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<document id="7584A1B783146FB910C7010580F57BC9" ID-DOI="10.1111/zoj.12299" ID-ISSN="0024-4082" ID-Zenodo-Dep="5339971" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630185318692" checkinUser="felipe" docAuthor="Cruz-López, Jesús A. &amp; Francke, Oscar F." docDate="2015" docId="03CA87F9A27FFF87FEA2FC8EC1BFFB9E" docLanguage="en" docName="zoj12299.pdf" docOrigin="Zoological Journal of the Linnean Society 175 (4)" docSource="http://dx.doi.org/10.1111/zoj.12299" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Crettaros valdezi Cruz-López &amp; Francke 2015, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="870" masterDocId="FFF3FF81A255FFACFFBAFFF9C27CFFCC" masterDocTitle="Cladistic analysis and taxonomic revision of the genus Karos Goodnight &amp; Goodnight, 1944 (Opiliones, Laniatores, Stygnopsidae)" masterLastPageNumber="891" masterPageNumber="827" pageNumber="869" updateTime="1699179109858" updateUser="plazi">
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<mods:title id="52623C9B24E8A6F73746BD851A5798DC">Cladistic analysis and taxonomic revision of the genus Karos Goodnight &amp; Goodnight, 1944 (Opiliones, Laniatores, Stygnopsidae)</mods:title>
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<mods:namePart id="69FFB8D94A353378FEEB44B794F3ABF7">Cruz-López, Jesús A.</mods:namePart>
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<mods:namePart id="8E154F13FEEAA4EE87023A1907801CC7">Francke, Oscar F.</mods:namePart>
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<mods:title id="87C8085043619F656922FB21E400228F">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="9DFF73C9F70B8EE184986C6B77F04B60">2015</mods:date>
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<mods:number id="2F1C7EA58F433F9AD89CC45083616DDD">2015-11-18</mods:number>
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<heading id="D0948183A27FFF86FEA2FC8EC012FC41" box="[280,622,887,909]" centered="true" fontSize="9" level="2" pageId="42" pageNumber="869" reason="4">
<taxonomicName id="4C634D6CA27FFF86FEA2FC8EC075FC40" ID-CoL="7VH5J" authority="Cruz-López &amp; Francke, 2015" authorityName="Cruz-López &amp; Francke" authorityYear="2015" box="[280,521,887,908]" class="Arachnida" family="Stygnopsidae" genus="Crettaros" kingdom="Animalia" order="Opiliones" pageId="42" pageNumber="888" phylum="Arthropoda" rank="species" species="valdezi" status="sp. nov.">CRETTAROS VALDEZI</taxonomicName>
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<paragraph id="8BDC36EFA27FFF86FE9EFC6EC01EFC61" blockId="42.[280,622,887,941]" box="[292,610,919,941]" pageId="42" pageNumber="869">
(
<figureCitation id="13582A6AA27FFF86FE97FC6EC3F4FC60" box="[301,392,919,940]" captionStart="Figure 43" captionStartId="42.[144,221,1824,1843]" captionTargetBox="[305,1262,1060,1791]" captionTargetId="figure-377@42.[303,1264,1057,1794]" captionTargetPageId="42" captionText="Figure 43. Crettaros valdezi sp. nov. A, habitus, dorsal view. B, legs IV, mesal view. C, habitus, lateral view. D, habitus, ventral view. E, scutum, dorsal view. F, ocularium, frontal view. Scale bars: A = 2.5 mm, B, D, and E = 1.5 mm, C = 1 mm, F = 0.5 mm. The dark lines on (D) indicate the stigmatic area." figureDoi="http://doi.org/10.5281/zenodo.5340063" httpUri="https://zenodo.org/record/5340063/files/figure.png" pageId="42" pageNumber="869">FIGS 43</figureCitation>
,
<figureCitation id="13582A6AA27FFF86FE2FFC6EC3C8FC60" box="[405,436,919,940]" captionStart="Figure 44" captionStartId="43.[164,243,847,866]" captionTargetBox="[164,1445,195,817]" captionTargetId="figure-431@43.[164,1445,195,817]" captionTargetPageId="43" captionText="Figure 44. Crettaros valdezi sp. nov. male genitalia. A, dorsal view. B, ventral view. C, lateral view. White arrows indicate ventral microsetae in (B) and parastylar setae in (C)." figureDoi="http://doi.org/10.5281/zenodo.5340065" httpUri="https://zenodo.org/record/5340065/files/figure.png" pageId="42" pageNumber="869">44</figureCitation>
,
