224 lines
26 KiB
XML
224 lines
26 KiB
XML
<document id="9B1A95FE427A17F416EEE7DE52786ED7" ID-DOI="10.5281/zenodo.191909" ID-GBIF-Dataset="b5235354-ef06-4a49-b9d1-5498e97c3639" ID-ISSN="1175-5326" ID-Zenodo-Dep="191909" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461137864308" checkinUser="plazi" docAuthor="Ng, Peter K. L., Rahayu, Dwi Listyo & Naser, Murtada D." docDate="2009" docId="03F58791D3290C18C9CF9B6FFBBCFB39" docLanguage="en" docName="zt02312p026.pdf" docOrigin="Zootaxa 2312" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Manningis arabicum Jones & Clayton 1983" docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="FFCCFFE9D3200C12C9589874FFC1FFD4" masterDocTitle="The Camptandriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-Khayat & Jones, 1996 (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="26" masterPageNumber="1" pageNumber="10" updateTime="1698598725934" updateUser="plazi">
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<mods:titleInfo id="F5F7C83DC57C8369F07700BE00EBB6D3">
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<mods:title id="FA2B83889FD044BD9D9913CCA576D7BB">The Camptandriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-Khayat & Jones, 1996 (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:name id="47976F7FFA3510D82EAB31AF708B9A0E" type="personal">
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<mods:namePart id="315E5AE921973D40B937E35E51D53C6C">Ng, Peter K. L.</mods:namePart>
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<mods:name id="D5ABB4D1B505B9F9C2CF91BBA9E084C4" type="personal">
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<mods:namePart id="C94530099AEF1842C4E0E59458C2D3EF">Rahayu, Dwi Listyo</mods:namePart>
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<mods:roleTerm id="738A2EA971EF6ECCDD25145BF100BEFE">Author</mods:roleTerm>
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<mods:namePart id="4214A322EF308866E0E692E266536F2F">Naser, Murtada D.</mods:namePart>
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<mods:typeOfResource id="2BBBF942EDBA8F3486E668B42A119D4A">text</mods:typeOfResource>
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<mods:title id="50553FEE711C34D6824B77E2424DFE4E">Zootaxa</mods:title>
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<mods:part id="3B5C2D49283B0DD23C47CB373E873A44">
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<mods:date id="1298F9636B8DF38B813C0D1767A13701">2009</mods:date>
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<mods:detail id="E9A6109CDE529DCC9D20FEC51161BFFE" type="volume">
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<mods:number id="9B57CCC4205B82366DC914902B47B6B5">2312</mods:number>
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<mods:classification id="F897D2BB2BF1BC92FFDF8490C6BBAA2C">journal article</mods:classification>
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<mods:identifier id="2A278979C347F888F43C7A1F45982114" type="DOI">10.5281/zenodo.191909</mods:identifier>
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<mods:identifier id="7633BE6C422F223FF3E1FCBDDC3DCCAA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A425DECE39BA225741DB706F26264EE6" type="Zenodo-Dep">191909</mods:identifier>
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<treatment id="03F58791D3290C18C9CF9B6FFBBCFB39" ID-DOI="http://doi.org/10.5281/zenodo.6226791" ID-GBIF-Taxon="119627623" ID-Zenodo-Dep="6226791" LSID="urn:lsid:plazi:treatment:03F58791D3290C18C9CF9B6FFBBCFB39" httpUri="http://treatment.plazi.org/id/03F58791D3290C18C9CF9B6FFBBCFB39" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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<subSubSection id="C346650CD3290C1BC9CF9B6FFEEAFC8F" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="8BE33687D3290C1BC9CF9B6FFD0BFCE1" blockId="9.[151,714,795,859]" box="[151,714,795,821]" pageId="9" pageNumber="10">
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<heading id="D0AB81EBD3290C1BC9CF9B6FFD0BFCE1" bold="true" box="[151,714,795,821]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
