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<document id="36B08272F9CE9B2E52512969EBB18E4C" ID-CLB-Dataset="7153" ID-DOI="10.5852/ejt.2021.747.1333" ID-GBIF-Dataset="aca67cc8-96ca-4f24-b8af-2b532ecd355c" ID-ISSN="2118-9773" ID-Zenodo-Dep="4836744" ID-ZooBank="3F0B6EAF-C616-4865-811A-414A094B590C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619605517524" checkinUser="plazi" docAuthor="Moctezuma, Victor &amp; Halffter, Gonzalo" docDate="2021" docId="03CA87F2FFE8FFD8FDDA448BBDA9398B" docLanguage="en" docName="ejt-747_1-71.pdf" docOrigin="European Journal of Taxonomy 747" docSource="http://zoobank.org/3f0b6eaf-c616-4865-811a-414a094b590c" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phanaeus jackenioi Moctezuma &amp; Halffter 2021, sp. nov." docType="treatment" docUuid="CF4B33E3-1245-4631-A019-2CAF43709603" docUuidSource="ZooBank" docVersion="10" lastPageNumber="26" masterDocId="FFF3FF8AFFFEFFC1FFA8420BBF723A3A" masterDocTitle="Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species" masterLastPageNumber="71" masterPageNumber="1" pageNumber="23" updateTime="1698937721231" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3908367415624224447D1F093DEE8A24">Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species</mods:title>
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<mods:namePart id="73A3E6C87F467A0D2F662BA1B46D2D5E">Moctezuma, Victor</mods:namePart>
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<mods:affiliation id="FE666279772DE88756933B38426BFD80">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:namePart id="DCD183D40462A9DEA987075E3B58B3A4">Halffter, Gonzalo</mods:namePart>
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<mods:affiliation id="0A01B43536BE03D1114F34C864E43ADF">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:title id="5E8234A621A238F031F970CA7B4103A4">European Journal of Taxonomy</mods:title>
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<mods:date id="2F1D6F37E32DE07753DE73439860805D">2021</mods:date>
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<mods:number id="8B680E1908B3EC72D341CCCAE8B8BED0">2021-04-28</mods:number>
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<treatment id="03CA87F2FFE8FFD8FDDA448BBDA9398B" ID-DOI="http://doi.org/10.5281/zenodo.4726227" ID-GBIF-Taxon="181753770" ID-Zenodo-Dep="4726227" LSID="urn:lsid:zoobank.org:act:CF4B33E3-1245-4631-A019-2CAF43709603" httpUri="http://treatment.plazi.org/id/03CA87F2FFE8FFD8FDDA448BBDA9398B" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23">
<subSubSection id="C379656FFFE8FFD7FDDA448BBCB03CA1" box="[626,962,1664,1691]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BDC36E4FFE8FFD7FDDA448BBCB03CA1" blockId="22.[376,1211,1664,1764]" box="[626,962,1664,1691]" pageId="22" pageNumber="23">
<heading id="D0948188FFE8FFD7FDDA448BBCB03CA1" box="[626,962,1664,1691]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="23" reason="2">
<taxonomicName id="4C634D67FFE8FFD7FDDA448BBC133CA1" ID-CoL="874F5" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[626,865,1664,1691]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="jackenioi" status="sp. nov.">
<emphasis id="B917EAF6FFE8FFD7FDDA448BBC133CA1" bold="true" box="[626,865,1664,1691]" italics="true" pageId="22" pageNumber="23">Phanaeus jackenioi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224578DFFE8FFD7FCC0448ABCB03CA1" box="[872,962,1665,1691]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C379656FFFE8FFD7FED044AEBCCE3CDE" pageId="22" pageNumber="23" type="description">
<paragraph id="8BDC36E4FFE8FFD7FED044AEBBC93CFA" blockId="22.[376,1211,1664,1764]" box="[376,1211,1701,1728]" pageId="22" pageNumber="23">
<uri id="FFF23AE6FFE8FFD7FED044AEBBC93CFA" box="[376,1211,1701,1728]" pageId="22" pageNumber="23">
urn:lsid:zoobank.org:act:
<uuid id="FFC50C31FFE8FFD7FD3E44AEBBC93CFA" box="[662,1211,1701,1728]" pageId="22" pageNumber="23">CF4B33E3-1245-4631-A019-2CAF43709603</uuid>
</uri>
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<paragraph id="8BDC36E4FFE8FFD7FDDF44C2BCCE3CDE" blockId="22.[376,1211,1664,1764]" box="[631,956,1737,1764]" pageId="22" pageNumber="23">
<figureCitation id="13582A61FFE8FFD7FDDF44C2BDA53CDE" box="[631,727,1737,1764]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma &amp; Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="22" pageNumber="23">Figs 1H</figureCitation>
,
<figureCitation id="13582A61FFE8FFD7FD4C44C1BC783CDE" box="[740,778,1738,1764]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="22" pageNumber="23">2D</figureCitation>
,
<figureCitation id="13582A61FFE8FFD7FCBE44C2BC553CD9" box="[790,807,1737,1763]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="22" pageNumber="23">6</figureCitation>
,
<figureCitation id="13582A61FFE8FFD7FC9B44C2BC203CDE" box="[819,850,1737,1764]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017;P. edmondsi Moctezuma, Deloya &amp; Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="22" pageNumber="23">15</figureCitation>
,
<figureCitation id="13582A61FFE8FFD7FCF744C1BCFB3CDE" box="[863,905,1738,1764]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma &amp; Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="22" pageNumber="23">18I</figureCitation>
,
<figureCitation id="13582A61FFE8FFD7FC3D44C1BCCE3CDE" box="[917,956,1738,1764]" captionStart="Fig" captionStartId="56.[189,231,1678,1704]" captionTargetBox="[306,1282,265,1646]" captionTargetId="figure-158@56.[306,1282,265,1646]" captionTargetPageId="56" captionText="Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma &amp; Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836804" httpUri="https://zenodo.org/record/4836804/files/figure.png" pageId="22" pageNumber="23">19I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFE8FFD7FF154507BC273DD3" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BDC36E4FFE8FFD7FF154507BE443D1C" blockId="22.[189,310,1804,1830]" box="[189,310,1804,1830]" pageId="22" pageNumber="23">
<heading id="D0948188FFE8FFD7FF154507BE443D1C" bold="true" box="[189,310,1804,1830]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B917EAF6FFE8FFD7FF154507BE443D1C" bold="true" box="[189,310,1804,1830]" pageId="22" pageNumber="23">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE8FFD7FF154537BC273DD3" blockId="22.[189,1398,1852,2025]" pageId="22" pageNumber="23">
