity>
. Ears of Coxs Leaf-nosed Bat are covered with hairs. Noseleaf is enlarged with two lateral leaflets. Anterior leaf fully covers muzzle. Intermediate leaf is bulbous and thickened. Posterior is large with swollen thick bump in middle. Wing membrane is dark. Pelage color is dark brown with ventral side paler.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="246" type="biology_ecology">
<paragraph blockId="19.[1920,3544,2785,4193]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[1922,2075,3156,3195]" pageId="19" pageNumber="246">Habitat.</emphasis>
Coxs Leaf-nosed Bat forages in forest gaps understory, e.g. trails. It has been netted in mangrove forest near rock crevices. It occurs in lowlands, from sea level up to a hill cave at
<quantity box="[2219,2358,3262,3301]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="246" unit="m" value="1200.0">1200 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="246" type="food_feeding">
<paragraph blockId="19.[1920,3544,2785,4193]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[1922,2279,3314,3353]" pageId="19" pageNumber="246">Food and Feeding.</emphasis>
Coxs Leaf-nosed Bat feeds on small beedes, moths, and other insects over shrubs, creeks and water bodies by using aerial and gleaning tactics. It can fly slowly and hover during gleaning.
</paragraph>
</subSubSection>
<subSubSection box="[1922,2426,3473,3512]" pageId="19" pageNumber="246" type="breeding">
<paragraph blockId="19.[1920,3544,2785,4193]" box="[1922,2426,3473,3512]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[1922,2106,3473,3512]" pageId="19" pageNumber="246">Breeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="246" type="activity">
<paragraph blockId="19.[1920,3544,2785,4193]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[1920,2237,3525,3564]" pageId="19" pageNumber="246">Activity patterns.</emphasis>
Coxs Leaf-nosed Bat is a cave-dwelling species. Echolocation is a typical CF/FM type, with a call frequency of the F segment of46-52 kHz.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="246" type="biology_ecology">
<paragraph blockId="19.[1920,3544,2785,4193]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[1922,2916,3631,3670]" pageId="19" pageNumber="246">Movements, Home range and Social organization.</emphasis>
Coxs Leaf-nosed Bat roosts in caves, typically in small groups of 2-5 individuals. At Wind Cave,
<collectingCountry box="[3243,3405,3684,3723]" name="Malaysia" pageId="19" pageNumber="246">Sarawak</collectingCountry>
, it was recorded roosting on the cave ceiling in a group of less than 20 individuals near an opening.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="246" type="conservation">
<paragraph blockId="19.[1920,3544,2785,4193]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[1922,2397,3842,3881]" pageId="19" pageNumber="246">Status and Conservation.</emphasis>
Classified as Endangered on The IUCNRed List. Coxs Leafnosed Bat is endemic to Borneo and is rare; its population is very small and decreasing (area of occupancy is less than
<quantity box="[2502,2640,3947,3986]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="246" unit="km" value="500.0">500 km</quantity>
2). Besides habitat loss, the main threats include mining and bird-nest harvesting. Recorded in Bako, Tanjung Datu, and Gunung Mulu national parks and Wind and Fairy cave reserves.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="246" type="bibRefCitation_list">
<paragraph blockId="19.[1920,3544,2785,4193]" pageId="19" pageNumber="246">
<emphasis bold="true" box="[1923,2129,4111,4140]" pageId="19" pageNumber="246">Bibliography.</emphasis>
Corbet &amp; Hill (1992), Khan eta/. (2008), MacArthur (2016a), Phillipps &amp; Phillipps (2016), Rosli et al. (2018), Simmons (2005).
</paragraph>
</subSubSection>
</treatment>
</document>