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<taxonomicName id="4C5C4D04D3290C1BC9CF9B6FFD0BFCE1" ID-CoL="3XVPC" authority="Jones & Clayton, 1983" authorityName="Jones & Clayton" authorityYear="1983" box="[151,714,795,821]" class="Malacostraca" family="Camptandriidae" genus="Manningis" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="arabicum">
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<emphasis id="B928EA95D3290C1BC9CF9B6FFD0BFCE1" bold="true" box="[151,714,795,821]" pageId="9" pageNumber="10">
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||
<emphasis id="B928EA95D3290C1BC9CF9B6FFE57FCE1" bold="true" box="[151,406,795,821]" italics="true" pageId="9" pageNumber="10">Manningis arabicum</emphasis>
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(
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<bibRefCitation id="EFCD4B76D3290C1BC8FD9B6FFD03FCE1" author="Jones" box="[421,706,795,821]" pageId="9" pageNumber="10" refString="Jones, D. A. & Clayton, D. (1983) The systematics and ecology of crabs belonging to the genera Cleistostoma de Haan and Paracleistostoma de Man on Kuwait mudflats. Crustaceana, 45 (2), 183 - 199." type="journal article" year="1983">Jones & Clayton, 1983</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BE33687D3290C1BC9CF9B35FEEAFC8F" blockId="9.[151,714,795,859]" box="[151,299,833,859]" pageId="9" pageNumber="10">
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(
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<figureCitation id="13672A02D3290C1BC9C79B35FF2CFC8F" box="[159,237,833,859]" captionStart="FIGURE 1" captionStartId="2.[151,259,1488,1512]" captionTargetBox="[151,1434,335,1470]" captionTargetId="figure@2.[151,1436,335,1470]" captionTargetPageId="2" captionText="FIGURE 1. Overall views. A, Opusia indicum (Alcock, 1900), male (6.6 x 11.3 mm) (ZRC 2009.832), Iraq; B, Manningis arabicum (Jones & Clayton, 1983), male (3.5 x 5.3 mm) (ZRC 2009.1097), Iraq." httpUri="https://zenodo.org/record/191910/files/figure.png" pageId="9" pageNumber="10">Figs. 1</figureCitation>
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B, 4)
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</paragraph>
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</subSubSection>
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<subSubSection id="C346650CD3290C1BC9CF9BF9FB91FC1E" pageId="9" pageNumber="10" type="reference_group">
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<paragraph id="8BE33687D3290C1BC9CF9BF9FCFDFC70" blockId="9.[151,1104,909,970]" box="[151,828,909,932]" pageId="9" pageNumber="10">
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<treatmentCitationGroup id="AB4C11A9D3290C1BC9CF9BF9FCFDFC70" box="[151,828,909,932]" pageId="9" pageNumber="10">
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<taxonomicName id="4C5C4D04D3290C1BC9CF9BF9FD30FC70" ID-CoL="4CQ8Z" authority="Jones & Clayton, 1983: 190" authorityName="Jones & Clayton" authorityPageNumber="190" authorityYear="1983" box="[151,753,909,932]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="arabicum">
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<emphasis id="B928EA95D3290C1BC9CF9BF9FE7FFC70" box="[151,446,909,932]" italics="true" pageId="9" pageNumber="10">Paracleistostoma arabicum</emphasis>
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<treatmentCitation id="0AFD1096D3290C1BC89D9BF9FD30FC70" author="Jones" box="[453,753,909,932]" page="190" pageId="9" pageNumber="10" year="1983">
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<bibRefCitation id="EFCD4B76D3290C1BC89D9BF9FD7CFC70" author="Jones" box="[453,701,909,932]" pageId="9" pageNumber="10" refString="Jones, D. A. & Clayton, D. (1983) The systematics and ecology of crabs belonging to the genera Cleistostoma de Haan and Paracleistostoma de Man on Kuwait mudflats. Crustaceana, 45 (2), 183 - 199." type="journal article" year="1983">Jones & Clayton, 1983</bibRefCitation>
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: 190
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</treatmentCitation>
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</taxonomicName>