Easily diagnosed species by the bright metallic green colour; striae wide, roughened, impressed basally as distinct fossae, with distinctly impressed punctation; the major male with posterolateral angles of pronotum sharply acute, elongate, projected posterolaterally (
<figureCitation id="13582A61FFE8FFD7FC03458EBB603DA5" box="[939,1042,1925,1951]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="22" pageNumber="23">Figs 2D</figureCitation>
,
<figureCitation id="13582A61FFE8FFD7FB8B458EBB383D9A" box="[1059,1098,1925,1952]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="22" pageNumber="23">6A</figureCitation>
); the major female with pronotal process without frontal concavity; rounded pronotal tubercles, nearly aligned, with middle tubercle more developed than lateral tubercles (
<figureCitation id="13582A61FFE8FFD7FD4345C5BC353DD3" box="[747,839,1998,2025]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="22" pageNumber="23">Fig. 6B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFE9FFD6FF154302BA2F3B4F" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BDC36E4FFE9FFD6FF154302BE363B19" blockId="23.[189,324,265,291]" box="[189,324,265,291]" pageId="23" pageNumber="24">
<heading id="D0948188FFE9FFD6FF154302BE363B19" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B917EAF6FFE9FFD6FF154302BE363B19" bold="true" box="[189,324,265,291]" pageId="23" pageNumber="24">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF154333BA2F3B4F" blockId="23.[189,1398,311,373]" pageId="23" pageNumber="24">
We are honoured to dedicate this new species to Jack Schuster and Enio Cano. They have significantly contributed to the knowledge of the Guatemalan scarab beetles, particularly of the family
<taxonomicName id="4C634D67FFE9FFD6FB734350BA2B3B4F" authorityName="Leach" authorityYear="1815" box="[1243,1369,347,373]" class="Insecta" family="Passalidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFE9FFD6FF154397BEA03E9F" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BDC36E4FFE9FFD6FF154397BD553B8D" blockId="23.[189,551,412,439]" box="[189,551,412,439]" pageId="23" pageNumber="24">
<emphasis id="B917EAF6FFE9FFD6FF154397BE1E3B8C" bold="true" box="[189,364,412,438]" pageId="23" pageNumber="24">Type material</emphasis>
(
<specimenCount id="9D65FD6DFFE9FFD6FED44397BEBC3B8D" box="[380,462,412,439]" pageId="23" pageNumber="24" type="male">11 ♂♂</specimenCount>
,
<specimenCount id="9D65FD6DFFE9FFD6FE734397BD693B8D" box="[475,539,412,439]" pageId="23" pageNumber="24" type="female">5 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF4243C0BE2E3BDF" blockId="23.[189,1103,459,521]" box="[234,348,459,485]" pageId="23" pageNumber="24">
<emphasis id="B917EAF6FFE9FFD6FF4243C0BE2E3BDF" bold="true" box="[234,348,459,485]" pageId="23" pageNumber="24">
<typeStatus id="54D88846FFE9FFD6FF4243C0BE2E3BDF" box="[234,348,459,485]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF1543E4BB3D3833" blockId="23.[189,1103,459,521]" box="[189,1103,494,521]" pageId="23" pageNumber="24">
<materialsCitation id="3B0B3CB9FFE9FFD6FF1543E4BB3D3833" ID-GBIF-Occurrence="3093980541" box="[189,1103,494,521]" collectingDate="1978-11-10" collectionCode="UVGC" country="Guatemala" location="Eugenia de Minondo" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Guatemala" typeStatus="holotype">
<collectingCountry id="F3747674FFE9FFD6FF1543E4BE063833" box="[189,372,495,521]" name="Guatemala" pageId="23" pageNumber="24">GUATEMALA</collectingCountry>
•
<specimenCount id="9D65FD6DFFE9FFD6FE2343E4BED63833" box="[395,420,495,521]" pageId="23" pageNumber="24" type="male">♂</specimenCount>
;
<collectingRegion id="49A7F806FFE9FFD6FE1843E5BD5D3832" box="[432,559,494,520]" country="Guatemala" name="Guatemala" pageId="23" pageNumber="24">Guatemala</collectingRegion>
; “
<date id="FFDD1024FFE9FFD6FDE143E4BDA53833" box="[585,727,495,521]" pageId="23" pageNumber="24" value="1978-11-10">
<collectingDate id="EF99E9CCFFE9FFD6FDE143E4BDA53833" box="[585,727,495,521]" pageId="23" pageNumber="24" value="1978-11-10">10/XI/1978</collectingDate>
</date>
.
<location id="8EBC603FFFE9FFD6FD4A43E5BCAC3832" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FD4A43E5BCAC3832" box="[738,990,494,520]" country="Guatemala" name="Eugenia de Minondo" pageId="23" pageNumber="24" stateProvince="Guatemala">Eugenia de Minondo</location>
”;
<collectionCode id="ED72AE21FFE9FFD6FC5C43E4BB393833" box="[1012,1099,495,521]" collectionName="Guatemala, Guatemala City, Universidad del Valle de Guatemala, Collecion de Artropodos" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="23" pageNumber="24">UVGC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF42403BBE153870" blockId="23.[189,1399,560,1012]" box="[234,359,560,586]" pageId="23" pageNumber="24">
<emphasis id="B917EAF6FFE9FFD6FF42403BBE153870" bold="true" box="[234,359,560,586]" pageId="23" pageNumber="24">
<typeStatus id="54D88846FFE9FFD6FF42403BBE153870" box="[234,359,560,586]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF15405FBA7639C9" blockId="23.[189,1399,560,1012]" pageId="23" pageNumber="24">
<materialsCitation id="3B0B3CB9FFE9FFD6FF15405FBE7638AB" ID-GBIF-Occurrence="3093980464" collectingDate="1978-09-13" collectionCode="VMC" country="Guatemala" location="El Injertal" municipality="Parramos" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Chimaltenango" typeStatus="paratype">
<collectingCountry id="F3747674FFE9FFD6FF15405FBE003854" box="[189,370,596,622]" name="Guatemala" pageId="23" pageNumber="24">GUATEMALA</collectingCountry>
–
<emphasis id="B917EAF6FFE9FFD6FE3D4058BD273857" bold="true" box="[405,597,595,621]" pageId="23" pageNumber="24">
<collectingRegion id="49A7F806FFE9FFD6FE3D4058BD273857" box="[405,597,595,621]" country="Guatemala" name="Chimaltenango" pageId="23" pageNumber="24">Chimaltenango</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFE9FFD6FDDC405FBDD63854" box="[628,676,596,622]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
; “
<collectingMunicipality id="6BB8AC9EFFE9FFD6FD17405FBC463854" box="[703,820,596,622]" pageId="23" pageNumber="24">Parramos</collectingMunicipality>
,
<location id="8EBC603FFFE9FFD6FCEA4058BCB23857" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FCEA4058BCB23857" box="[834,960,595,621]" country="Guatemala" municipality="Parramos" name="El Injertal" pageId="23" pageNumber="24" stateProvince="Chimaltenango">El Injertal</location>
.
