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<taxonomicName id="4C634D67FFF8FFC7FE2E46A4BBDF3EF0" ID-CoL="4FSJD" authority="Moctezuma, Sanchez-Huerta &amp; Halffter, 2017" authorityName="Moctezuma, Sanchez-Huerta &amp; Halffter" authorityYear="2017" box="[390,1197,1199,1226]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bravoensis">
<emphasis id="B917EAF6FFF8FFC7FE2E46A4BDF53EF0" bold="true" box="[390,647,1199,1226]" italics="true" pageId="6" pageNumber="7">Phanaeus bravoensis</emphasis>
<bibRefCitation id="EFF24B15FFF8FFC7FD2746BBBBDF3EF0" author="Moctezuma V. &amp; Sanchez-Huerta J. L. &amp; Halffter G." box="[655,1197,1199,1226]" pageId="6" pageNumber="7" pagination="113 - 135" refId="ref40712" refString="Moctezuma V., Sanchez-Huerta J. L. &amp; Halffter G. 2017. Two new species of the Phanaeus endymion species group (Coleoptera, Scarabaeidae, Scarabaeinae). ZooKeys 702: 113 - 135. https: // doi. org / 10.3897 / zookeys. 702.14728" type="journal article" year="2017">Moctezuma, Sánchez-Huerta &amp; Halffter, 2017</bibRefCitation>
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<paragraph id="8BDC36E4FFF8FFC7FD3D46D8BCEC3ED7" blockId="6.[390,1197,1199,1262]" box="[661,926,1235,1262]" pageId="6" pageNumber="7">
<figureCitation id="13582A61FFF8FFC7FD3D46D8BD873ED7" box="[661,757,1235,1261]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma &amp; Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="6" pageNumber="7">Figs 1A</figureCitation>
,
<figureCitation id="13582A61FFF8FFC7FCAA46D8BC533ED4" box="[770,801,1235,1262]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017;P. edmondsi Moctezuma, Deloya &amp; Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="6" pageNumber="7">15</figureCitation>
,
<figureCitation id="13582A61FFF8FFC7FC8646D8BC133ED7" box="[814,865,1235,1261]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma &amp; Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="6" pageNumber="7">18B</figureCitation>
,
<figureCitation id="13582A61FFF8FFC7FCC546D8BCEC3ED7" box="[877,926,1235,1261]" captionStart="Fig" captionStartId="56.[189,231,1678,1704]" captionTargetBox="[306,1282,265,1646]" captionTargetId="figure-158@56.[306,1282,265,1646]" captionTargetPageId="56" captionText="Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma &amp; Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836804" httpUri="https://zenodo.org/record/4836804/files/figure.png" pageId="6" pageNumber="7">19B</figureCitation>
</paragraph>
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<paragraph id="8BDC36E4FFF8FFC7FF15471EBC643F0A" blockId="6.[189,790,1301,1328]" box="[189,790,1301,1328]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB7311CAFFF8FFC7FF15471EBC643F0A" box="[189,790,1301,1328]" pageId="6" pageNumber="7">
<treatmentCitation id="0AC210F5FFF8FFC7FF15471EBC643F0A" LSID="urn:lsid:zoobank.org:act:41BF39B8-9D18-4275-9A1A-01BA830C3867" author="Moctezuma et al." box="[189,790,1301,1328]" httpUri="http://treatment.plazi.org/id/E8980C9FB007F63E573701F45D7C30AD" page="115" pageId="6" pageNumber="7" year="2017">
<taxonomicName id="4C634D67FFF8FFC7FF15471EBC603F0A" ID-CoL="4FSJD" authority="Moctezuma et al., 2017: 115" authorityName="Moctezuma, Sanchez-Huerta &amp; Halffter" authorityPageNumber="115" authorityYear="2017" box="[189,786,1301,1328]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bravoensis">
<emphasis id="B917EAF6FFF8FFC7FF15471EBEC53F15" box="[189,439,1301,1327]" italics="true" pageId="6" pageNumber="7">Phanaeus bravoensis</emphasis>
Moctezuma
<emphasis id="B917EAF6FFF8FFC7FDF8471DBDF63F15" box="[592,644,1301,1327]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
., 2017: 115
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.