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, fig. 4.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE33687D3290C1BC9CF9BC7FB91FC1E" blockId="9.[151,1104,909,970]" box="[151,1104,947,970]" pageId="9" pageNumber="10">
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<treatmentCitationGroup id="AB4C11A9D3290C1BC9CF9BC7FB91FC1E" box="[151,1104,947,970]" pageId="9" pageNumber="10">
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<taxonomicName id="4C5C4D04D3290C1BC9CF9BC7FEB4FC1E" ID-CoL="3XVPC" box="[151,373,947,970]" class="Malacostraca" family="Camptandriidae" genus="Manningis" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="arabicum">
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<emphasis id="B928EA95D3290C1BC9CF9BC7FEB4FC1E" box="[151,373,947,970]" italics="true" pageId="9" pageNumber="10">Manningis arabicum</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0AFD1096D3290C1BC8C59BC7FD21FC1E" author="Al-Khayat" box="[413,736,947,970]" page="798" pageId="9" pageNumber="10" year="1996">
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<bibRefCitation id="EFCD4B76D3290C1BC8C59BC7FD6AFC1E" author="Al-Khayat" box="[413,683,947,970]" pageId="9" pageNumber="10" refString="Al-Khayat, J. A. & Jones, D. A. (1996) Two new genera, Manningis and Leptochryseus (Decapoda: Camptandriinae), and descriptions of the first zoea of six brachyurans from the Arabian Gulf. Journal of Crustacean Biology, 16 (4), 797 - 813." type="journal article" year="1996">Al-Khayat & Jones 1996</bibRefCitation>
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: 798
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</treatmentCitation>
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, 809, fig. 7. – Ng
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<emphasis id="B928EA95D3290C1BCAF99BC7FC16FC1E" box="[929,983,947,970]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
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2008: 233.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C346650CD3290C1BC9CF9B8AFD86FB02" pageId="9" pageNumber="10" type="materials_examined">
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<paragraph id="8BE33687D3290C1BC9CF9B8AFDD2FB64" blockId="9.[151,1437,1022,2033]" pageId="9" pageNumber="10">
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<emphasis id="B928EA95D3290C1BC9CF9B8AFE31FBCC" bold="true" box="[151,496,1022,1048]" pageId="9" pageNumber="10">
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Material examined.
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<collectingCountry id="F34B7617D3290C1BC8F99B8AFE31FBCC" box="[417,496,1022,1048]" name="Iraq" pageId="9" pageNumber="10">IRAQ</collectingCountry>
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</emphasis>
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:
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<specimenCount id="9D5AFD0ED3290C1BCB599B8BFD94FBCD" box="[513,597,1023,1049]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
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(3.5 x
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<quantity id="4CA49B62D3290C1BCBF79B8BFCD1FBCD" box="[687,784,1023,1049]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="9" pageNumber="10" unit="mm" value="5.3">5.3 mm</quantity>
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) (
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<collectionCode id="ED4DAE42D3290C1BCA719B8BFCA7FBCD" box="[809,870,1023,1049]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="10">ZRC</collectionCode>
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2009.1097), Shatt Al-Basrah,
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<geoCoordinate id="EE685040D3290C1BCDBD9B8BFA5DFBCD" box="[1253,1436,1023,1049]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="1" value="30.405947">30°24'21.41"N</geoCoordinate>
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<geoCoordinate id="EE685040D3290C1BC9CF9C50FE86FBEA" box="[151,327,1060,1086]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="1" value="47.778675">47°46'43.23"E</geoCoordinate>
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, coll. M. D. Naser, 2009.