<date id="FFDD1024FFE9FFD6FC65405FBB2A3854" box="[973,1112,596,622]" pageId="23" pageNumber="24" value="1978-09-13">
<collectingDate id="EF99E9CCFFE9FFD6FC65405FBB2A3854" box="[973,1112,596,622]" pageId="23" pageNumber="24" value="1978-09-13">13/IX/1978</collectingDate>
</date>
.
<location id="8EBC603FFFE9FFD6FBCE4058BA1A3857" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FBCE4058BA1A3857" box="[1126,1384,595,621]" country="Guatemala" municipality="Parramos" name="Eugenia de Minondo" pageId="23" pageNumber="24" stateProvince="Chimaltenango">Eugenia de Minondo</location>
”;
<collectionCode id="ED72AE21FFE9FFD6FF15407CBE7638AB" box="[189,260,631,657]" pageId="23" pageNumber="24">VMC</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B0B3CB9FFE9FFD6FE80407CBB3138AB" ID-GBIF-Occurrence="3093980607" box="[296,1091,631,657]" collectingDate="2007-02-07" collectingDateMax="2007-02-14" collectingDateMin="2007-02-07" collectionCode="VMC" collectorName="R. Chicas" country="Guatemala" location="Chiquimula." pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Chiquimula" typeStatus="paratype">
<emphasis id="B917EAF6FFE9FFD6FE80407CBECC38AB" bold="true" box="[296,446,631,657]" pageId="23" pageNumber="24">
<collectingRegion id="49A7F806FFE9FFD6FE80407CBECC38AB" box="[296,446,631,657]" country="Guatemala" name="Chiquimula" pageId="23" pageNumber="24">Chiquimula</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFE9FFD6FE72407CBD7838AB" box="[474,522,631,657]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFE9FFD6FD8A407CBDC538AB" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FD8A407CBDC538AB" box="[546,695,631,657]" country="Guatemala" name="Chiquimula." pageId="23" pageNumber="24" stateProvince="Chiquimula">Chiquimula.</location>
<date id="FFDD1024FFE9FFD6FD68407CBC2C38AB" box="[704,862,631,657]" pageId="23" pageNumber="24" value="2007-02-07" valueMax="2007-02-14" valueMin="2007-02-07">
<collectingDate id="EF99E9CCFFE9FFD6FD68407CBC2C38AB" box="[704,862,631,657]" pageId="23" pageNumber="24" value="2007-02-07" valueMax="2007-02-14" valueMin="2007-02-07">7–14 II 2007</collectingDate>
</date>
.
<collectorName id="26965332FFE9FFD6FCC3407CBC9638AB" box="[875,996,631,657]" pageId="23" pageNumber="24">R. Chicas</collectorName>
”;
<collectionCode id="ED72AE21FFE9FFD6FC54407CBB3138AB" box="[1020,1091,631,657]" pageId="23" pageNumber="24">VMC</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B0B3CB9FFE9FFD6FBCE407CBBCF388E" ID-GBIF-Occurrence="3093980575" collectingDate="1993-06-22" collectionCode="VMC" collectorName="E. Poll" country="Guatemala" location="Barranca Sta. Catarina. Pinula" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Guatemala" typeStatus="paratype">
<emphasis id="B917EAF6FFE9FFD6FBCE407CBB8338AB" bold="true" box="[1126,1265,631,657]" pageId="23" pageNumber="24">
<collectingRegion id="49A7F806FFE9FFD6FBCE407CBB8338AB" box="[1126,1265,631,657]" country="Guatemala" name="Guatemala" pageId="23" pageNumber="24">Guatemala</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFE9FFD6FAA5407CBA4F38AB" box="[1293,1341,631,657]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; “nr.
<location id="8EBC603FFFE9FFD6FF154090BD6D388E" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FF154090BD6D388E" box="[189,543,666,693]" country="Guatemala" name="Barranca Sta. Catarina. Pinula" pageId="23" pageNumber="24" stateProvince="Guatemala">Barranca Sta. Catarina. Pinula</location>
, Z-14.
<date id="FFDD1024FFE9FFD6FDD84091BD84388E" box="[624,758,666,693]" pageId="23" pageNumber="24" value="1993-06-22">
<collectingDate id="EF99E9CCFFE9FFD6FDD84091BD84388E" box="[624,758,666,693]" pageId="23" pageNumber="24" value="1993-06-22">22 VI 1993</collectingDate>
</date>
.
<collectorName id="26965332FFE9FFD6FCA94090BC3D388E" box="[769,847,666,693]" pageId="23" pageNumber="24">E. Póll</collectorName>
leg. Luz incandescente”;
<collectionCode id="ED72AE21FFE9FFD6FBD24091BBCF388E" box="[1146,1213,666,692]" pageId="23" pageNumber="24">VMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE9FFD6FB7C4091BD5C38E2" ID-GBIF-Occurrence="3093980431" collectingDate="1982-10-13" collectionCode="TAMU" collectorName="S. Roesch" country="Guatemala" location="Ciudad." pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Guatemala" typeStatus="paratype">
<specimenCount id="9D65FD6DFFE9FFD6FB7C4091BA70388F" box="[1236,1282,666,693]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFE9FFD6FAB04091BA05388E" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FAB04091BA05388E" box="[1304,1399,666,692]" country="Guatemala" name="Ciudad." pageId="23" pageNumber="24" stateProvince="Guatemala">Ciudad.</location>
<date id="FFDD1024FFE9FFD6FF1540B5BE4F38E2" box="[189,317,702,728]" pageId="23" pageNumber="24" value="1982-10-13">
<collectingDate id="EF99E9CCFFE9FFD6FF1540B5BE4F38E2" box="[189,317,702,728]" pageId="23" pageNumber="24" value="1982-10-13">13 X 1982</collectingDate>
</date>
.