<figureCitation id="13582A6AA27FFF86FE7AFC6EC3A3FC61" box="[448,479,919,941]" captionStart="Figure 57" captionStartId="54.[144,221,1193,1212]" captionTargetBox="[144,1425,195,1163]" captionTargetId="figure-243@54.[144,1425,195,1163]" captionTargetPageId="54" captionText="Figure 57. Distribution map of the species of the Karos genus-group. Black triangles, Monterella tuberculata; black square, Mictlana inops comb. nov. (only locality of material examined was mapped); black circles, Crettaros santibanezi sp. nov.; white circles with black outline, Crettaros valdezi sp. nov.; black crosses, Chapulobunus unispinosus; inverted black triangles, Karos parvus." figureDoi="http://doi.org/10.5281/zenodo.5340091" httpUri="https://zenodo.org/record/5340091/files/figure.png" pageId="42" pageNumber="869">57</figureCitation>
,
<figureCitation id="13582A6AA27FFF86FE56FC6EC061FC61" box="[492,541,919,941]" captionStart="Figure 63" captionStartId="60.[144,223,1276,1295]" captionTargetBox="[144,1425,196,1246]" captionTargetId="figure-0@60.[144,1425,196,1246]" captionTargetPageId="60" captionText="Figure 63. A, dorsal view of the anal bulge on Chapulobunus poblano sp. nov.; arrow indicates the bulge. B, lateral view of Huasteca silhavyi sp. nov., line indicates the very high lateral channel area. C, laterodorsal view of scutum of Crettaros valdezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. D, posterodorsal view of scutum of Crettaros santibanezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. E, dorsal view of trochanter IV of Karos barbarikos; arrow indicates the tridentate apophysis. F, anterodorsal view of scutum of Karos barbarikos, the black points mark the tubercles forming a V. G, dorsal ornamentation of Karos tersum sp. nov. showing the transversal row of tubercles on mesotergal areas; transversal row II indicated by black dots. H, detail of the ventral ornamentation on femur IV of Karos barbarikos. I, detail of the ectal ornamentation on tibia IV of Karos barbarikos; arrows indicate the spiniform tubercles. J, detail of the ventral ornamentation on leg IV in Chapulobunus poblano sp. nov. K, detail of the two apical spines on femur IV in Chapulobunus poblano sp. nov. These images are not at the same scale." figureDoi="http://doi.org/10.5281/zenodo.5340103" httpUri="https://zenodo.org/record/5340103/files/figure.png" pageId="42" pageNumber="869">63C</figureCitation>
,
<figureCitation id="13582A6AA27FFF86FD93FC6EC026FC61" box="[553,602,919,941]" captionStart="Figure 64" captionStartId="61.[164,241,1568,1587]" captionTargetBox="[266,1342,195,1560]" captionTargetId="figure-0@61.[266,1342,195,1560]" captionTargetPageId="61" captionText="Figure 64. A, mesodorsal view of pedipalp femur and patella of Huasteca silhavyi sp. nov.; arrows indicate the setiferous tubercles on mesal side of these segments. B, dorsal view of pedipalp femur of Karos barbarikos; arrow indicates the mesodistal setiferous tubercle. C, apical portion of metatarsus IV of Karos singularis sp. nov., the circle indicates the swollen area. D, dorsal view of pars distalis of Karos barbarikos; line indicates the exposed base of the follis. E, dorsal view of pars distalis of Philora tuxtlae; line indicates the base of the follis in apical depression. F, dorsal view of pars distalis of Paramitraceras granulatum; arrows indicate the pars distalis width/follis width ratio. G, dorsal view of pars distalis of Crettaros valdezi sp. nov.; arrows indicate the pars distalis width/follis width ratio. H, ventral view of pars distalis of Chapulobunus unispinosus; arrow indicates the width at the middle of the ventral plate. I, ventral view of pars distalis of Mictlana inops the white arrow indicates the slight width in the middle of ventral plate, the black dotted line indicates the apical end of ventral plate. J, ventral view of pars distalis of Karos barbarikos, the white dotted line indicates the apical end of ventral plate. K, ventral view of pars distalis of Chapulobunus poblano sp. nov., the white dotted line indicates the apical end of ventral plate. L, dorsal view of pars distalis of Potosa dybasi; arrows indi-" figureDoi="http://doi.org/10.5281/zenodo.5340105" httpUri="https://zenodo.org/record/5340105/files/figure.png" pageId="42" pageNumber="869">64G</figureCitation>
)
</paragraph>
<paragraph id="8BDC36EFA27FFF86FF2AFC46C08AFC3E" blockId="42.[144,759,957,1010]" pageId="42" pageNumber="869">
<emphasis id="B917EAFDA27FFF86FF2AFC46C34DFC1F" box="[144,305,958,980]" italics="true" pageId="42" pageNumber="869">Type material:</emphasis>
<materialsCitation id="3B0B3CB2A27FFF86FE83FC47C08FFC18" box="[313,755,957,980]" country="Mexico" location="San Luis Potosi" pageId="42" pageNumber="869" specimenCount="1" specimenCount-male="1" stateProvince="San Luis Potosi" typeStatus="holotype">
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:
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<collectingRegion id="49A7F80DA27FFF86FE0AFC47C01FFC1F" box="[432,611,958,979]" country="Mexico" name="San Luis Potosi" pageId="42" pageNumber="869">San Luís Potosí</collectingRegion>
:
</emphasis>
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<typeStatus id="54D8884DA27FFF86FD2BFC47C08FFC18" box="[657,755,958,980]" pageId="42" pageNumber="869" type="holotype">holotype</typeStatus>
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,
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<specimenCount id="9D65FD66A27FFF86FF2AFC24C2BEFC3E" box="[144,194,988,1010]" pageId="42" pageNumber="869" type="male">1 ♂</specimenCount>
and
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<typeStatus id="54D8884DA27FFF86FE84FC24C3CCFC3E" box="[318,432,989,1010]" pageId="42" pageNumber="869" type="paratype">paratypes</typeStatus>
[
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and CNAN-
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<paragraph id="8BDC36EFA27FFF86FC9CFF3FC76BFED4" blockId="42.[806,1421,198,281]" pageId="42" pageNumber="869">
T0800 (
<date id="FFDD102FA27FFF86FCC4FF3FC197FF17" box="[894,1003,198,219]" pageId="42" pageNumber="869" value="2012-05-13">13.v.2012</date>
; J. Cruz, G. Contreras, J. Mendoza, and R. Monjaraz)] (
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,
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). Municipio Villa de Zaragoza, Km 58 Cave.
</paragraph>
<paragraph id="8BDC36EFA27FFF86FC9CFEBEC7F1FDA3" blockId="42.[806,1421,326,624]" pageId="42" pageNumber="869">
<emphasis id="B917EAFDA27FFF86FC9CFEBEC67BFE90" box="[806,1031,327,348]" italics="true" pageId="42" pageNumber="869">Material examined:</emphasis>
<materialsCitation id="3B0B3CB2A27FFF86FBA8FEBEC11AFE74" collectingDate="2010-05-13" collectionCode="CNAN" collectorName="O. Francke &amp; A. Valdez &amp; C. Santibanez &amp; J. Cruz" country="Mexico" pageId="42" pageNumber="869" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="San Luis Potosi">
<collectingCountry id="F374767FA27FFF86FBA8FEBEC607FE90" box="[1042,1147,327,348]" name="Mexico" pageId="42" pageNumber="869">MEXICO</collectingCountry>
:
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<collectingRegion id="49A7F80DA27FFF86FB34FEBEC736FE90" box="[1166,1354,327,348]" country="Mexico" name="San Luis Potosi" pageId="42" pageNumber="869">San Luís Potosí</collectingRegion>
:
</emphasis>
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and
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[
<collectionCode id="ED72AE2AA27FFF86FC28FE9CC19CFEB6" box="[914,992,357,378]" pageId="42" pageNumber="869">CNAN</collectionCode>
(
<date id="FFDD102FA27FFF86FC4BFE9CC621FEB6" box="[1009,1117,357,378]" pageId="42" pageNumber="869" value="2010-05-13">
<collectingDate id="EF99E9C7A27FFF86FC4BFE9CC621FEB6" box="[1009,1117,357,378]" pageId="42" pageNumber="869" value="2010-05-13">13.v.2010</collectingDate>
</date>
;
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,
<collectorName id="26965339A27FFF86FB43FE9CC714FEB7" box="[1273,1384,357,379]" pageId="42" pageNumber="869">A. Valdez</collectorName>
,
<collectorName id="26965339A27FFF86FACEFE9CC1D5FE56" pageId="42" pageNumber="869">C. Santibáñez</collectorName>
, and
<collectorName id="26965339A27FFF86FC55FE7DC636FE55" box="[1007,1098,387,409]" pageId="42" pageNumber="869">J. Cruz</collectorName>
], same locality data as the
<typeStatus id="54D8884DA27FFF86FC9CFE5AC11AFE74" box="[806,870,419,440]" pageId="42" pageNumber="869">types</typeStatus>
</materialsCitation>
.
<materialsCitation id="3B0B3CB2A27FFF86FCCCFE5AC193FDD8" collectingDate="2012-05-13" collectionCode="CNAN, R" collectorName="J. Cruz &amp; G. Contreras &amp; J. Mendoza &amp; Monjaraz &amp; Km" country="Mexico" latitude="22.01677" location="Municipio Villa de Zaragoza" longLatPrecision="1" longitude="-100.59166" pageId="42" pageNumber="869" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="San Luis Potosi">
<specimenCount id="9D65FD66A27FFF86FCCCFE5AC1D4FE74" box="[886,936,418,440]" pageId="42" pageNumber="869" type="male">1 ♂</specimenCount>
and
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[
<collectionCode id="ED72AE2AA27FFF86FB93FE5AC60BFE74" box="[1065,1143,419,440]" pageId="42" pageNumber="869">CNAN</collectionCode>
(
<date id="FFDD102FA27FFF86FB31FE5AC685FE74" box="[1163,1273,419,440]" pageId="42" pageNumber="869" value="2012-05-13">
<collectingDate id="EF99E9C7A27FFF86FB31FE5AC685FE74" box="[1163,1273,419,440]" pageId="42" pageNumber="869" value="2012-05-13">13.v.2012</collectingDate>
</date>
;
<collectorName id="26965339A27FFF86FAB2FE5AC71FFE74" box="[1288,1379,419,440]" pageId="42" pageNumber="869">J. Cruz</collectorName>
,
<collectorName id="26965339A27FFF86FAC9FE5AC1EAFE1A" pageId="42" pageNumber="869">G. Contreras</collectorName>
,
<collectorName id="26965339A27FFF86FC18FE38C65FFE1B" box="[930,1059,449,471]" pageId="42" pageNumber="869">J. Mendoza</collectorName>
, and
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.
<collectorName id="26965339A27FFF86FBC4FE39C695FE1B" box="[1150,1257,448,471]" pageId="42" pageNumber="869">Monjaraz</collectorName>
)] (
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,
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).
<location id="8EBC6034A27FFF86FC4EFE19C73CFE39" LSID="urn:lsid:plazi:treatment:03CA87F9A27FFF87FEA2FC8EC1BFFB9E:8EBC6034A27FFF86FC4EFE19C73CFE39" box="[1012,1344,480,502]" country="Mexico" latitude="22.01677" longLatPrecision="1" longitude="-100.59166" name="Municipio Villa de Zaragoza" pageId="42" pageNumber="869" stateProvince="San Luis Potosi">Municipio Villa de Zaragoza</location>
, small cave from
<collectorName id="26965339A27FFF86FC25FE06C1B4FDD8" box="[927,968,511,532]" pageId="42" pageNumber="869">Km</collectorName>
58
</materialsCitation>
.
<materialsCitation id="3B0B3CB2A27FFF86FC41FE07C7F5FDA3" collectingDate="2012-05-14" collectionCode="CNAN, R" collectorName="J. Cruz &amp; G. Contreras &amp; J. Mendoza &amp; Monjaraz &amp; Cueva de la Laguna &amp; Valle &amp; Fantasmas" country="Mexico" latitude="22.0713" location="Municipio Villa de Zaragoza" longLatPrecision="1" longitude="-100.62499" pageId="42" pageNumber="869" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="San Luis Potosi">
<specimenCount id="9D65FD66A27FFF86FC41FE07C657FDD8" box="[1019,1067,510,532]" pageId="42" pageNumber="869" type="male">2 ♂</specimenCount>
and
<specimenCount id="9D65FD66A27FFF86FBDCFE07C6EEFDD8" box="[1126,1170,510,532]" pageId="42" pageNumber="869" type="female">1 ♀</specimenCount>
[
<collectionCode id="ED72AE2AA27FFF86FB1AFE07C692FDDF" box="[1184,1262,510,531]" pageId="42" pageNumber="869">CNAN</collectionCode>
(
<date id="FFDD102FA27FFF86FB44FE07C710FDDF" box="[1278,1388,510,531]" pageId="42" pageNumber="869" value="2012-05-14">
<collectingDate id="EF99E9C7A27FFF86FB44FE07C710FDDF" box="[1278,1388,510,531]" pageId="42" pageNumber="869" value="2012-05-14">14.v.2012</collectingDate>
</date>
;
<collectorName id="26965339A27FFF86FAC2FE06C11CFDFE" pageId="42" pageNumber="869">J. Cruz</collectorName>
,
<collectorName id="26965339A27FFF86FCCBFDE4C677FDFE" box="[881,1035,541,562]" pageId="42" pageNumber="869">G. Contreras</collectorName>
,
<collectorName id="26965339A27FFF86FBA1FDE4C6DAFDFF" box="[1051,1190,541,563]" pageId="42" pageNumber="869">J. Mendoza</collectorName>
, and
<collectionCode id="ED72AE2AA27FFF86FB56FDE4C77DFDFE" box="[1260,1281,541,562]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="42" pageNumber="869">R</collectionCode>
.
<collectorName id="26965339A27FFF86FAAAFDE5C703FDFF" box="[1296,1407,540,563]" pageId="42" pageNumber="869">Monjaraz</collectorName>
)] (
<geoCoordinate id="EE575028A27FFF86FC94FDC2C1BDFD9D" box="[814,961,571,593]" degrees="22" direction="north" minutes="4" orientation="latitude" pageId="42" pageNumber="869" precision="1" seconds="16.68" value="22.0713">22°416.68″N</geoCoordinate>
,
<geoCoordinate id="EE575028A27FFF86FC74FDC2C6FEFD9D" box="[974,1154,571,593]" degrees="100" direction="west" minutes="37" orientation="longitude" pageId="42" pageNumber="869" precision="1" seconds="29.96" value="-100.62499">100°3729.96″W</geoCoordinate>
).
<location id="8EBC6034A27FFF86FB2EFDC2C108FDA3" LSID="urn:lsid:plazi:treatment:03CA87F9A27FFF87FEA2FC8EC1BFFB9E:8EBC6034A27FFF86FB2EFDC2C108FDA3" country="Mexico" latitude="22.0713" longLatPrecision="1" longitude="-100.62499" name="Municipio Villa de Zaragoza" pageId="42" pageNumber="869" stateProvince="San Luis Potosi">Municipio Villa de Zaragoza</location>
,
<collectorName id="26965339A27FFF86FC3BFDA3C612FDA3" box="[897,1134,602,624]" pageId="42" pageNumber="869">Cueva de la Laguna</collectorName>
,
<collectorName id="26965339A27FFF86FBC0FDA3C6CFFDBC" box="[1146,1203,602,624]" pageId="42" pageNumber="869">Valle</collectorName>
de los
<collectorName id="26965339A27FFF86FABDFDA3C7F5FDA3" box="[1287,1417,602,623]" pageId="42" pageNumber="869">Fantasmas</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36EFA27FFF86FC9CFD67C7F2FD3D" blockId="42.[806,1422,670,753]" pageId="42" pageNumber="869">
<emphasis id="B917EAFDA27FFF86FC9CFD67C1D8FD7F" box="[806,932,670,691]" italics="true" pageId="42" pageNumber="869">Etymology:</emphasis>
Patronym, dedicated to Alejandro Valdez- Mondragón, friend and colleague, who has contribut- ed to the knowledge of spiders and ricinuleids in
<collectingCountry id="F374767FA27FFF86FA81FD22C7F6FD3D" box="[1339,1418,731,753]" name="Mexico" pageId="42" pageNumber="869">Mexico</collectingCountry>
.
</paragraph>
<paragraph id="8BDC36EFA27FFF86FC9CFCE6C6C2FC5C" blockId="42.[806,1421,798,912]" pageId="42" pageNumber="869">
<emphasis id="B917EAFDA27FFF86FC9CFCE6C1E5FCF8" box="[806,921,799,820]" italics="true" pageId="42" pageNumber="869">Diagnosis:</emphasis>
Troglomorphic species. Dorsal tubercles only present in the central portion of mesotergal areas. On male penis the two basal-most pairs of lateral setae very close to each other (
<figureCitation id="13582A6AA27FFF86FBF4FC83C6CDFC5C" box="[1102,1201,890,912]" captionStart="Figure 44" captionStartId="43.[164,243,847,866]" captionTargetBox="[164,1445,195,817]" captionTargetId="figure-431@43.[164,1445,195,817]" captionTargetPageId="43" captionText="Figure 44. Crettaros valdezi sp. nov. male genitalia. A, dorsal view. B, ventral view. C, lateral view. White arrows indicate ventral microsetae in (B) and parastylar setae in (C)." figureDoi="http://doi.org/10.5281/zenodo.5340065" httpUri="https://zenodo.org/record/5340065/files/figure.png" pageId="42" pageNumber="869">Fig. 44B</figureCitation>
).
</paragraph>
<paragraph id="8BDC36EFA27FFF87FC9CFC47C307F8A2" blockId="42.[806,1421,958,1010]" lastBlockId="43.[164,779,932,1902]" lastPageId="43" lastPageNumber="870" pageId="42" pageNumber="869">
<emphasis id="B917EAFDA27FFF86FC9CFC47C617FC1F" box="[806,1131,958,979]" italics="true" pageId="42" pageNumber="869">
Description: Male
<typeStatus id="54D8884DA27FFF86FBBEFC47C619FC1F" box="[1028,1125,958,979]" pageId="42" pageNumber="869" type="holotype">holotype</typeStatus>
:
</emphasis>
Measurements: scutum length 2.6, maximum scutum width 2.2. Legs. I 1.20/ 0.50/0.85/1.10, II 1.50/0.75/1.30/1.35, III 1.30/0.57/1.15/ 1.27, IV 1.80/0.75/1.55/1.97.
<emphasis id="B917EAFDA27EFF87FE64FC3AC042FC14" box="[478,574,963,984]" italics="true" pageId="43" pageNumber="870">Dorsum.</emphasis>
Rugose, with only small spiniform tubercles on the central portion of mesotergal areas IIV. Areas II and III with median humps slightly elevated (
<figureCitation id="13582A6AA27EFF87FE03FBE6C064FBF8" box="[441,536,1054,1077]" captionStart="Figure 63" captionStartId="60.[144,223,1276,1295]" captionTargetBox="[144,1425,196,1246]" captionTargetId="figure-0@60.[144,1425,196,1246]" captionTargetPageId="60" captionText="Figure 63. A, dorsal view of the anal bulge on Chapulobunus poblano sp. nov.; arrow indicates the bulge. B, lateral view of Huasteca silhavyi sp. nov., line indicates the very high lateral channel area. C, laterodorsal view of scutum of Crettaros valdezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. D, posterodorsal view of scutum of Crettaros santibanezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. E, dorsal view of trochanter IV of Karos barbarikos; arrow indicates the tridentate apophysis. F, anterodorsal view of scutum of Karos barbarikos, the black points mark the tubercles forming a V. G, dorsal ornamentation of Karos tersum sp. nov. showing the transversal row of tubercles on mesotergal areas; transversal row II indicated by black dots. H, detail of the ventral ornamentation on femur IV of Karos barbarikos. I, detail of the ectal ornamentation on tibia IV of Karos barbarikos; arrows indicate the spiniform tubercles. J, detail of the ventral ornamentation on leg IV in Chapulobunus poblano sp. nov. K, detail of the two apical spines on femur IV in Chapulobunus poblano sp. nov. These images are not at the same scale." figureDoi="http://doi.org/10.5281/zenodo.5340103" httpUri="https://zenodo.org/record/5340103/files/figure.png" pageId="43" pageNumber="870">Fig. 63C</figureCitation>
). Opisthosoma convex in lateral view. Ocularium in the middle of prosoma, low, slightly projected beyond the eyes, blunt, covered by small tubercles. Lateral clear areas projected from the scutum in triangular shape, apex rounded. Lateral clear areas on apices of area V and free tergites III rounded, slightly protruding. Free tergites with one median row of small tubercles.
<emphasis id="B917EAFDA27EFF87FDA3FB0CC018FAC6" box="[537,612,1269,1290]" italics="true" pageId="43" pageNumber="870">Venter.</emphasis>
Entire venter covered by small rounded tubercles, with a few small setae. Stigmatic area an inverted Y shape, with lateral margins straight, short (
<figureCitation id="13582A6AA27EFF87FE0AFAA9C073FAAB" box="[432,527,1360,1383]" captionStart="Figure 43" captionStartId="42.[144,221,1824,1843]" captionTargetBox="[305,1262,1060,1791]" captionTargetId="figure-377@42.[303,1264,1057,1794]" captionTargetPageId="42" captionText="Figure 43. Crettaros valdezi sp. nov. A, habitus, dorsal view. B, legs IV, mesal view. C, habitus, lateral view. D, habitus, ventral view. E, scutum, dorsal view. F, ocularium, frontal view. Scale bars: A = 2.5 mm, B, D, and E = 1.5 mm, C = 1 mm, F = 0.5 mm. The dark lines on (D) indicate the stigmatic area." figureDoi="http://doi.org/10.5281/zenodo.5340063" httpUri="https://zenodo.org/record/5340063/files/figure.png" pageId="43" pageNumber="870">Fig. 43D</figureCitation>
). Coxa IV with a small dorso-ectal spiniform tooth.
<emphasis id="B917EAFDA27EFF87FE60FA96C032FA48" box="[474,590,1391,1412]" italics="true" pageId="43" pageNumber="870">Pedipalps.</emphasis>
Patella with one mesal spiniform tubercle.
<emphasis id="B917EAFDA27EFF87FE69FA77C072FA6F" box="[467,526,1422,1443]" italics="true" pageId="43" pageNumber="870">Legs.</emphasis>
Very similar in ornamentation and size, covered by numerous small rounded tubercles and setae. Ventral ornamentation of femur IV slightly larger. Femora III and IV curved. Metatarsus II without clear annuli. Tarsal count 4(2):6(3):6:6.
<emphasis id="B917EAFDA27EFF87FF1EF9DEC337F9F0" box="[164,331,1575,1596]" italics="true" pageId="43" pageNumber="870">Male genitalia.</emphasis>
Pars distalis elongated, spatular shape; apical margin almost straight. Lateral setae slender and slightly compressed distally, the two basal-most pairs very close to each other. Two pairs of parastylar setae flanking the follis. Spiniform projections not exposed and inconspicuous (
<figureCitation id="13582A6AA27EFF87FE5EF939C048F91A" box="[484,564,1728,1750]" captionStart="Figure 44" captionStartId="43.[164,243,847,866]" captionTargetBox="[164,1445,195,817]" captionTargetId="figure-431@43.[164,1445,195,817]" captionTargetPageId="43" captionText="Figure 44. Crettaros valdezi sp. nov. male genitalia. A, dorsal view. B, ventral view. C, lateral view. White arrows indicate ventral microsetae in (B) and parastylar setae in (C)." figureDoi="http://doi.org/10.5281/zenodo.5340065" httpUri="https://zenodo.org/record/5340065/files/figure.png" pageId="43" pageNumber="870">Fig. 44</figureCitation>
).
<emphasis id="B917EAFDA27EFF87FDF2F939C177F919" box="[584,779,1728,1749]" italics="true" pageId="43" pageNumber="870">
Female
<typeStatus id="54D8884DA27EFF87FD18F939C179F919" box="[674,773,1728,1749]" pageId="43" pageNumber="870" type="paratype">paratype</typeStatus>
:
</emphasis>
Measurements: scutum length 2.6, maximum scutum width 2.2. Very similar to male, with the following differences: stigmatic area shorter and narrower than on males. Legs IV slightly thinner (
<figureCitation id="13582A6AA27EFF87FD9FF8C3C0D0F883" box="[549,684,1850,1872]" captionStart="Figure 43" captionStartId="42.[144,221,1824,1843]" captionTargetBox="[305,1262,1060,1791]" captionTargetId="figure-377@42.[303,1264,1057,1794]" captionTargetPageId="42" captionText="Figure 43. Crettaros valdezi sp. nov. A, habitus, dorsal view. B, legs IV, mesal view. C, habitus, lateral view. D, habitus, ventral view. E, scutum, dorsal view. F, ocularium, frontal view. Scale bars: A = 2.5 mm, B, D, and E = 1.5 mm, C = 1 mm, F = 0.5 mm. The dark lines on (D) indicate the stigmatic area." figureDoi="http://doi.org/10.5281/zenodo.5340063" httpUri="https://zenodo.org/record/5340063/files/figure.png" pageId="43" pageNumber="870">Fig. 43B, D</figureCitation>
). Tarsal count 4(2):6(3):6:6.
</paragraph>
<caption id="DF1C6667A27FFF86FF2AF8D9C14CF8A2" ID-DOI="http://doi.org/10.5281/zenodo.5340063" ID-Zenodo-Dep="5340063" httpUri="https://zenodo.org/record/5340063/files/figure.png" pageId="42" pageNumber="869" startId="42.[144,221,1824,1843]" targetBox="[305,1262,1060,1791]" targetPageId="42">
<paragraph id="8BDC36EFA27FFF86FF2AF8D9C14CF8A2" blockId="42.[144,1422,1824,1902]" pageId="42" pageNumber="869">
<emphasis id="B917EAFDA27FFF86FF2AF8D9C053F8FF" bold="true" box="[144,559,1824,1843]" pageId="42" pageNumber="869">
Figure 43.
<taxonomicName id="4C634D6CA27FFF86FEB7F8D9C3AEF8FF" authority="Cruz-López &amp; Francke, 2015" authorityName="Cruz-López &amp; Francke" authorityYear="2015" box="[269,466,1824,1843]" class="Arachnida" family="Stygnopsidae" genus="Crettaros" kingdom="Animalia" order="Opiliones" pageId="42" pageNumber="869" phylum="Arthropoda" rank="species" species="valdezi" status="sp. nov.">
<emphasis id="B917EAFDA27FFF86FEB7F8D9C3AEF8FF" bold="true" box="[269,466,1824,1843]" italics="true" pageId="42" pageNumber="869">Crettaros valdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2245786A27FFF86FE63F8D9C053F8FF" box="[473,559,1824,1843]" pageId="42" pageNumber="869" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, habitus, dorsal view. B, legs IV, mesal view. C, habitus, lateral view. D, habitus, ventral view. E, scutum, dorsal view. F, ocularium, frontal view. Scale bars: A = 2.5 mm, B, D, and E = 1.5 mm, C = 1 mm, F = 0.5 mm. The dark lines on (D) indicate the stigmatic area.
</paragraph>
</caption>
<caption id="DF1C6667A27EFF87FF1EFCB6C13CFC4C" ID-DOI="http://doi.org/10.5281/zenodo.5340065" ID-Zenodo-Dep="5340065" httpUri="https://zenodo.org/record/5340065/files/figure.png" pageId="43" pageNumber="870" startId="43.[164,243,847,866]" targetBox="[164,1445,195,817]" targetPageId="43">
<paragraph id="8BDC36EFA27EFF87FF1EFCB6C13CFC4C" blockId="43.[164,1441,847,896]" pageId="43" pageNumber="870">
<emphasis id="B917EAFDA27EFF87FF1EFCB6C02FFCAF" bold="true" box="[164,595,847,867]" pageId="43" pageNumber="870">
Figure 44.
<taxonomicName id="4C634D6CA27EFF87FE9EFCB6C38CFCAE" authority="Cruz-López &amp; Francke, 2015" authorityName="Cruz-López &amp; Francke" authorityYear="2015" box="[292,496,847,866]" class="Arachnida" family="Stygnopsidae" genus="Crettaros" kingdom="Animalia" order="Opiliones" pageId="43" pageNumber="870" phylum="Arthropoda" rank="species" species="valdezi" status="sp. nov.">
<emphasis id="B917EAFDA27EFF87FE9EFCB6C38CFCAE" bold="true" box="[292,496,847,866]" italics="true" pageId="43" pageNumber="870">Crettaros valdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2245786A27EFF87FE40FCA9C02FFCAF" box="[506,595,848,867]" pageId="43" pageNumber="870" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male genitalia. A, dorsal view. B, ventral view. C, lateral view. White arrows indicate ventral microsetae in (B) and parastylar setae in (C).
</paragraph>
</caption>
<paragraph id="8BDC36EFA27EFF87FC80FC5DC1BFFB9E" blockId="43.[826,1441,932,1106]" pageId="43" pageNumber="870">
<emphasis id="B917EAFDA27EFF87FC80FC5DC18AFC75" box="[826,1014,932,953]" italics="true" pageId="43" pageNumber="870">Natural history:</emphasis>
All specimens were collected under rocks in the dark zone, never found outside the cave. Generally,
<specimenCount id="9D65FD66A27EFF87FC00FC1BC62BFC3B" box="[954,1111,993,1015]" pageId="43" pageNumber="870" type="generic">one specimen</specimenCount>
was found under one rock, rarely two or three. When uncovered they showed thanatosis behaviour similar to that of
<taxonomicName id="4C634D6CA27EFF87FA88FBE6C1C0FB9E" authorityName="Cruz-López &amp; Francke" authorityYear="2015" class="Arachnida" family="Stygnopsidae" genus="Crettaros" kingdom="Animalia" order="Opiliones" pageId="43" pageNumber="870" phylum="Arthropoda" rank="species" species="santibanezi">
<emphasis id="B917EAFDA27EFF87FA88FBE6C1C0FB9E" italics="true" pageId="43" pageNumber="870">Crettaros santibanezi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>