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<emphasis id="B928EA95D3290C1BCB269C50FD39FBEA" bold="true" box="[638,760,1060,1086]" pageId="9" pageNumber="10">
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<collectingCountry id="F34B7617D3290C1BCB269C50FD39FBEA" box="[638,760,1060,1086]" name="Kuwait" pageId="9" pageNumber="10">KUWAIT</collectingCountry>
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</emphasis>
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:
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<typeStatus id="54E78825D3290C1BCA509C50FCB5FBEA" box="[776,884,1060,1086]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
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: male (5.4 x
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<quantity id="4CA49B62D3290C1BCD489C50FBAFFBEA" box="[1040,1134,1060,1086]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="9" pageNumber="10" unit="mm" value="8.9">8.9 mm</quantity>
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) (NHM 1981.496), sand/ mudflats, Al Kiran,
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<date id="FFE21047D3290C1BC8DB9C3EFDDCFBB0" box="[387,541,1098,1124]" pageId="9" pageNumber="10" value="1981-04-08">8 April 1981</date>
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;
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<typeStatus id="54E78825D3290C1BCB719C3EFD5FFBB0" box="[553,670,1098,1124]" pageId="9" pageNumber="10" type="paratype">paratypes</typeStatus>
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:
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<specimenCount id="9D5AFD0ED3290C1BCBF19C3EFCC2FBB0" box="[681,771,1098,1124]" pageId="9" pageNumber="10" type="male">2 males</specimenCount>
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(4.4 x 7.0 mm, 4.4 x 7.0 mm),
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<specimenCount id="9D5AFD0ED3290C1BCD2D9C3EFB26FBB0" box="[1141,1255,1098,1124]" pageId="9" pageNumber="10" type="female">3 females</specimenCount>
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(4.2 x
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<quantity id="4CA49B62D3290C1BCC629C3EFA59FBB0" box="[1338,1432,1098,1124]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="9" pageNumber="10" unit="mm" value="6.9">6.9 mm</quantity>
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, 4.4 x
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<quantity id="4CA49B62D3290C1BC9859C04FEFCFB5E" box="[221,317,1136,1162]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="9" pageNumber="10" unit="mm" value="7.2">7.2 mm</quantity>
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, 4.6 x
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<quantity id="4CA49B62D3290C1BC8C89C04FE2EFB5E" box="[400,495,1136,1162]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="9" pageNumber="10" unit="mm" value="8.1">8.1 mm</quantity>
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) (NHM 1981.497), sand/mudflats, Al Kiran,
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<date id="FFE21047D3290C1BCD4F9C04FB76FB5E" box="[1047,1207,1136,1162]" pageId="9" pageNumber="10" value="1981-04-08">8 April 1981</date>
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||
;
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||
<specimenCount id="9D5AFD0ED3290C1BCD9C9C04FAE4FB5E" box="[1220,1317,1136,1162]" pageId="9" pageNumber="10" type="male">2 males</specimenCount>
|
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,
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<specimenCount id="9D5AFD0ED3290C1BCC6A9C04FA5DFB5E" box="[1330,1436,1136,1162]" pageId="9" pageNumber="10" type="female">1 female</specimenCount>
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(NHM 1974.389), no other data.
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</paragraph>
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<paragraph id="8BE33687D3290C1BC99E9CC8FD86FB02" blockId="9.[151,1437,1022,2033]" box="[198,583,1212,1238]" pageId="9" pageNumber="10">
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<emphasis id="B928EA95D3290C1BC99E9CC8FEADFB02" bold="true" box="[198,364,1212,1238]" pageId="9" pageNumber="10">
|
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<typeStatus id="54E78825D3290C1BC99E9CC8FEC5FB02" box="[198,260,1212,1238]" pageId="9" pageNumber="10">Type</typeStatus>
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locality.
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</emphasis>
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Al Kiran,
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<collectingCountry id="F34B7617D3290C1BC8B19CC8FD82FB02" box="[489,579,1212,1238]" name="Kuwait" pageId="9" pageNumber="10">Kuwait</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C346650CD3290C18C99E9C95FD64FD56" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
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<paragraph id="8BE33687D3290C1BC99E9C95FCB7F916" blockId="9.[151,1437,1022,2033]" pageId="9" pageNumber="10">
|
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<emphasis id="B928EA95D3290C1BC99E9C95FE9EFB2F" bold="true" box="[198,351,1249,1275]" pageId="9" pageNumber="10">Description.</emphasis>
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Carapace much broader (1.6–1.7 times) than long, glabrous, dorsal surface sloping downwards from posterior to anterior portion, hepatic region immediately posterior to external orbital tooth sunken, epigastric lobes indistinct, posterior branchial region with central group of rounded granules; gastric region moderately convex, anterior, lateral grooves of cardiac region, lateral grooves of intestinal region distinct; anterolateral margin convex, granular, with 2 weak lobes posterior to external orbital tooth, dorsolateral margin defined by row of granules running along anterior half of posterior branchial region; posterolateral margin convex, rounded; posterior carapace margin thickened, broad, straight. Front about as wide as orbital, deflexed, margin thick, weakly bilobed from dorsal view, lateral angles sharp, junction between base of front, supraorbital margin rounded, without distinct notch; frontomedial margin distinctly bilobed. Supraorbital margin sloped posteriorly, sinuous, thick, outer corner with tuft of setae curled over eye; infraorbital margin finely granular, setose, median third with larger granules; inner infraorbital tooth not very distinct from rest of margin, separated by shallow, wide indentation, suborbital ridge moderately developed, sparsely setose, granular along outer third, not visible dorsally.