<collectorName id="26965332FFE9FFD6FEE040B5BEB638E2" box="[328,452,702,728]" pageId="23" pageNumber="24">S. Roesch</collectorName>
”;
<collectionCode id="ED72AE21FFE9FFD6FE7240B5BD5C38E2" box="[474,558,702,728]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="23" pageNumber="24">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE9FFD6FDEF40B5BBD738E2" ID-GBIF-Occurrence="3093980352" box="[583,1189,702,728]" collectingDate="1984-08" collectorName="J. Perez" country="Guatemala" location="Guatemala." pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Guatemala" typeStatus="paratype">
<specimenCount id="9D65FD6DFFE9FFD6FDEF40B5BD0538E2" box="[583,631,702,728]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFE9FFD6FD2540B5BC6538E2" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FD2540B5BC6538E2" box="[653,791,702,728]" country="Guatemala" name="Guatemala." pageId="23" pageNumber="24" stateProvince="Guatemala">Guatemala.</location>
214
<date id="FFDD1024FFE9FFD6FCF940B5BCBB38E2" box="[849,969,702,728]" pageId="23" pageNumber="24" value="1984-08">
<collectingDate id="EF99E9CCFFE9FFD6FCF940B5BCBB38E2" box="[849,969,702,728]" pageId="23" pageNumber="24" value="1984-08">VIII 1984</collectingDate>
</date>
.
<collectorName id="26965332FFE9FFD6FC7D40B5BB4438E2" box="[981,1078,702,728]" pageId="23" pageNumber="24">J. Perez</collectorName>
”; CNMN
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE9FFD6FB1640B5BC5A38C1" ID-GBIF-Occurrence="3093980455" collectingDate="2000-11-22" collectionCode="UVGC" collectorName="M. L. Muller" country="Guatemala" location="Kanajuyu" municipality="Sta. Rosa" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Guatemala" typeStatus="paratype">
<specimenCount id="9D65FD6DFFE9FFD6FB1640B5BB9E38E2" box="[1214,1260,702,728]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
; “
<collectingMunicipality id="6BB8AC9EFFE9FFD6FAAE40B5BA0538E2" box="[1286,1399,702,728]" pageId="23" pageNumber="24">Sta. Rosa</collectingMunicipality>
z.16.
<location id="8EBC603FFFE9FFD6FF5640EABE0238C1" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FF5640EABE0238C1" box="[254,368,737,763]" country="Guatemala" municipality="Sta. Rosa" name="Kanajuyu" pageId="23" pageNumber="24" stateProvince="Guatemala">Kanajuyú</location>
2.
<date id="FFDD1024FFE9FFD6FE3D40EABD6D38C1" box="[405,543,737,763]" pageId="23" pageNumber="24" value="2000-11-22">
<collectingDate id="EF99E9CCFFE9FFD6FE3D40EABD6D38C1" box="[405,543,737,763]" pageId="23" pageNumber="24" value="2000-11-22">22.XI.2000</collectingDate>
</date>
.
<collectorName id="26965332FFE9FFD6FD8240EABDC838C1" box="[554,698,737,763]" pageId="23" pageNumber="24">M.L.Muller</collectorName>
”;
<collectionCode id="ED72AE21FFE9FFD6FD7940EABC5A38C1" box="[721,808,737,763]" collectionName="Guatemala, Guatemala City, Universidad del Valle de Guatemala, Collecion de Artropodos" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="23" pageNumber="24">UVGC</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B0B3CB9FFE9FFD6FCE240EABA533925" ID-GBIF-Occurrence="3093980483" collectedFrom="Bosque seco" collectingDate="1998-07-22" collectingMethod="Cano Heces de perro" collectionCode="UVGC" country="Guatemala" location="Nenton" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Huehuetenango" typeStatus="paratype">
<emphasis id="B917EAF6FFE9FFD6FCE240EABB7D38C1" bold="true" box="[842,1039,737,763]" pageId="23" pageNumber="24">
<collectingRegion id="49A7F806FFE9FFD6FCE240EABB7D38C1" box="[842,1039,737,763]" country="Guatemala" name="Huehuetenango" pageId="23" pageNumber="24">Huehuetenango</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFE9FFD6FB8140EABB2A38C6" box="[1065,1112,737,764]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFE9FFD6FBC740EABBBE38C1" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FBC740EABBBE38C1" box="[1135,1228,737,763]" country="Guatemala" name="Nenton" pageId="23" pageNumber="24" stateProvince="Huehuetenango">Nentón</location>
.
<locationDeviation id="9A6E2FC6FFE9FFD6FB7F40EABD693924" pageId="23" pageNumber="24">Camino entre Nentón y San José Chaquial.</locationDeviation>
<date id="FFDD1024FFE9FFD6FD8D410EBDCE3925" box="[549,700,773,799]" pageId="23" pageNumber="24" value="1998-07-22">
<collectingDate id="EF99E9CCFFE9FFD6FD8D410EBDCE3925" box="[549,700,773,799]" pageId="23" pageNumber="24" value="1998-07-22">22 VII 1998</collectingDate>
</date>
.
<collectedFrom id="387F7519FFE9FFD6FD63410EBC173925" box="[715,869,773,799]" pageId="23" pageNumber="24">Bosque seco</collectedFrom>
. E.B.
<collectingMethod id="52224EF3FFE9FFD6FC19410EBBC63925" box="[945,1204,772,799]" pageId="23" pageNumber="24">Cano Heces de perro</collectingMethod>
”;
<collectionCode id="ED72AE21FFE9FFD6FB66410EBA533925" box="[1230,1313,773,799]" collectionName="Guatemala, Guatemala City, Universidad del Valle de Guatemala, Collecion de Artropodos" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="23" pageNumber="24">UVGC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE9FFD6FA97410EBDCA3978" ID-GBIF-Occurrence="3093980531" collectingDate="1993-10-31" collectionCode="UVGC" collectorName="E. Cano" country="Guatemala" location="Chivacabe." pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Huehuetenango" typeStatus="paratype">
<specimenCount id="9D65FD6DFFE9FFD6FA97410EBA033925" box="[1343,1393,773,799]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFE9FFD6FF6F4123BE3C3978" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FF6F4123BE3C3978" box="[199,334,808,834]" country="Guatemala" name="Chivacabe." pageId="23" pageNumber="24" stateProvince="Huehuetenango">Chivacabé.</location>
<date id="FFDD1024FFE9FFD6FEFE4123BEAB3978" box="[342,473,808,834]" pageId="23" pageNumber="24" value="1993-10-31">
<collectingDate id="EF99E9CCFFE9FFD6FEFE4123BEAB3978" box="[342,473,808,834]" pageId="23" pageNumber="24" value="1993-10-31">31 X 1993</collectingDate>
</date>
.