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</paragraph>
<paragraph id="8BDC36E4FFF8FFC7FF15475CBBA33F48" blockId="6.[189,1233,1367,1394]" box="[189,1233,1367,1394]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB7311CAFFF8FFC7FF15475CBBA33F48" box="[189,1233,1367,1394]" pageId="6" pageNumber="7">
<taxonomicName id="4C634D67FFF8FFC7FF15475CBEC53F4B" ID-CoL="4FSJD" authorityName="Moctezuma, Sanchez-Huerta &amp; Halffter" authorityYear="2017" box="[189,439,1367,1393]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bravoensis">
<emphasis id="B917EAF6FFF8FFC7FF15475CBEC53F4B" box="[189,439,1367,1393]" italics="true" pageId="6" pageNumber="7">Phanaeus bravoensis</emphasis>
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–
<treatmentCitation id="0AC210F5FFF8FFC7FE7C475CBC773F4B" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." box="[468,773,1367,1394]" httpUri="http://treatment.plazi.org/id/DE1922127F68F012FF66FAB790EF4555" page="69" pageId="6" pageNumber="7" year="2018">
<bibRefCitation id="EFF24B15FFF8FFC7FE7C475CBC773F4B" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." box="[468,773,1367,1394]" pageId="6" pageNumber="7" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. &amp; Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
Kohlmann
<emphasis id="B917EAF6FFF8FFC7FDF04753BDF93F4B" box="[600,651,1367,1393]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2018: 69
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AC210F5FFF8FFC7FC9C4753BBF33F48" author="Moctezuma V. &amp; Deloya C. &amp; Sanchez-Huerta J. L. &amp; Halffter G." box="[820,1153,1367,1394]" page="253" pageId="6" pageNumber="7" year="2019">
<bibRefCitation id="EFF24B15FFF8FFC7FC9C4753BBF33F48" author="Moctezuma V. &amp; Deloya C. &amp; Sanchez-Huerta J. L. &amp; Halffter G." box="[820,1153,1367,1394]" pageId="6" pageNumber="7" pagination="249 - 254" refId="ref40763" refString="Moctezuma V., Deloya C., Sanchez-Huerta J. L. &amp; Halffter G. 2019. A new species of the Phanaeus endymion species group (Coleoptera: Scarabaeidae: Scarabaeinae), with comments on ecology and distribution. Annales de la Societe entomologique de France (N. S.) 55: 249 - 254. https: // doi. org / 10.1080 / 00379271.2019.1577170" type="journal article" year="2019">
Moctezuma
<emphasis id="B917EAF6FFF8FFC7FC6E4753BC883F4B" box="[966,1018,1367,1393]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2019: 253
</bibRefCitation>
</treatmentCitation>
, fig. 5.
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</paragraph>
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<subSubSection id="C379656FFFF8FFC7FF154792BDAD3CA1" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BDC36E4FFF8FFC7FF154792BE1E3F89" blockId="6.[189,364,1433,1459]" box="[189,364,1433,1459]" pageId="6" pageNumber="7">
<heading id="D0948188FFF8FFC7FF154792BE1E3F89" bold="true" box="[189,364,1433,1459]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B917EAF6FFF8FFC7FF154792BE1E3F89" bold="true" box="[189,364,1433,1459]" pageId="6" pageNumber="7">Type material</emphasis>
</heading>
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<emphasis id="B917EAF6FFF8FFC7FF4247C3BE2E3FD8" bold="true" box="[234,348,1480,1506]" pageId="6" pageNumber="7">
<typeStatus id="54D88846FFF8FFC7FF4247C3BE2E3FD8" box="[234,348,1480,1506]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
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<materialsCitation id="3B0B3CB9FFF8FFC7FF1547E7BE793C10" ID-GBIF-Occurrence="3093980407" collectionCode="IEXA" country="MEXCIO" county="Moctezuma" location="Chilpancingo de los Bravo" municipality="Chilpancingo" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Guerrero" typeStatus="holotype">
<collectingCountry id="F3747674FFF8FFC7FF1547E7BE433C3C" box="[189,305,1516,1542]" name="Mexico" pageId="6" pageNumber="7">MEXCIO</collectingCountry>
•
<specimenCount id="9D65FD6DFFF8FFC7FEE547E6BE113C3D" box="[333,355,1517,1543]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
, Moctezuma
<emphasis id="B917EAF6FFF8FFC7FDA147E6BD4E3C3C" box="[521,572,1516,1542]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2017: 118, figs 6–8 (revised);
<collectingRegion id="49A7F806FFF8FFC7FC1D47E7BB503C3C" box="[949,1058,1516,1542]" country="Mexico" name="Guerrero" pageId="6" pageNumber="7">Guerrero</collectingRegion>
,
<location id="8EBC603FFFF8FFC7FB9847E7BA003C3D" LSID="urn:lsid:plazi:treatment:03CA87F2FFF8FFC6FE2E46A4BEC23B95:8EBC603FFFF8FFC7FB9847E7BA003C3D" box="[1072,1394,1516,1543]" country="MEXCIO" county="Moctezuma" municipality="Chilpancingo" name="Chilpancingo de los Bravo" pageId="6" pageNumber="7" stateProvince="Guerrero">Chilpancingo de los Bravo</location>
;
<collectionCode id="ED72AE21FFF8FFC7FF15441BBE753C10" box="[189,263,1552,1578]" pageId="6" pageNumber="7">IEXA</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BDC36E4FFF8FFC7FF154459BE2F3C51" blockId="6.[189,349,1617,1644]" box="[189,349,1617,1644]" pageId="6" pageNumber="7">
<heading id="D0948188FFF8FFC7FF154459BE2F3C51" bold="true" box="[189,349,1617,1644]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B917EAF6FFF8FFC7FF154459BE2F3C51" bold="true" box="[189,349,1617,1644]" pageId="6" pageNumber="7">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFF8FFC7FF15448ABDAD3CA1" blockId="6.[189,735,1665,1691]" box="[189,735,1665,1691]" pageId="6" pageNumber="7">
<collectingCountry id="F3747674FFF8FFC7FF15448ABE683CA1" box="[189,282,1665,1691]" name="Mexico" pageId="6" pageNumber="7">Mexico</collectingCountry>
,
<collectingRegion id="49A7F806FFF8FFC7FE8D448ABEE03CA1" box="[293,402,1665,1691]" country="Mexico" name="Guerrero" pageId="6" pageNumber="7">Guerrero</collectingRegion>
, Chilpancingo de los Bravo.
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFF8FFC7FF1544C8BC2B3D37" box="[189,857,1731,1805]" pageId="6" pageNumber="7" type="distribution">
<paragraph id="8BDC36E4FFF8FFC7FF1544C8BE243CE7" blockId="6.[189,342,1731,1757]" box="[189,342,1731,1757]" pageId="6" pageNumber="7">
<heading id="D0948188FFF8FFC7FF1544C8BE243CE7" bold="true" box="[189,342,1731,1757]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B917EAF6FFF8FFC7FF1544C8BE243CE7" bold="true" box="[189,342,1731,1757]" pageId="6" pageNumber="7">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFF8FFC7FF1544F9BC2B3D37" blockId="6.[189,857,1778,1805]" box="[189,857,1778,1805]" pageId="6" pageNumber="7">
<collectingCountry id="F3747674FFF8FFC7FF1544F9BE683D36" box="[189,282,1778,1804]" name="Mexico" pageId="6" pageNumber="7">Mexico</collectingCountry>
, Sierra Madre del sur, central
<collectingRegion id="49A7F806FFF8FFC7FDD244F9BD963D36" box="[634,740,1778,1804]" country="Mexico" name="Guerrero" pageId="6" pageNumber="7">Guerrero</collectingRegion>
(
<figureCitation id="13582A61FFF8FFC7FD5C44F9BC393D37" box="[756,843,1778,1805]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017;P. edmondsi Moctezuma, Deloya &amp; Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="6" pageNumber="7">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFF8FFC6FF15453FBEC23B95" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8BDC36E4FFF8FFC7FF15453FBE5D3D74" blockId="6.[189,303,1844,1870]" box="[189,303,1844,1870]" pageId="6" pageNumber="7">
<heading id="D0948188FFF8FFC7FF15453FBE5D3D74" bold="true" box="[189,303,1844,1870]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B917EAF6FFF8FFC7FF15453FBE5D3D74" bold="true" box="[189,303,1844,1870]" pageId="6" pageNumber="7">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFF8FFC6FF154568BEC23B95" blockId="6.[188,1400,1891,2025]" lastBlockId="7.[189,1401,265,431]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
This species was recently described by Moctezuma
<emphasis id="B917EAF6FFF8FFC7FCB1456FBC3E3D47" box="[793,844,1891,1917]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. (2017). To the original description we add that the elytral striae are impressed basally as distinct fossae; right lobe of endophallite copulatrix is more developed than left lobe; right lobe of endophallite copulatrix obtusely triangular, arched superiorly; left lobe of endophallite copulatrix lobed, strongly reduced; central ridge more developed than central column (
<figureCitation id="13582A61FFF9FFC6FE8E4302BEF63B19" box="[294,388,265,291]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma &amp; Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="7" pageNumber="8">Fig. 1A</figureCitation>
). For the female, the trituberculate cephalic carina with nearly aligned, similar sized, conical tubercles; frons punctures scarce, almost effaced; pronotal surface distinctly punctate; pronotal process trituberculate with a posterior concavity; all pronotal tubercles rounded, with middle tubercle slightly more developed and projected frontally than lateral tubercles; posterior pronotal midline distinctly impressed.
</paragraph>
</subSubSection>
</treatment>
</document>
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