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</paragraph>
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<paragraph id="8BE33687D3290C1BC99E9EBAFDC8F854" blockId="9.[151,1437,1022,2033]" pageId="9" pageNumber="10">Pterygostomian region smooth, sparsely setose, with small granules just under outer end of supraorbital margin, anterior region with shallow, transverse, closed sulcus. Anterior half of epistome separated from posterior half by distinct groove, followed posteriorly by transverse ridge. Posterolateral margin separated from anterior pterygostomian margin by open sulcus, posteromedial tooth broadly triangular, posterior margins on either side concave.</paragraph>
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<paragraph id="8BE33687D3290C1BC99E9FFFFC43F825" blockId="9.[151,1437,1022,2033]" pageId="9" pageNumber="10">Third maxillipeds with thick inner margin; merus slightly shorter than ischium, outer distal angle slightly broad, rounded, outer surface without distinct grooves; ischium with inner distal angle slightly produced; basal segment of palp not excavated to form spatulate structure.</paragraph>
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<paragraph id="8BE33687D32A0C18C99E98ECFBABFE98" blockId="10.[151,1437,152,1261]" pageId="10" pageNumber="11">Chelipeds robust in males, well developed; outer surface of merus granular, particularly on dorsal surface; carpus globular from dorsal view, with small tuft of setae on inner surface; palm about as high as long, smooth, with row of setae on inner surface, close to dorsal margin; fingers with setose inner surfaces, tips spatulate, movable finger with large, molariform tooth closer to proximal end, immovable finger with cutting edge curved slightly, forming gape. Female chelipeds slender, setose, tips spatulate.</paragraph>
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<paragraph id="8BE33687D32A0C18C99E992DFCFCFE4E" blockId="10.[151,1437,152,1261]" pageId="10" pageNumber="11">Ambulatory legs relatively stout, short, sparsely setose; P3 longest, P5 shortest, second, third meri with anterior, dorsal surface granular, never spinular or serrate.</paragraph>
|
||
<paragraph id="8BE33687D32A0C18C99E99D2FC9DFDE0" blockId="10.[151,1437,152,1261]" pageId="10" pageNumber="11">Thoracic sternum swollen, such that portion of sternum, abdomen visible dorsally. Male abdominal somite 1 about as wide as somite 2, somites 2–4 immobile, sutures complete, not as deep as those between somites 1, 2, 5–7, somites 5, 6, telson freely mobile, lateral margins of somite 6 convex, angular. Female abdomen subcircular, with all somites free, telson triangular.</paragraph>
|
||
<paragraph id="8BE33687D32A0C18C99E9A35FD64FD56" blockId="10.[151,1437,152,1261]" pageId="10" pageNumber="11">G1 ending in truncate apex, apex covered with strong, short, backward curving spines, row of long, strong comb-shaped spines subapically on one side.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346650CD32A0C18C99E9AFAFBBCFB39" pageId="10" pageNumber="11" type="discussion">
|
||
<paragraph id="8BE33687D32A0C18C99E9AFAFADDFD22" blockId="10.[151,1437,152,1261]" pageId="10" pageNumber="11">
|
||
<emphasis id="B928EA95D32A0C18C99E9AFAFE81FD7C" bold="true" box="[198,320,654,680]" pageId="10" pageNumber="11">Remarks.</emphasis>
|
||
The male
|
||
<typeStatus id="54E78825D32A0C18C8E69AFAFDE5FD7C" box="[446,548,654,680]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