<collectorName id="26965332FFE9FFD6FE4E4123BD3F3978" box="[486,589,808,834]" pageId="23" pageNumber="24">E. Cano</collectorName>
”;
<collectionCode id="ED72AE21FFE9FFD6FDCD4123BDCA3978" box="[613,696,808,834]" collectionName="Guatemala, Guatemala City, Universidad del Valle de Guatemala, Collecion de Artropodos" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="23" pageNumber="24">UVGC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE9FFD6FD7C4123BA563978" ID-GBIF-Occurrence="3093980420" box="[724,1316,808,834]" collectingDate="1993-10-31" collectionCode="VMC" collectorName="E. Cano" country="Guatemala" location="Chivacabe." pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Huehuetenango" typeStatus="paratype">
<specimenCount id="9D65FD6DFFE9FFD6FD7C4123BC703978" box="[724,770,808,834]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFE9FFD6FB494123BA563978" box="[1249,1316,808,834]" pageId="23" pageNumber="24">VMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE9FFD6FAE84123BECD39B3" ID-GBIF-Occurrence="3093980537" collectingDate="2011-09-24" collectionCode="TAMU" collectorName="Col. M. Acevedo &amp; H. Enriquez." country="Guatemala" county="La" location="La libertad" municipality="La Mesilla" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Huehuetenango" typeStatus="paratype">
<specimenCount id="9D65FD6DFFE9FFD6FAE84123BA033978" box="[1344,1393,808,834]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFE9FFD6FF6F4140BE22395F" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FF6F4140BE22395F" box="[199,336,843,869]" country="Guatemala" county="La" municipality="La Mesilla" name="La libertad" pageId="23" pageNumber="24" stateProvince="Huehuetenango">La libertad</location>
,
<collectingMunicipality id="6BB8AC9EFFE9FFD6FEF74147BE91395F" box="[351,483,843,870]" pageId="23" pageNumber="24">La Mesilla</collectingMunicipality>
,
<location id="8EBC603FFFE9FFD6FE5A4140BDC7395F" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FE5A4140BDC7395F" box="[498,693,843,869]" country="Guatemala" county="La" municipality="La Mesilla" name="finca El Bolson." pageId="23" pageNumber="24" stateProvince="Huehuetenango">finca El Bolsón.</location>
<date id="FFDD1024FFE9FFD6FD174140BC3C395F" box="[703,846,843,870]" pageId="23" pageNumber="24" value="2011-09-24">
<collectingDate id="EF99E9CCFFE9FFD6FD174140BC3C395F" box="[703,846,843,870]" pageId="23" pageNumber="24" value="2011-09-24">24 IX 2011</collectingDate>
</date>
. Cafetal. Luz UV + HG.
<collectorName id="26965332FFE9FFD6FB2E4140BA27395F" box="[1158,1365,843,870]" pageId="23" pageNumber="24">Col. M. Acevedo</collectorName>
&amp;
<collectorName id="26965332FFE9FFD6FF154164BE2539B3" box="[189,343,879,905]" pageId="23" pageNumber="24">H. Enríquez.</collectorName>
”;
<collectionCode id="ED72AE21FFE9FFD6FEC34164BECD39B3" box="[363,447,879,905]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="23" pageNumber="24">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE9FFD6FE7E4164BB6E39B3" ID-GBIF-Occurrence="3093980308" box="[470,1052,879,905]" collectingDate="2011-09-24" collectionCode="UVGC" collectorName="Col. M. Acevedo &amp; H. Enriquez." country="Guatemala" county="La" location="El Bolson." municipality="La Mesilla" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Huehuetenango" typeStatus="paratype">
<specimenCount id="9D65FD6DFFE9FFD6FE7E4164BD7339B3" box="[470,513,879,905]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFE9FFD6FC614164BB6E39B3" box="[969,1052,879,905]" collectionName="Guatemala, Guatemala City, Universidad del Valle de Guatemala, Collecion de Artropodos" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="23" pageNumber="24">UVGC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFE9FFD6FB9B4164BEDF3996" ID-GBIF-Occurrence="3093980572" collectingDate="2011-09-24" collectionCode="VMC" collectorName="Col. M. Acevedo &amp; H. Enriquez." country="Guatemala" county="La" location="El Bolson." municipality="La Mesilla" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Huehuetenango" typeStatus="paratype">
<specimenCount id="9D65FD6DFFE9FFD6FB9B4164BB2C39B3" box="[1075,1118,879,905]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFE9FFD6FECE4199BEDF3996" box="[358,429,914,940]" pageId="23" pageNumber="24">VMC</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B0B3CB9FFE9FFD6FE61419ABDA939EA" ID-GBIF-Occurrence="3093980359" collectingDate="1978-10-29" collectionCode="TAMU" collectorName="E. Duarte" country="Guatemala" county="Bosque" location="Bosque pasado" municipality="Comunidad Ruiz" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Sacatepequez" typeStatus="paratype">
<emphasis id="B917EAF6FFE9FFD6FE61419ABD003996" bold="true" box="[457,626,913,940]" pageId="23" pageNumber="24">
<collectingRegion id="49A7F806FFE9FFD6FE61419ABD003996" box="[457,626,913,940]" country="Guatemala" name="Sacatepequez" pageId="23" pageNumber="24">Sacatepéquez</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFE9FFD6FD224199BDC43997" box="[650,694,914,941]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
; “
<location id="8EBC603FFFE9FFD6FD654199BCF13996" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FD654199BCF13996" box="[717,899,914,940]" country="Guatemala" county="Bosque" municipality="Comunidad Ruiz" name="Bosque pasado" pageId="23" pageNumber="24" stateProvince="Sacatepequez">Bosque pasado</location>
.
<collectingMunicipality id="6BB8AC9EFFE9FFD6FC244199BB273996" box="[908,1109,914,940]" pageId="23" pageNumber="24">Comunidad Ruiz</collectingMunicipality>
,
<locationDeviation id="9A6E2FC6FFE9FFD6FBC84199BE2E39EA" pageId="23" pageNumber="24">km35 a Mixco Viejo via San Juan Sac</locationDeviation>
.
<date id="FFDD1024FFE9FFD6FECF41BDBE9939EA" box="[359,491,950,976]" pageId="23" pageNumber="24" value="1978-10-29">
<collectingDate id="EF99E9CCFFE9FFD6FECF41BDBE9939EA" box="[359,491,950,976]" pageId="23" pageNumber="24" value="1978-10-29">29-X-1978</collectingDate>
</date>
.
<collectorName id="26965332FFE9FFD6FE5E41BDBD1839EA" box="[502,618,950,976]" pageId="23" pageNumber="24">E. Duarte</collectorName>
”;
<collectionCode id="ED72AE21FFE9FFD6FD2B41BDBDA939EA" box="[643,731,950,976]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="23" pageNumber="24">TAMU</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B0B3CB9FFE9FFD6FD5341BDBA7239C9" ID-GBIF-Occurrence="3093980523" collectedFrom="Bosque de pino" collectingDate="2001-12-05" collectingMethod="En heces de vaca" collectionCode="UVGC" collectorName="A. Higueros" country="Guatemala" location="cerca de La Marmolera." municipality="San Lorenzo" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Zacapa" typeStatus="paratype">
<emphasis id="B917EAF6FFE9FFD6FD5341BDBC2A39EA" bold="true" box="[763,856,950,976]" pageId="23" pageNumber="24">
<collectingRegion id="49A7F806FFE9FFD6FD5341BDBC2A39EA" box="[763,856,950,976]" country="Guatemala" name="Zacapa" pageId="23" pageNumber="24">Zacapa</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFE9FFD6FCDA41BDBCD339EA" box="[882,929,950,976]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; “
<collectingMunicipality id="6BB8AC9EFFE9FFD6FC1041BDBB2139EA" box="[952,1107,950,976]" pageId="23" pageNumber="24">San Lorenzo</collectingMunicipality>
,
<location id="8EBC603FFFE9FFD6FBF641BDBA0539F5" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FBF641BDBA0539F5" box="[1118,1399,949,976]" country="Guatemala" municipality="San Lorenzo" name="cerca de La Marmolera." pageId="23" pageNumber="24" stateProvince="Zacapa">cerca de La Marmolera.</location>
<date id="FFDD1024FFE9FFD6FF1541D2BE3339C9" box="[189,321,985,1012]" pageId="23" pageNumber="24" value="2001-12-05">
<collectingDate id="EF99E9CCFFE9FFD6FF1541D2BE3339C9" box="[189,321,985,1012]" pageId="23" pageNumber="24" value="2001-12-05">5 XII 2001</collectingDate>
</date>
.
<collectingMethod id="52224EF3FFE9FFD6FEE441D1BD6B39CE" box="[332,537,985,1012]" pageId="23" pageNumber="24">En heces de vaca</collectingMethod>
.
<collectorName id="26965332FFE9FFD6FD8B41D2BDC739C9" box="[547,693,985,1011]" pageId="23" pageNumber="24">A. Higueros</collectorName>
, leg.
<collectedFrom id="387F7519FFE9FFD6FD5B41D1BCDE39C9" box="[755,940,985,1012]" pageId="23" pageNumber="24">Bosque de pino</collectedFrom>
, aprox. 1700msnm”;
<collectionCode id="ED72AE21FFE9FFD6FB0141D2BA7239C9" box="[1193,1280,985,1011]" collectionName="Guatemala, Guatemala City, Universidad del Valle de Guatemala, Collecion de Artropodos" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="23" pageNumber="24">UVGC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF154610BB2A3E0F" blockId="23.[189,1112,1050,1077]" box="[189,1112,1050,1077]" pageId="23" pageNumber="24">
<materialsCitation id="3B0B3CB9FFE9FFD6FF154610BB263E0F" ID-GBIF-Occurrence="3093980346" box="[189,1108,1050,1077]" collectingDate="1962-08" collectionCode="TAMU" collectorName="G. Halffter" country="Mexico" location="Santa Rosa" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas">
<collectingCountry id="F3747674FFE9FFD6FF154610BE433E0F" box="[189,305,1051,1077]" name="Mexico" pageId="23" pageNumber="24">MEXICO</collectingCountry>
–
<emphasis id="B917EAF6FFE9FFD6FEE64611BEC73E0E" bold="true" box="[334,437,1050,1076]" pageId="23" pageNumber="24">
<collectingRegion id="49A7F806FFE9FFD6FEE64611BEC73E0E" box="[334,437,1050,1076]" country="Mexico" name="Chiapas" pageId="23" pageNumber="24">Chiapas</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFE9FFD6FE664610BE8F3E0F" box="[462,509,1051,1077]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFE9FFD6FDBB4610BDEF3E0F" LSID="urn:lsid:plazi:treatment:03CA87F2FFE8FFD8FDDA448BBDA9398B:8EBC603FFFE9FFD6FDBB4610BDEF3E0F" box="[531,669,1051,1077]" country="Mexico" name="Santa Rosa" pageId="23" pageNumber="24" stateProvince="Chiapas">Santa Rosa</location>
,
<date id="FFDD1024FFE9FFD6FD0F4610BC503E0F" box="[679,802,1051,1077]" pageId="23" pageNumber="24" value="1962-08">
<collectingDate id="EF99E9CCFFE9FFD6FD0F4610BC503E0F" box="[679,802,1051,1077]" pageId="23" pageNumber="24" value="1962-08">VIII-1962</collectingDate>
</date>
.
<collectorName id="26965332FFE9FFD6FC854610BCDF3E0F" box="[813,941,1051,1077]" pageId="23" pageNumber="24">G. Halffter</collectorName>
leg.”;
<collectionCode id="ED72AE21FFE9FFD6FC544610BB263E0F" box="[1020,1108,1051,1077]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="23" pageNumber="24">TAMU</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF154657BE2F3E4C" blockId="23.[189,349,1116,1142]" box="[189,349,1116,1142]" pageId="23" pageNumber="24">
<heading id="D0948188FFE9FFD6FF154657BE2F3E4C" bold="true" box="[189,349,1116,1142]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B917EAF6FFE9FFD6FF154657BE2F3E4C" bold="true" box="[189,349,1116,1142]" pageId="23" pageNumber="24">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF154680BEA03E9F" blockId="23.[189,466,1163,1189]" box="[189,466,1163,1189]" pageId="23" pageNumber="24">
<collectingCountry id="F3747674FFE9FFD6FF154680BE4D3E9F" box="[189,319,1163,1189]" name="Guatemala" pageId="23" pageNumber="24">Guatemala</collectingCountry>
,
<collectingCountry id="F3747674FFE9FFD6FEE34680BEBC3E9F" box="[331,462,1163,1189]" name="Guatemala" pageId="23" pageNumber="24">Guatemala</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFE9FFD8FF1546C7BDFF38BC" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="description">
<paragraph id="8BDC36E4FFE9FFD6FF1546C7BE3C3EDC" blockId="23.[189,334,1228,1254]" box="[189,334,1228,1254]" pageId="23" pageNumber="24">
<heading id="D0948188FFE9FFD6FF1546C7BE3C3EDC" bold="true" box="[189,334,1228,1254]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B917EAF6FFE9FFD6FF1546C7BE3C3EDC" bold="true" box="[189,334,1228,1254]" pageId="23" pageNumber="24">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF4246F0BE8D3F2C" blockId="23.[189,511,1275,1337]" box="[234,511,1275,1302]" pageId="23" pageNumber="24">
<emphasis id="B917EAF6FFE9FFD6FF4246F0BE0F3F2C" bold="true" box="[234,381,1275,1302]" pageId="23" pageNumber="24">Major male</emphasis>
(
<typeStatus id="54D88846FFE9FFD6FE2546F0BE853F2C" box="[397,503,1275,1302]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF154714BEF53F03" blockId="23.[189,511,1275,1337]" box="[189,391,1311,1337]" pageId="23" pageNumber="24">
Length
<quantity id="4C9B9B01FFE9FFD6FEBF4714BEF13F03" box="[279,387,1311,1337]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" pageId="23" pageNumber="24" unit="mm" value="17.1">17.1 mm</quantity>
.
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF15476ABE903FDC" blockId="23.[189,1398,1376,1510]" pageId="23" pageNumber="24">
HEAD. Clypeus bidentate, black on anterior margin, bright metallic green on posterior portion, with roughened sculpture. Genae bright metallic green, with roughened sculpture. Front black, with dark metallic blue-green on portions adjacent to cephalic horn. Cephalic horn black, curved posteriorly over pronotum (
<figureCitation id="13582A61FFE9FFD6FE9747C0BED23FDF" box="[319,416,1483,1509]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="23" pageNumber="24">Figs 2D</figureCitation>
,
<figureCitation id="13582A61FFE9FFD6FE0547C0BEA73FDC" box="[429,469,1483,1510]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="23" pageNumber="24">6A</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF154406BE1A3CC6" blockId="23.[189,1399,1548,1788]" pageId="23" pageNumber="24">
PRONOTUM. Uniformly bright metallic green, becoming black on lateral margins of posterolateral angles. Keel absent in the middle of anterior pronotal margin. Disc triangular, flat, with two distinctly developed tubercles on anterior portion. Triangle with lightly granulate, scabriculous, impunctate. Sides scabriculous, with smooth sculpture, superficially impressed punctures. Lateral lines of pronotal triangle straight. Posterolateral angles sharply acute, elongate, projected posterolaterally. Lateral fossae distinctly impressed. Basal fossae obtusely oval, deeply impressed. Posterior margin distinctly punctate (
<figureCitation id="13582A61FFE9FFD6FF6D44EABE553CC1" box="[197,295,1761,1787]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="23" pageNumber="24">Figs 2D</figureCitation>
,
<figureCitation id="13582A61FFE9FFD6FE9B44EABE293CC6" box="[307,347,1761,1788]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="23" pageNumber="24">6A</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF154528BD0F3D92" blockId="23.[189,1401,1826,1960]" pageId="23" pageNumber="24">
ELYTRA. Thick, roughened, bright metallic green striae, scabriculous, impressed basally as distinct fossae, with distinctly impressed punctation. Interstriae black, partially roughened, scabriculous, with almost effaced punctation; except for almost completely rough, bright metallic green first stria. Sutural margin without apical tooth (
<figureCitation id="13582A61FFE9FFD6FDBA4586BD033D92" box="[530,625,1933,1960]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="23" pageNumber="24">Fig. 6A</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFE9FFD6FF1545C4BDCA3DD2" blockId="23.[189,696,1998,2025]" box="[189,696,1998,2025]" pageId="23" pageNumber="24">PROTIBIAE. Quadridentate with apical spine.</paragraph>
<paragraph id="8BDC36E4FFE6FFD9FF154302BD573B7D" blockId="24.[189,1398,265,327]" pageId="24" pageNumber="25">TERGITE VIII. Bright metallic green; scabriculous; with rough, distinctly impressed punctures. Basal margin with thick, small setae.</paragraph>
<paragraph id="8BDC36E4FFE6FFD9FF154365BCFE3BF4" blockId="24.[186,1398,366,462]" pageId="24" pageNumber="25">
GENITALIA. Right lobe and left lobe of endophallite copulatrix similar in size. Right lobe obtusely triangular in shape, with apical portion projected posteriorly. Left lobe bent posteriorly, convex superiorly, lobed inferiorly. Central ridge and column similar in size (
<figureCitation id="13582A61FFE6FFD9FC8843BFBC0C3BF4" box="[800,894,436,462]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma &amp; Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="24" pageNumber="25">Fig. 1H</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFE6FFD9FF4243FEBE0F382A" blockId="24.[187,1399,501,599]" box="[234,381,501,528]" pageId="24" pageNumber="25">
<emphasis id="B917EAF6FFE6FFD9FF4243FEBE0F382A" bold="true" box="[234,381,501,528]" pageId="24" pageNumber="25">Minor male</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFE6FFD9FF154012BC17386D" blockId="24.[187,1399,501,599]" pageId="24" pageNumber="25">Like the major male, except for the reduction of secondary sexual characters (i.e., cephalic horn, pronotal triangle and tubercles, and pronotal posterolateral angles).</paragraph>
<caption id="DF1C666CFFE6FFD9FF1545C5BB5D3DD2" ID-DOI="http://doi.org/10.5281/zenodo.4836768" ID-Zenodo-Dep="4836768" box="[189,1071,1998,2025]" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,232,1998,2024]" targetBox="[217,1394,753,1972]" targetPageId="24">
<paragraph id="8BDC36E4FFE6FFD9FF1545C5BB5D3DD2" blockId="24.[189,1071,1998,2025]" box="[189,1071,1998,2025]" pageId="24" pageNumber="25">
<emphasis id="B917EAF6FFE6FFD9FF1545C5BE783DD2" bold="true" box="[189,266,1998,2024]" pageId="24" pageNumber="25">Fig. 6.</emphasis>
<taxonomicName id="4C634D67FFE6FFD9FEBA45C5BE8B3DD2" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[274,505,1998,2024]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="jackenioi" status="sp. nov.">
<emphasis id="B917EAF6FFE6FFD9FEBA45C5BE8B3DD2" box="[274,505,1998,2024]" italics="true" pageId="24" pageNumber="25">Phanaeus jackenioi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224578DFFE6FFD9FDA845C4BD283DD3" box="[512,602,1999,2025]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B917EAF6FFE6FFD9FDF745C5BD063DD2" bold="true" box="[607,628,1998,2024]" pageId="24" pageNumber="25">A</emphasis>
. Holotype, ♂ (UVGC).
<emphasis id="B917EAF6FFE6FFD9FC3A45C5BCD43DD2" bold="true" box="[914,934,1998,2024]" pageId="24" pageNumber="25">B</emphasis>
. ♀ (VMC).
</paragraph>
</caption>
<paragraph id="8BDC36E4FFE7FFD8FF424302BE373B19" blockId="25.[187,1399,265,544]" box="[234,325,265,291]" pageId="25" pageNumber="26">
<emphasis id="B917EAF6FFE7FFD8FF424302BE373B19" bold="true" box="[234,325,265,291]" pageId="25" pageNumber="26">Female</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF154327BD6A381A" blockId="25.[187,1399,265,544]" pageId="25" pageNumber="26">
Similar to the male, except for the head showing a cephalic trituberculate carina; carinate middle tubercle, less raised, slightly more frontally projected than lateral tubercles; lateral tubercles obtusely conical; frons with distinctly impressed punctures; pronotal sculpture completely smooth, with superficially impressed punctures; pronotum bright metallic green, with shining black, variable in size area on central portion of disc; pronotal process trituberculate, lacking concavity rounded pronotal tubercles, nearly aligned, with middle tubercle more developed than lateral tubercles; posterior pronotal midline completely effaced (
<figureCitation id="13582A61FFE7FFD8FE06400EBD78381A" box="[430,522,517,544]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="25" pageNumber="26">Fig. 6B</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF42404CBE12385B" blockId="25.[189,653,583,646]" box="[234,352,583,609]" pageId="25" pageNumber="26">
<emphasis id="B917EAF6FFE7FFD8FF42404CBE12385B" bold="true" box="[234,352,583,609]" pageId="25" pageNumber="26">Variation</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF154067BDFF38BC" blockId="25.[189,653,583,646]" box="[189,653,619,646]" pageId="25" pageNumber="26">
Mean length
<quantity id="4C9B9B01FFE7FFD8FEF04060BEB238BC" box="[344,448,619,646]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="25" pageNumber="26" unit="mm" value="17.3">17.3 mm</quantity>
(
<quantity id="4C9B9B01FFE7FFD8FE784060BDF338BC" box="[464,641,619,646]" metricMagnitude="-2" metricUnit="m" metricValue="1.7299999999999998" metricValueMax="1.8399999999999999" metricValueMin="1.6199999999999999" pageId="25" pageNumber="26" unit="mm" value="17.299999999999997" valueMax="18.4" valueMin="16.2">16.2–18.4 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFE7FFD8FF1540A6BA0438C2" box="[189,1398,685,760]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BDC36E4FFE7FFD8FF1540A6BE2438FD" blockId="25.[189,342,685,711]" box="[189,342,685,711]" pageId="25" pageNumber="26">
<heading id="D0948188FFE7FFD8FF1540A6BE2438FD" bold="true" box="[189,342,685,711]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B917EAF6FFE7FFD8FF1540A6BE2438FD" bold="true" box="[189,342,685,711]" pageId="25" pageNumber="26">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF1540D6BA0438C2" blockId="25.[189,1398,733,760]" box="[189,1398,733,760]" pageId="25" pageNumber="26">
Inner slope of the Sierra Madre de Chiapas, south-central
<collectingCountry id="F3747674FFE7FFD8FCF040D6BCA538CD" box="[856,983,733,759]" name="Guatemala" pageId="25" pageNumber="26">Guatemala</collectingCountry>
and southeastern Chiapas (
<figureCitation id="13582A61FFE7FFD8FABB40D6BA1A38C2" box="[1299,1384,733,760]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017;P. edmondsi Moctezuma, Deloya &amp; Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="25" pageNumber="26">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFE7FFD8FF154114BDA9398B" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8BDC36E4FFE7FFD8FF154114BE5D3903" blockId="25.[189,303,799,825]" box="[189,303,799,825]" pageId="25" pageNumber="26">
<heading id="D0948188FFE7FFD8FF154114BE5D3903" bold="true" box="[189,303,799,825]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B917EAF6FFE7FFD8FF154114BE5D3903" bold="true" box="[189,303,799,825]" pageId="25" pageNumber="26">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFE7FFD8FF154145BDA9398B" blockId="25.[188,1399,846,945]" pageId="25" pageNumber="26">
The type series of
<taxonomicName id="4C634D67FFE7FFD8FE394144BD6B3952" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[401,537,846,872]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="jackenioi" status="sp. nov.">
<emphasis id="B917EAF6FFE7FFD8FE394144BD6B3952" box="[401,537,846,872]" italics="true" pageId="25" pageNumber="26">P. jackenioi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224578DFFE7FFD8FD884144BD0B3953" box="[544,633,847,873]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
was originally labelled as
<taxonomicName id="4C634D67FFE7FFD8FC054144BB633952" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[941,1041,847,872]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFE7FFD8FC054144BB633952" box="[941,1041,847,872]" italics="true" pageId="25" pageNumber="26">P. pyrois</emphasis>
</taxonomicName>
. The new species was referred to as
<taxonomicName id="4C634D67FFE7FFD8FEA84178BEE239B6" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[256,400,882,908]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="endymion">
<emphasis id="B917EAF6FFE7FFD8FEA84178BEE239B6" box="[256,400,882,908]" italics="true" pageId="25" pageNumber="26">P. endymion</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF24B15FFE7FFD8FE684179BDFA39B7" author="Edmonds W. D." box="[448,648,882,909]" pageId="25" pageNumber="26" pagination="1 - 105" refId="ref37487" refString="Edmonds W. D. 1994. Revision of Phanaeus Macleay, a New World genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Contributions in Science, Natural History Museum of Los Angeles County 443: 1 - 105." type="journal article" year="1994">Edmonds (1994)</bibRefCitation>
,
<bibRefCitation id="EFF24B15FFE7FFD8FD3E4179BCB939B7" author="Edmonds W. D. &amp; Zidek J." box="[662,971,882,909]" pageId="25" pageNumber="26" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. &amp; Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds &amp; Zídek (2012)</bibRefCitation>
,
<bibRefCitation id="EFF24B15FFE7FFD8FC714179BBCD39B7" author="Lizardo V. &amp; Escobar F. &amp; Rojas-Soto O." box="[985,1215,882,909]" pageId="25" pageNumber="26" pagination="271 - 294" refId="ref39838" refString="Lizardo V., Escobar F. &amp; Rojas-Soto O. 2017. Diversity and distribution of Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) in Mexico. Zootaxa 4358 (3): 271 - 294. https: // doi. org / 10.11646 / zootaxa. 4358.2.3" type="journal article" year="2017">
Lizardo
<emphasis id="B917EAF6FFE7FFD8FB964178BB0339B6" box="[1086,1137,882,908]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. 2017
</bibRefCitation>
,
<bibRefCitation id="EFF24B15FFE7FFD8FB644178BE02398B" author="Moctezuma V. &amp; Halffter G." pageId="25" pageNumber="26" pagination="47 - 56" refId="ref40548" refString="Moctezuma V. &amp; Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47" type="journal article" year="2017">Moctezuma &amp; Halffter (2017)</bibRefCitation>
and
<bibRefCitation id="EFF24B15FFE7FFD8FE01419CBDAA398B" author="GBIF Secretariat" box="[425,728,918,945]" pageId="25" pageNumber="26" refId="ref38078" refString="GBIF Secretariat. 2019 a. Phanaeus endymion Harold, 1863. GBIF Backbone Taxonomy. Checklist dataset. Available from https: // doi. org / 10.15468 / 39 omei [accessed Jul. 2020]." type="book" year="2019">GBIF Secretariat (2019a)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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