agrees well with the original descriptions and figures provided by
|
||
<bibRefCitation id="EFCD4B76D32A0C18CC649AFAFE6DFD1B" author="Jones" pageId="10" pageNumber="11" refString="Jones, D. A. & Clayton, D. (1983) The systematics and ecology of crabs belonging to the genera Cleistostoma de Haan and Paracleistostoma de Man on Kuwait mudflats. Crustaceana, 45 (2), 183 - 199." type="journal article" year="1983" yearSuffix="a">Jones & Clayton (1983: fig. 4a)</bibRefCitation>
|
||
except that in their figure, the carapace grooves appear deeper than they are on the actual specimen (
|
||
<figureCitation id="13672A02D32A0C18C83A9AA8FE68FD22" box="[354,425,732,758]" captionStart="FIGURE 4" captionStartId="8.[151,255,1926,1950]" captionTargetBox="[172,1411,189,1872]" captionTargetId="figure@8.[157,1430,177,1901]" captionTargetPageId="8" captionText="FIGURE 4. Manningis arabicum (Jones & Clayton, 1983), holotype male (5.4 x 8.9 mm) (NHM 1981.496), Kuwait. A, carapace; B, frontal view of carapace showing orbit, eye and epistome; D, inner view of right chela; E, dorsal view of right cheliped; F, left G 1; G, distal part of left G 1. Scales: A, B, D – F = 1.0 mm; C = 0.5 mm; G = 0.15 mm." httpUri="https://zenodo.org/record/191913/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
|
||
A). Specimens preserved in alcohol were creamy white with deep-red eyes.
|
||
</paragraph>
|
||
<paragraph id="8BE33687D32A0C18C99E9B76FDC9FB12" blockId="10.[151,1437,152,1261]" pageId="10" pageNumber="11">
|
||
<emphasis id="B928EA95D32A0C18C99E9B76FE0FFCC8" bold="true" box="[198,462,770,796]" pageId="10" pageNumber="11">Habitat and Biology.</emphasis>
|
||
<bibRefCitation id="EFCD4B76D32A0C18C8819B76FCEEFCC8" author="Jones" box="[473,815,770,796]" pageId="10" pageNumber="11" refString="Jones, D. A. & Clayton, D. (1983) The systematics and ecology of crabs belonging to the genera Cleistostoma de Haan and Paracleistostoma de Man on Kuwait mudflats. Crustaceana, 45 (2), 183 - 199." type="journal article" year="1983">Jones & Clayton (1983:192)</bibRefCitation>
|
||
gave an account of the biology of this species and remarked that "growth of the large chelae seen in adult males again appears to be rapid, and to some extent unrelated to overall crab size". They added that
|
||
<taxonomicName id="4C5C4D04D32A0C18CBBF9B24FC24FCBD" box="[743,997,848,873]" class="Malacostraca" family="Camptandriidae" genus="Manningis" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="arabicum">
|
||
<emphasis id="B928EA95D32A0C18CBBF9B24FC24FCBD" box="[743,997,848,873]" italics="true" pageId="10" pageNumber="11">Manningis arabicum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5C4D04D32A0C18CD7A9B24FADFFCBD" box="[1058,1310,848,873]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32A0C18CD7A9B24FADFFCBD" box="[1058,1310,848,873]" italics="true" pageId="10" pageNumber="11">Nasima dotilliformis</emphasis>
|
||
</taxonomicName>
|
||
are found slightly lower intertidally on the mudflats. According to these authors,
|
||
<taxonomicName id="4C5C4D04D32A0C18CA829B03FBB0FC44" box="[986,1137,887,912]" class="Malacostraca" family="Camptandriidae" genus="Manningis" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="arabicum">
|
||
<emphasis id="B928EA95D32A0C18CA829B03FBB0FC44" box="[986,1137,887,912]" italics="true" pageId="10" pageNumber="11">M. arabicum</emphasis>
|
||
</taxonomicName>
|
||
is a deposit feeder, using its forcep-like chela to pick up sand grains. Together with
|
||
<taxonomicName id="4C5C4D04D32A0C18CA0C9BE9FBD3FC62" box="[852,1042,925,950]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32A0C18CA0C9BE9FBD3FC62" box="[852,1042,925,950]" italics="true" pageId="10" pageNumber="11">N. dotilliformis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C4D04D32A0C18CD7B9BE9FB7FFC62" box="[1059,1214,925,950]" class="Malacostraca" family="Camptandriidae" genus="Manningis" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="arabicum">
|
||
<emphasis id="B928EA95D32A0C18CD7B9BE9FB7FFC62" box="[1059,1214,925,950]" italics="true" pageId="10" pageNumber="11">M. arabicum</emphasis>
|
||
</taxonomicName>
|
||
was found to feed "on the surface of the sand and mudflats as soon as the ebbing tide left their [its] burrows". They also remarked that
|
||
<taxonomicName id="4C5C4D04D32A0C18C8169B9EFE2EFBD7" box="[334,495,1002,1027]" class="Malacostraca" family="Camptandriidae" genus="Manningis" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="arabicum">
|
||
<emphasis id="B928EA95D32A0C18C8169B9EFE2EFBD7" box="[334,495,1002,1027]" italics="true" pageId="10" pageNumber="11">M. arabicum</emphasis>
|
||
</taxonomicName>
|
||
was not found with
|
||
<taxonomicName id="4C5C4D04D32A0C18CBAE9B9EFB81FBD7" box="[758,1088,1002,1027]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kuwaitense">
|
||
<emphasis id="B928EA95D32A0C18CBAE9B9EFB81FBD7" box="[758,1088,1002,1027]" italics="true" pageId="10" pageNumber="11">Leptochryseus kuwaitense</emphasis>
|
||
</taxonomicName>
|
||
and that it colonised sandy mudflats at Al Kiran and areas of the salt marsh at Doha,
|
||
<collectingCountry id="F34B7617D32A0C18CA1A9C65FC5DFBFF" box="[834,924,1041,1067]" name="Kuwait" pageId="10" pageNumber="11">Kuwait</collectingCountry>
|
||
. In
|
||
<collectingCountry id="F34B7617D32A0C18CA919C65FC3AFBFF" box="[969,1019,1041,1067]" name="Iraq" pageId="10" pageNumber="11">Iraq</collectingCountry>
|
||
, the specimens were obtained from soft silty clay on intertidal salt marshes. As has been discussed earlier, because of the many development projects upstream of the major rivers, there is not much freshwater now flowing to the sea and coupled with the strong tidal influence, the area’s salinity averages 32 ‰ despite its distance from the sea (see also habitat notes for
|
||
<taxonomicName id="4C5C4D04D32A0C18C85D9CD8FE36FB11" box="[261,503,1196,1221]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dotilliformis">
|
||
<emphasis id="B928EA95D32A0C18C85D9CD8FE36FB11" box="[261,503,1196,1221]" italics="true" pageId="10" pageNumber="11">Nasima dotilliformis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE33687D32A0C18C99E9CA6FBBCFB39" blockId="10.[151,1437,152,1261]" box="[198,1149,1234,1261]" pageId="10" pageNumber="11">
|
||
<emphasis id="B928EA95D32A0C18C99E9CA6FDD3FB38" bold="true" box="[198,530,1234,1260]" pageId="10" pageNumber="11">Geographical distribution.</emphasis>
|
||
Only previously known from
|
||
<collectingCountry id="F34B7617D32A0C18CA2E9CA7FC0EFB39" box="[886,975,1235,1261]" name="Kuwait" pageId="10" pageNumber="11">Kuwait</collectingCountry>
|
||
, and now
|
||
<collectingCountry id="F34B7617D32A0C18CD1E9CA7FBB8FB39" box="[1094,1145,1235,1261]" name="Iraq" pageId="10" pageNumber="11">Iraq</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |