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<document id="25DCEBEE5BC5BE40F1DFAC661A1F2F71" ID-CLB-Dataset="7153" ID-DOI="10.5852/ejt.2021.747.1333" ID-GBIF-Dataset="aca67cc8-96ca-4f24-b8af-2b532ecd355c" ID-ISSN="2118-9773" ID-Zenodo-Dep="4836744" ID-ZooBank="3F0B6EAF-C616-4865-811A-414A094B590C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619605517524" checkinUser="plazi" docAuthor="Moctezuma, Victor &amp; Halffter, Gonzalo" docDate="2021" docId="03CA87F2FFDDFFE7FDC94688BD063B71" docLanguage="en" docName="ejt-747_1-71.pdf" docOrigin="European Journal of Taxonomy 747" docSource="http://zoobank.org/3f0b6eaf-c616-4865-811a-414a094b590c" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phanaeus panamensis Moctezuma &amp; Halffter 2021, sp. nov." docType="treatment" docUuid="C10343D8-4C59-4F57-987F-4234062447E3" docUuidSource="ZooBank" docVersion="11" lastPageNumber="39" masterDocId="FFF3FF8AFFFEFFC1FFA8420BBF723A3A" masterDocTitle="Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species" masterLastPageNumber="71" masterPageNumber="1" pageNumber="36" updateTime="1698937721231" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2028BE86A2DAA954B33C472836A44658">Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species</mods:title>
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<mods:namePart id="FB1F6D90C8D0384EF0BA4F9B5F992379">Moctezuma, Victor</mods:namePart>
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<mods:affiliation id="21801CC0284D44633C6187CE805FE9D7">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:namePart id="0CEF3EAE5A15DF8D949CFF05CEB8EF07">Halffter, Gonzalo</mods:namePart>
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<mods:affiliation id="36FF0F97CA653ED37C6444A520F65497">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:date id="E6437A15FBD4B7C5975F15A7D710084D">2021</mods:date>
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<treatment id="03CA87F2FFDDFFE7FDC94688BD063B71" ID-DOI="http://doi.org/10.5281/zenodo.4726235" ID-GBIF-Taxon="181753774" ID-Zenodo-Dep="4726235" LSID="urn:lsid:zoobank.org:act:C10343D8-4C59-4F57-987F-4234062447E3" httpUri="http://treatment.plazi.org/id/03CA87F2FFDDFFE7FDC94688BD063B71" lastPageId="38" lastPageNumber="39" pageId="35" pageNumber="36">
<subSubSection id="C379656FFFDDFFE2FDC94688BCA13EA4" box="[609,979,1155,1182]" pageId="35" pageNumber="36" type="nomenclature">
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<heading id="D0948188FFDDFFE2FDC94688BCA13EA4" box="[609,979,1155,1182]" centered="true" fontSize="11" level="2" pageId="35" pageNumber="36" reason="2">
<taxonomicName id="4C634D67FFDDFFE2FDC94688BC023EA7" ID-CoL="874GT" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[609,880,1155,1182]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B917EAF6FFDDFFE2FDC94688BC023EA7" bold="true" box="[609,880,1155,1182]" italics="true" pageId="35" pageNumber="36">Phanaeus panamensis</emphasis>
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<taxonomicNameLabel id="A224578DFFDDFFE2FCD0468FBCA13EA4" box="[888,979,1156,1182]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C379656FFFDDFFE2FED546ACBCA13EDC" pageId="35" pageNumber="36" type="description">
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urn:lsid:zoobank.org:act:
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<paragraph id="8BDC36E4FFDDFFE2FDC946C7BCA13EDC" blockId="35.[381,1206,1155,1254]" box="[609,979,1227,1254]" pageId="35" pageNumber="36">
<figureCitation id="13582A61FFDDFFE2FDC946C7BDCF3EDC" box="[609,701,1228,1254]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma &amp; Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="35" pageNumber="36">Figs 1L</figureCitation>
,
<figureCitation id="13582A61FFDDFFE2FD6246C7BD823EDC" box="[714,752,1228,1254]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="35" pageNumber="36">2H</figureCitation>
,
<figureCitation id="13582A61FFDDFFE2FD5546C7BC6F3EDC" box="[765,797,1228,1254]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="35" pageNumber="36">10</figureCitation>
,
<figureCitation id="13582A61FFDDFFE2FC8146C0BC3B3EDF" box="[809,841,1227,1253]" captionStart="Fig" captionStartId="51.[189,232,1233,1259]" captionTargetBox="[193,1395,270,1197]" captionTargetId="figure-241@51.[189,1398,265,1199]" captionTargetPageId="51" captionText="Fig. 16. Distribution of the Phanaeus endymion species group in Central America (in part)." figureDoi="http://doi.org/10.5281/zenodo.4836798" httpUri="https://zenodo.org/record/4836798/files/figure.png" pageId="35" pageNumber="36">16</figureCitation>
,
<figureCitation id="13582A61FFDDFFE2FCFD46C7BCFC3EDC" box="[853,910,1228,1254]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma &amp; Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="35" pageNumber="36">18M</figureCitation>
,
<figureCitation id="13582A61FFDDFFE2FC3346C7BCA13EDC" box="[923,979,1228,1254]" captionStart="Fig" captionStartId="56.[189,231,1678,1704]" captionTargetBox="[306,1282,265,1646]" captionTargetId="figure-158@56.[306,1282,265,1646]" captionTargetPageId="56" captionText="Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma &amp; Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836804" httpUri="https://zenodo.org/record/4836804/files/figure.png" pageId="35" pageNumber="36">19M</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFDDFFE2FF154705BD2B3C0B" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="8BDC36E4FFDDFFE2FF154705BE443F12" blockId="35.[189,310,1294,1320]" box="[189,310,1294,1320]" pageId="35" pageNumber="36">
<heading id="D0948188FFDDFFE2FF154705BE443F12" bold="true" box="[189,310,1294,1320]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B917EAF6FFDDFFE2FF154705BE443F12" bold="true" box="[189,310,1294,1320]" pageId="35" pageNumber="36">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFDDFFE2FF154735BD2B3C0B" blockId="35.[189,1399,1342,1585]" pageId="35" pageNumber="36">
The new species is easily diagnosed within the
<taxonomicName id="4C634D67FFDDFFE2FD4F4735BC043F62" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[743,886,1342,1368]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="endymion">
<emphasis id="B917EAF6FFDDFFE2FD4F4735BC043F62" box="[743,886,1342,1368]" italics="true" pageId="35" pageNumber="36">P. endymion</emphasis>
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species group by the dull black colour with bright metallic red-green sheen dorsally (
<figureCitation id="13582A61FFDDFFE2FD084769BC703F46" box="[672,770,1378,1404]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="35" pageNumber="36">Figs 2H</figureCitation>
,
<figureCitation id="13582A61FFDDFFE2FCA74769BC5D3F46" box="[783,815,1378,1404]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="35" pageNumber="36">10</figureCitation>
); and elytral striae not strongly impressed basally as distinct fossae (
<figureCitation id="13582A61FFDDFFE2FE36478DBE8B3F9A" box="[414,505,1414,1440]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="35" pageNumber="36">Fig. 10</figureCitation>
).
<taxonomicName id="4C634D67FFDDFFE2FDB9478DBC683F9A" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[529,794,1414,1440]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B917EAF6FFDDFFE2FDB9478DBC683F9A" box="[529,794,1414,1440]" italics="true" pageId="35" pageNumber="36">Phanaeus panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224578DFFDDFFE2FC8D478CBCF13F9B" box="[805,899,1415,1441]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished from
<taxonomicName id="4C634D67FFDDFFE2FB3E478CBA5A3F9A" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[1174,1320,1414,1440]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B917EAF6FFDDFFE2FB3E478CBA5A3F9A" box="[1174,1320,1414,1440]" italics="true" pageId="35" pageNumber="36">P. olsoufieffi</emphasis>
</taxonomicName>
by its smaller body size (rarely attaining
<quantity id="4C9B9B01FFDDFFE2FDFF47A1BDA53FFF" box="[599,727,1450,1477]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="35" pageNumber="36" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">20–21 mm</quantity>
in length); and the major males with obsolete keel in the middle of anterior pronotal margin (
<figureCitation id="13582A61FFDDFFE2FDDC47C5BDA43FD3" box="[628,726,1486,1513]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="35" pageNumber="36">Fig. 2H</figureCitation>
); and posterolateral angles weakly developed, widened, slightly projected posteriorly (
<figureCitation id="13582A61FFDDFFE2FD8247F8BDE93C37" box="[554,667,1523,1549]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="35" pageNumber="36">Fig. 10A</figureCitation>
). Furthermore, the pronotal disc in
<taxonomicName id="4C634D67FFDDFFE2FBE647FFBB923C37" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[1102,1248,1523,1549]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B917EAF6FFDDFFE2FBE647FFBB923C37" box="[1102,1248,1523,1549]" italics="true" pageId="35" pageNumber="36">P. olsoufieffi</emphasis>
</taxonomicName>
is distinctly darker and smoother (
<figureCitation id="13582A61FFDDFFE2FE16441CBD6A3C0B" box="[446,536,1559,1585]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="35" pageNumber="36">Figs 2F</figureCitation>
,
<figureCitation id="13582A61FFDDFFE2FD8C441CBD3E3C0B" box="[548,588,1559,1585]" captionStart="Fig" captionStartId="31.[189,232,1962,1988]" captionTargetBox="[199,1366,462,1913]" captionTargetId="figure-52@31.[189,1399,462,1925]" captionTargetPageId="31" captionText="Fig. 8. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Lectotype ♂ (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836778" httpUri="https://zenodo.org/record/4836778/files/figure.png" pageId="35" pageNumber="36">8A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFDDFFE2FF154451BBB03C99" box="[189,1218,1626,1700]" pageId="35" pageNumber="36" type="etymology">
<paragraph id="8BDC36E4FFDDFFE2FF154451BE363C4E" blockId="35.[189,324,1626,1652]" box="[189,324,1626,1652]" pageId="35" pageNumber="36">
<heading id="D0948188FFDDFFE2FF154451BE363C4E" bold="true" box="[189,324,1626,1652]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B917EAF6FFDDFFE2FF154451BE363C4E" bold="true" box="[189,324,1626,1652]" pageId="35" pageNumber="36">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFDDFFE2FF154482BBB03C99" blockId="35.[189,1218,1673,1700]" box="[189,1218,1673,1700]" pageId="35" pageNumber="36">
The specific epithet refers to
<collectingCountry id="F3747674FFDDFFE2FDBA4481BD003C9E" box="[530,626,1674,1700]" name="Panama" pageId="35" pageNumber="36">Panama</collectingCountry>
, where a majority of the
<typeStatus id="54D88846FFDDFFE2FC3F4481BCB83C9E" box="[919,970,1674,1700]" pageId="35" pageNumber="36">type</typeStatus>
series was collected.
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFDDFFE5FF1544C7BD273987" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BDC36E4FFDDFFE2FF1544C7BD453CDC" blockId="35.[189,567,1739,1766]" box="[189,567,1739,1766]" pageId="35" pageNumber="36">
<emphasis id="B917EAF6FFDDFFE2FF1544C7BE1E3CDF" bold="true" box="[189,364,1739,1766]" pageId="35" pageNumber="36">Type material</emphasis>
(
<specimenCount id="9D65FD6DFFDDFFE2FED444C7BEBD3CDC" box="[380,463,1740,1766]" pageId="35" pageNumber="36" type="male">17 ♂♂</specimenCount>
,
<specimenCount id="9D65FD6DFFDDFFE2FE7444C7BD593CDC" box="[476,555,1740,1766]" pageId="35" pageNumber="36" type="female">14 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BDC36E4FFDDFFE2FF4244F0BE2E3D2F" blockId="35.[189,1399,1787,1886]" box="[234,348,1787,1813]" pageId="35" pageNumber="36">
<emphasis id="B917EAF6FFDDFFE2FF4244F0BE2E3D2F" bold="true" box="[234,348,1787,1813]" pageId="35" pageNumber="36">
<typeStatus id="54D88846FFDDFFE2FF4244F0BE2E3D2F" box="[234,348,1787,1813]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFDDFFE2FF154514BD3B3D64" blockId="35.[189,1399,1787,1886]" pageId="35" pageNumber="36">
<materialsCitation id="3B0B3CB9FFDDFFE2FF154514BD3B3D64" ID-GBIF-Occurrence="3093980596" collectingDate="2019-09" collectingMethod="Manure trap" collectionCode="TAMU" country="Panama" elevation="850" latitude="8.666667" location="Cerro Campana" longLatPrecision="1300" longitude="-79.933334" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Panama" typeStatus="holotype">
<collectingCountry id="F3747674FFDDFFE2FF154514BE4B3D03" box="[189,313,1823,1849]" name="Panama" pageId="35" pageNumber="36">PANAMA</collectingCountry>
•
<specimenCount id="9D65FD6DFFDDFFE2FEF1452BBE003D00" box="[345,370,1824,1850]" pageId="35" pageNumber="36" type="male">♂</specimenCount>
;
<collectingRegion id="49A7F806FFDDFFE2FE2A4514BE903D03" box="[386,482,1823,1849]" country="Panama" name="Panama" pageId="35" pageNumber="36">Panamá</collectingRegion>
,
<location id="8EBC603FFFDDFFE2FE594514BDC13D03" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDDFFE2FE594514BDC13D03" box="[497,691,1823,1849]" country="Panama" latitude="8.666667" longLatPrecision="1300" longitude="-79.933334" name="Cerro Campana" pageId="35" pageNumber="36" stateProvince="Panama">Cerro Campana</location>
; “
<location id="8EBC603FFFDDFFE2FD644514BCFD3D03" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDDFFE2FD644514BCFD3D03" box="[716,911,1823,1849]" country="Panama" latitude="8.666667" longLatPrecision="1300" longitude="-79.933334" name="Cerro Campana" pageId="35" pageNumber="36" stateProvince="Panama">Cerro Campana</location>
,
<quantity id="4C9B9B01FFDDFFE2FC364514BC973D00" box="[926,997,1823,1850]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="35" pageNumber="36" unit="m" value="850.0">
<elevation id="004ED1D7FFDDFFE2FC364514BC973D00" box="[926,997,1823,1850]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="35" pageNumber="36" unit="m" value="850.0">850m</elevation>
</quantity>
.
<geoCoordinate id="EE575023FFDDFFE2FC5C4514BB3F3D03" box="[1012,1101,1823,1849]" degrees="8" direction="north" minutes="40" orientation="latitude" pageId="35" pageNumber="36" precision="925" value="8.666667">8°40’N</geoCoordinate>
,
<geoCoordinate id="EE575023FFDDFFE2FBF54514BBBB3D00" box="[1117,1225,1823,1850]" degrees="79" direction="west" minutes="56" orientation="longitude" pageId="35" pageNumber="36" precision="925" value="-79.933334">79°56’W</geoCoordinate>
.
<date id="FFDD1024FFDDFFE2FB714514BA4F3D03" box="[1241,1341,1823,1849]" pageId="35" pageNumber="36" value="2019-09">
<collectingDate id="EF99E9CCFFDDFFE2FB714514BA4F3D03" box="[1241,1341,1823,1849]" pageId="35" pageNumber="36" value="2019-09">19 Sept.</collectingDate>
</date>
’71. Stockwell.
<collectingMethod id="52224EF3FFDDFFE2FEE9454FBEA63D64" box="[321,468,1860,1886]" pageId="35" pageNumber="36">Manure trap</collectingMethod>
”;
<collectionCode id="ED72AE21FFDDFFE2FE45454FBD373D64" box="[493,581,1860,1886]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="35" pageNumber="36">TAMU</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BDC36E4FFDDFFE2FF42458DBE153D9A" blockId="35.[189,1398,1926,2025]" box="[234,359,1926,1952]" pageId="35" pageNumber="36">
<emphasis id="B917EAF6FFDDFFE2FF42458DBE153D9A" bold="true" box="[234,359,1926,1952]" pageId="35" pageNumber="36">
<typeStatus id="54D88846FFDDFFE2FF42458DBE153D9A" box="[234,359,1926,1952]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFDDFFE2FF1545A1BBFB3DD2" blockId="35.[189,1398,1926,2025]" pageId="35" pageNumber="36">
<materialsCitation id="3B0B3CB9FFDDFFE2FF1545A1BD553DD2" ID-GBIF-Occurrence="3093980411" collectingDate="1969-09-12" collectionCode="GHC" collectorName="G. Halffter &amp; P. Reyes" country="Costa Rica" latitude="10.466666" location="Rio Puerto Viejo" longLatPrecision="0" longitude="-84.0" municipality="Estacion La Selva" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="paratype">
<collectingCountry id="F3747674FFDDFFE2FF1545A1BE153DFE" box="[189,359,1962,1988]" name="Costa Rica" pageId="35" pageNumber="36">COSTA RICA</collectingCountry>
–
<emphasis id="B917EAF6FFDDFFE2FE2B45A1BE953DFE" bold="true" box="[387,487,1962,1988]" pageId="35" pageNumber="36">
<collectingRegion id="49A7F806FFDDFFE2FE2B45A1BE953DFE" box="[387,487,1962,1988]" country="Costa Rica" name="Heredia" pageId="35" pageNumber="36">Heredia</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFDDFFE2FDA845A1BD423DFF" box="[512,560,1962,1989]" pageId="35" pageNumber="36" type="male">1 ♂</specimenCount>
; “
<collectingMunicipality id="6BB8AC9EFFDDFFE2FDEE45A1BC523DFE" box="[582,800,1962,1989]" pageId="35" pageNumber="36">Estación La Selva</collectingMunicipality>
,
<location id="8EBC603FFFDDFFE2FC8445A1BC863DFE" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDDFFE2FC8445A1BC863DFE" box="[812,1012,1962,1989]" country="Costa Rica" latitude="10.466666" longLatPrecision="0" longitude="-84.0" municipality="Estacion La Selva" name="Rio Puerto Viejo" pageId="35" pageNumber="36" stateProvince="Heredia">Río Puerto Viejo</location>
,
<geoCoordinate id="EE575023FFDDFFE2FBA845A1BB3B3DFE" box="[1024,1097,1962,1988]" degrees="84" direction="west" orientation="longitude" pageId="35" pageNumber="36" precision="55555" value="-84.0">84°W</geoCoordinate>
/
<geoCoordinate id="EE575023FFDDFFE2FBFA45A1BBC43DFE" box="[1106,1206,1962,1988]" degrees="10" direction="north" minutes="28" orientation="latitude" pageId="35" pageNumber="36" precision="925" value="10.466666">10°28’N</geoCoordinate>
.
<date id="FFDD1024FFDDFFE2FB6945A1BA3D3DFE" box="[1217,1359,1962,1988]" pageId="35" pageNumber="36" value="1969-09-12">
<collectingDate id="EF99E9CCFFDDFFE2FB6945A1BA3D3DFE" box="[1217,1359,1962,1988]" pageId="35" pageNumber="36" value="1969-09-12">12-IX-1969</collectingDate>
</date>
.
<collectorName id="26965332FFDDFFE2FAF245A1BE6B3DD2" pageId="35" pageNumber="36">G. Halffter</collectorName>
y
<collectorName id="26965332FFDDFFE2FE9F45C4BEE83DD3" box="[311,410,1999,2025]" pageId="35" pageNumber="36">P. Reyes</collectorName>
col.”;
<collectionCode id="ED72AE21FFDDFFE2FE4145C5BD553DD2" box="[489,551,1998,2024]" pageId="35" pageNumber="36">GHC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDDFFE2FDE845C5BBF73DD2" ID-GBIF-Occurrence="3093980319" box="[576,1157,1998,2025]" collectingDate="1969-09-12" collectionCode="VMC" collectorName="G. Halffter &amp; P. Reyes" country="Costa Rica" latitude="10.466666" location="Rio Puerto Viejo" longLatPrecision="0" longitude="-84.0" municipality="Estacion La Selva" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="9D65FD6DFFDDFFE2FDE845C5BD1D3DD3" box="[576,623,1998,2025]" pageId="35" pageNumber="36" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFDDFFE2FB9645C5BBF73DD2" box="[1086,1157,1998,2024]" pageId="35" pageNumber="36">VMC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36E4FFDAFFE5FF154302BA693970" blockId="36.[189,1399,264,843]" pageId="36" pageNumber="37">
<materialsCitation id="3B0B3CB9FFDAFFE5FF154302BE633B72" ID-GBIF-Occurrence="3093980489" collectingDate="1982-06-17" collectingDateMax="1982-06-21" collectingDateMin="1982-06-17" collectionCode="TAMU" collectorName="B. Gill." country="Panama" elevation="1200" location="Cerro Hornito" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui">
<collectingCountry id="F3747674FFDAFFE5FF154302BE4B3B19" box="[189,313,265,291]" name="Panama" pageId="36" pageNumber="37">PANAMA</collectingCountry>
–
<emphasis id="B917EAF6FFDAFFE5FEF54303BEBB3B19" bold="true" box="[349,457,264,291]" pageId="36" pageNumber="37">
<collectingRegion id="49A7F806FFDAFFE5FEF54303BEBB3B19" box="[349,457,264,291]" country="Panama" name="Chiriqui" pageId="36" pageNumber="37">Chiriquí</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFDAFFE5FE484302BD633B19" box="[480,529,265,291]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FD844302BDA93B19" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FD844302BDA93B19" box="[556,731,265,291]" country="Panama" name="Cerro Hornito" pageId="36" pageNumber="37" stateProvince="Chiriqui">Cerro Hornito</location>
,
<locationDeviation id="9A6E2FC6FFDAFFE5FD434302BCA13B19" box="[747,979,265,292]" location="Gualaca." pageId="36" pageNumber="37">
<quantity id="4C9B9B01FFDAFFE5FD434302BC4B3B19" box="[747,825,265,292]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="36" pageNumber="37" unit="km" value="15.0">15 km</quantity>
W Gualaca.
</locationDeviation>
<date id="FFDD1024FFDAFFE5FC754302BBE83B19" box="[989,1178,265,291]" pageId="36" pageNumber="37" value="1982-06-17" valueMax="1982-06-21" valueMin="1982-06-17">
<collectingDate id="EF99E9CCFFDAFFE5FC754302BBE83B19" box="[989,1178,265,291]" pageId="36" pageNumber="37" value="1982-06-17" valueMax="1982-06-21" valueMin="1982-06-17">17–21 VI 1982</collectingDate>
</date>
.
<collectorName id="26965332FFDAFFE5FB014302BB8B3B19" box="[1193,1273,265,291]" pageId="36" pageNumber="37">B.Gill.</collectorName>
<quantity id="4C9B9B01FFDAFFE5FAAC4302BA173B19" box="[1284,1381,265,291]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="36" pageNumber="37" unit="m" value="1200.0">
<elevation id="004ED1D7FFDAFFE5FAAC4302BA173B19" box="[1284,1381,265,291]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="36" pageNumber="37" unit="m" value="1200.0">1200 m</elevation>
</quantity>
”;
<collectionCode id="ED72AE21FFDAFFE5FF154325BE633B72" box="[189,273,302,328]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FE9A4325BA2B3B72" ID-GBIF-Occurrence="3093980477" box="[306,1369,301,328]" collectingDate="1982-06-21" collectingDateMax="1982-07-16" collectingDateMin="1982-06-21" collectionCode="TAMU" collectorName="B. Gill." country="Panama" elevation="1200" location="Cerro Hornito" pageId="36" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Chiriqui">
<specimenCount id="9D65FD6DFFDAFFE5FE9A4325BE113B72" box="[306,355,302,328]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FED74325BD5C3B72" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FED74325BD5C3B72" box="[383,558,302,328]" country="Panama" name="Cerro Hornito" pageId="36" pageNumber="37" stateProvince="Chiriqui">Cerro Hornito</location>
,
<locationDeviation id="9A6E2FC6FFDAFFE5FD964326BC573B72" box="[574,805,301,328]" location="Gualaca." pageId="36" pageNumber="37">
<quantity id="4C9B9B01FFDAFFE5FD964326BDF93B72" box="[574,651,301,328]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="36" pageNumber="37" unit="km" value="15.0">15 km</quantity>
W Gualaca.
</locationDeviation>
<date id="FFDD1024FFDAFFE5FC984325BB683B72" box="[816,1050,302,328]" pageId="36" pageNumber="37" value="1982-06-21" valueMax="1982-07-16" valueMin="1982-06-21">
<collectingDate id="EF99E9CCFFDAFFE5FC984325BB683B72" box="[816,1050,302,328]" pageId="36" pageNumber="37" value="1982-06-21" valueMax="1982-07-16" valueMin="1982-06-21">21 VI-16 VII 1982</collectingDate>
</date>
.
<collectorName id="26965332FFDAFFE5FB824325BB0B3B72" box="[1066,1145,302,328]" pageId="36" pageNumber="37">B.Gill.</collectorName>
<quantity id="4C9B9B01FFDAFFE5FB2D4325BB973B72" box="[1157,1253,302,328]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="36" pageNumber="37" unit="m" value="1200.0">
<elevation id="004ED1D7FFDAFFE5FB2D4325BB973B72" box="[1157,1253,302,328]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="36" pageNumber="37" unit="m" value="1200.0">1200 m</elevation>
</quantity>
”;
<collectionCode id="ED72AE21FFDAFFE5FAA94325BA2B3B72" box="[1281,1369,302,328]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B0B3CB9FFDAFFE5FF15435ABB703B57" ID-GBIF-Occurrence="3093980563" box="[189,1026,337,365]" collectingDate="1985-06-10" collectingDateMax="1985-06-13" collectingDateMin="1985-06-10" collectionCode="CNMC" collectorName="E. Kiley &amp; D. Rider" country="Panama" location="El Valle" pageId="36" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Cocle">
<emphasis id="B917EAF6FFDAFFE5FF15435ABE713B56" bold="true" box="[189,259,337,364]" pageId="36" pageNumber="37">
<collectingRegion id="49A7F806FFDAFFE5FF15435ABE713B56" box="[189,259,337,364]" country="Panama" name="Cocle" pageId="36" pageNumber="37">Coclé</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFDAFFE5FE894358BE223B57" box="[289,336,339,365]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FEC44359BEA23B56" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FEC44359BEA23B56" box="[364,464,338,364]" country="Panama" name="El Valle" pageId="36" pageNumber="37" stateProvince="Cocle">El Valle</location>
,
<date id="FFDD1024FFDAFFE5FE764359BDEE3B57" box="[478,668,338,365]" pageId="36" pageNumber="37" value="1985-06-10" valueMax="1985-06-13" valueMin="1985-06-10">
<collectingDate id="EF99E9CCFFDAFFE5FE764359BDEE3B57" box="[478,668,338,365]" pageId="36" pageNumber="37" value="1985-06-10" valueMax="1985-06-13" valueMin="1985-06-10">VI-10–13–1985</collectingDate>
</date>
,
<collectorName id="26965332FFDAFFE5FD024359BC773B56" box="[682,773,338,364]" pageId="36" pageNumber="37">E.Kiley</collectorName>
&amp;
<collectorName id="26965332FFDAFFE5FC874359BCE13B56" box="[815,915,338,364]" pageId="36" pageNumber="37">D.Rider</collectorName>
”;
<collectionCode id="ED72AE21FFDAFFE5FC044358BB703B57" box="[940,1026,339,365]" collectionName="USA, Colorado, Fruita, Colorado National Monument" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" lsid="urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="36" pageNumber="37">CNMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FB884358BECC3BAB" ID-GBIF-Occurrence="3093980391" collectingDate="1985-06-10" collectingDateMax="1985-06-13" collectingDateMin="1985-06-10" collectionCode="TAMU" collectorName="E. Kiley &amp; D. Rider" country="Panama" location="Cocle" pageId="36" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Cocle">
<specimenCount id="9D65FD6DFFDAFFE5FB884358BB3D3B57" box="[1056,1103,339,365]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFDAFFE5FEC2437CBECC3BAB" box="[362,446,375,401]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FE70437CBB1A3BAB" ID-GBIF-Occurrence="3093980456" box="[472,1128,375,402]" collectingDate="1985-06-10" collectingDateMax="1985-06-13" collectingDateMin="1985-06-10" collectionCode="UVGC" collectorName="E. Kiley &amp; D. Rider" country="Panama" location="Cocle" pageId="36" pageNumber="37" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cocle">
<specimenCount id="9D65FD6DFFDAFFE5FE70437CBD743BA8" box="[472,518,375,402]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D65FD6DFFDAFFE5FDBB437CBD333BA8" box="[531,577,375,402]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFDAFFE5FBBD437CBB1A3BAB" box="[1045,1128,375,401]" collectionName="Guatemala, Guatemala City, Universidad del Valle de Guatemala, Collecion de Artropodos" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="36" pageNumber="37">UVGC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FB2A437CBD7B3B8C" ID-GBIF-Occurrence="3093980335" collectingDate="1985-06-10" collectingDateMax="1985-06-13" collectingDateMin="1985-06-10" collectionCode="VMC" collectorName="E. Kiley &amp; D. Rider" country="Panama" location="Cocle" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Cocle">
<specimenCount id="9D65FD6DFFDAFFE5FB2A437CBBC03BA8" box="[1154,1202,375,402]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFDAFFE5FE6E4397BD7B3B8C" box="[454,521,412,438]" pageId="36" pageNumber="37">VMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FD8C4397BA163B8C" ID-GBIF-Occurrence="3093980304" box="[548,1380,412,439]" collectingDate="1945-06-05" collectionCode="TAMU" collectorName="C. D. Michener" country="Panama" elevation="762" location="El Valle de Anton" pageId="36" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Cocle">
<specimenCount id="9D65FD6DFFDAFFE5FD8C4397BD203B8C" box="[548,594,412,438]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FDC54397BC363B8C" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FDC54397BC363B8C" box="[621,836,412,438]" country="Panama" name="El Valle de Anton" pageId="36" pageNumber="37" stateProvince="Cocle">El Valle de Antón</location>
,
<date id="FFDD1024FFDAFFE5FCF94397BCBF3B8D" box="[849,973,412,439]" pageId="36" pageNumber="37" value="1945-06-05">
<collectingDate id="EF99E9CCFFDAFFE5FCF94397BCBF3B8D" box="[849,973,412,439]" pageId="36" pageNumber="37" value="1945-06-05">VI 5 1945</collectingDate>
</date>
.
<quantity id="4C9B9B01FFDAFFE5FC724397BB453B8C" box="[986,1079,412,439]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="36" pageNumber="37" unit="ft" value="2500.0">
<elevation id="004ED1D7FFDAFFE5FC724397BB453B8C" box="[986,1079,412,439]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="36" pageNumber="37" unit="ft" value="2500.0">2500 ft.</elevation>
</quantity>
<collectorName id="26965332FFDAFFE5FB974397BB843B8C" box="[1087,1270,412,438]" pageId="36" pageNumber="37">C.D. Michener</collectorName>
”;
<collectionCode id="ED72AE21FFDAFFE5FAA44397BA163B8C" box="[1292,1380,412,438]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
.–
<materialsCitation id="3B0B3CB9FFDAFFE5FF1543CBBB383BE1" ID-GBIF-Occurrence="3093980597" box="[189,1098,448,475]" collectingDate="1966-09-05" collectionCode="TAMU" collectorName="M. G. Naumann" country="Panama" location="Cerro Campana." pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<emphasis id="B917EAF6FFDAFFE5FF1543CBBE563BE1" bold="true" box="[189,292,448,475]" pageId="36" pageNumber="37">
<collectingRegion id="49A7F806FFDAFFE5FF1543CBBE563BE1" box="[189,292,448,475]" country="Panama" name="Panama" pageId="36" pageNumber="37">Panamá</collectingRegion>
</emphasis>
•
<specimenCount id="9D65FD6DFFDAFFE5FEEA43CABE013BE1" box="[322,371,449,475]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FE2443CABD203BE1" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FE2443CABD203BE1" box="[396,594,449,475]" country="Panama" name="Cerro Campana." pageId="36" pageNumber="37" stateProvince="Panama">Cerro Campana.</location>
<date id="FFDD1024FFDAFFE5FDF443CBBDA93BE1" box="[604,731,448,475]" pageId="36" pageNumber="37" value="1966-09-05">
<collectingDate id="EF99E9CCFFDAFFE5FDF443CBBDA93BE1" box="[604,731,448,475]" pageId="36" pageNumber="37" value="1966-09-05">5-IX-1966</collectingDate>
</date>
.
<collectorName id="26965332FFDAFFE5FD4143CABCD03BE1" box="[745,930,449,475]" pageId="36" pageNumber="37">M.G. Naumann</collectorName>
col.”;
<collectionCode id="ED72AE21FFDAFFE5FC5E43CABB383BE1" box="[1014,1098,449,475]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FBC043CABDA03BC5" ID-GBIF-Occurrence="3093980600" collectingDate="1929-05" collectionCode="TAMU" country="Panama" location="Barro Colorado. Canal Zone." pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FBC043CABBEB3BE1" box="[1128,1177,449,475]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FB1A43CABE3D383A" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FB1A43CABE3D383A" country="Panama" name="Barro Colorado. Canal Zone." pageId="36" pageNumber="37" stateProvince="Panama">Barro Colorado. Canal Zone.</location>
<date id="FFDD1024FFDAFFE5FEFF43EDBEA63BC5" box="[343,468,485,512]" pageId="36" pageNumber="37" value="1929-05">
<collectingDate id="EF99E9CCFFDAFFE5FEFF43EDBEA63BC5" box="[343,468,485,512]" pageId="36" pageNumber="37" value="1929-05">May 1929</collectingDate>
</date>
. Darlington”;
<collectionCode id="ED72AE21FFDAFFE5FDD643EEBDA03BC5" box="[638,722,485,511]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FD4743EEBE97381E" ID-GBIF-Occurrence="3093980497" collectingDate="1975-12-06" collectingMethod="Rat carrion trap" collectionCode="TAMU" collectorName="Col. O. P. Young" country="Panama" location="Canal Zone" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FD4743EEBC6D383A" box="[751,799,485,512]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FC9043EEBCB6383A" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FC9043EEBCB6383A" box="[824,964,485,512]" country="Panama" name="Canal Zone" pageId="36" pageNumber="37" stateProvince="Panama">Canal Zone</location>
B.C.
<date id="FFDD1024FFDAFFE5FBAC43EEBB59383A" box="[1028,1067,485,512]" pageId="36" pageNumber="37">
I.
<collectingDate id="EF99E9CCFFDAFFE5FBB543EEBB59383A" box="[1053,1067,485,512]" pageId="36" pageNumber="37" value="1975-12-06">6</collectingDate>
</date>
-XII-1975.
<collectingMethod id="52224EF3FFDAFFE5FB1B43EDBA00383A" box="[1203,1394,485,512]" pageId="36" pageNumber="37">Rat carrion trap</collectingMethod>
.
<collectorName id="26965332FFDAFFE5FF154001BE09381E" box="[189,379,522,548]" pageId="36" pageNumber="37">Col. O.P.Young</collectorName>
”;
<collectionCode id="ED72AE21FFDAFFE5FE394001BE97381E" box="[401,485,522,548]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FDA84001BE723873" ID-GBIF-Occurrence="3093980349" collectingDate="1975-11" collectingMethod="Human feces trap" collectionCode="VMC" collectorName="Col. O. P. Young" country="Panama" location="Canal Zone" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FDA84001BD5D381F" box="[512,559,522,549]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FDEF4001BDA0381F" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FDEF4001BDA0381F" box="[583,722,522,549]" country="Panama" name="Canal Zone" pageId="36" pageNumber="37" stateProvince="Panama">Canal Zone</location>
B.C.I. 1,2-
<date id="FFDD1024FFDAFFE5FCFE4001BCCD381F" box="[854,959,522,549]" pageId="36" pageNumber="37" value="1975-11">
<collectingDate id="EF99E9CCFFDAFFE5FCFE4001BCCD381F" box="[854,959,522,549]" pageId="36" pageNumber="37" value="1975-11">XI-1975</collectingDate>
</date>
.
<collectingMethod id="52224EF3FFDAFFE5FC624000BBEC381F" box="[970,1182,522,549]" pageId="36" pageNumber="37">Human feces trap</collectingMethod>
.
<collectorName id="26965332FFDAFFE5FB024001BA15381E" box="[1194,1383,522,548]" pageId="36" pageNumber="37">Col. O.P.Young</collectorName>
”;
<collectionCode id="ED72AE21FFDAFFE5FF154024BE723873" box="[189,256,559,585]" pageId="36" pageNumber="37">VMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FEB34024BBBF3873" ID-GBIF-Occurrence="3093980592" box="[283,1229,558,585]" collectingDate="1975-12-08" collectingMethod="Rat carrion trap" collectionCode="TAMU" collectorName="Col. O. P. Young" country="Panama" location="Canal Zone" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FEB34024BE393873" box="[283,331,559,585]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FECA4024BE9C3873" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FECA4024BE9C3873" box="[354,494,559,585]" country="Panama" name="Canal Zone" pageId="36" pageNumber="37" stateProvince="Panama">Canal Zone</location>
B.C.
<date id="FFDD1024FFDAFFE5FD854024BD203873" box="[557,594,558,585]" pageId="36" pageNumber="37">
I.
<collectingDate id="EF99E9CCFFDAFFE5FDED4025BD203873" box="[581,594,558,585]" pageId="36" pageNumber="37" value="1975-12-08">8</collectingDate>
</date>
-XII-1975.
<collectingMethod id="52224EF3FFDAFFE5FD734024BCEA3873" box="[731,920,559,585]" pageId="36" pageNumber="37">Rat carrion trap</collectingMethod>
.
<collectorName id="26965332FFDAFFE5FC0C4024BB103873" box="[932,1122,559,585]" pageId="36" pageNumber="37">Col. O.P.Young</collectorName>
”;
<collectionCode id="ED72AE21FFDAFFE5FBD14024BBBF3873" box="[1145,1229,559,585]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FB404024BC4E3854" ID-GBIF-Occurrence="3093980374" collectingDate="1985-06-04" collectingDateMax="1985-06-21" collectingDateMin="1985-06-04" collectionCode="IEXA" collectorName="E. G., Riley" country="Panama" location="Fort Kobbe" municipality="Canal Zone" pageId="36" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FB404024BA643873" box="[1256,1302,559,585]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; “
<collectingMunicipality id="6BB8AC9EFFDAFFE5FA874024BF8E3854" pageId="36" pageNumber="37">Canal Zone</collectingMunicipality>
,
<location id="8EBC603FFFDAFFE5FEAF405FBEE23857" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FEAF405FBEE23857" box="[263,400,595,622]" country="Panama" municipality="Canal Zone" name="Fort Kobbe" pageId="36" pageNumber="37" stateProvince="Panama">Fort Kobbe</location>
,
<date id="FFDD1024FFDAFFE5FE334058BD3E3854" box="[411,588,595,622]" pageId="36" pageNumber="37" value="1985-06-04" valueMax="1985-06-21" valueMin="1985-06-04">
<collectingDate id="EF99E9CCFFDAFFE5FE334058BD3E3854" box="[411,588,595,622]" pageId="36" pageNumber="37" value="1985-06-04" valueMax="1985-06-21" valueMin="1985-06-04">VI-4–21–1985</collectingDate>
</date>
.
<collectorName id="26965332FFDAFFE5FDFE405FBDAD3857" box="[598,735,595,622]" pageId="36" pageNumber="37">E.G., Riley</collectorName>
”;
<collectionCode id="ED72AE21FFDAFFE5FD5E405FBC4E3854" box="[758,828,596,622]" pageId="36" pageNumber="37">IEXA</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FCFA405FBEE738A8" ID-GBIF-Occurrence="3093980503" collectingDate="1985-06-04" collectingDateMax="1985-06-21" collectingDateMin="1985-06-04" collectionCode="TAMU" collectorName="E. G., Riley" country="Panama" location="Fort Kobbe" municipality="Canal Zone" pageId="36" pageNumber="37" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Panama">
<collectionCode id="ED72AE21FFDAFFE5FCFA405FBCD93854" box="[850,939,596,622]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
:
<specimenCount id="9D65FD6DFFDAFFE5FC1D405FBC913854" box="[949,995,596,622]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D65FD6DFFDAFFE5FC47405FBB433854" box="[1007,1073,596,622]" pageId="36" pageNumber="37" type="female">3 ♀♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFDAFFE5FEE94073BEE738A8" box="[321,405,632,658]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FE194073BC8C38A8" ID-GBIF-Occurrence="3093980555" box="[433,1022,632,659]" collectingDate="1985-06-04" collectingDateMax="1985-06-21" collectingDateMin="1985-06-04" collectionCode="VMC" collectorName="E. G., Riley" country="Panama" isTailEnumeration="true" location="Fort Kobbe" municipality="Canal Zone" pageId="36" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FE194073BE9238A9" box="[433,480,632,659]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFDAFFE5FC134073BC8C38A8" box="[955,1022,632,658]" pageId="36" pageNumber="37">VMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FBB24073BD2E388D" ID-GBIF-Occurrence="3093980618" collectingDate="1982-08-01" collectingDateMax="1982-08-22" collectingDateMin="1982-08-01" collectionCode="TAMU" collectorName="B-Gill" country="Panama" elevation="400" location="Chepo-Carti Rd." pageId="36" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FBB24073BB3B38A9" box="[1050,1097,632,659]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FBCA4073BA5B38A8" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FBCA4073BA5B38A8" box="[1122,1321,632,658]" country="Panama" name="Chepo-Carti Rd." pageId="36" pageNumber="37" stateProvince="Panama">Chepo-Carti Rd.</location>
<date id="FFDD1024FFDAFFE5FA9A4073BE4A388D" pageId="36" pageNumber="37" value="1982-08-01" valueMax="1982-08-22" valueMin="1982-08-01">
<collectingDate id="EF99E9CCFFDAFFE5FA9A4073BE4A388D" pageId="36" pageNumber="37" value="1982-08-01" valueMax="1982-08-22" valueMin="1982-08-01">1–22- VIII-1982</collectingDate>
</date>
.
<collectorName id="26965332FFDAFFE5FEEC4096BEE1388D" box="[324,403,669,695]" pageId="36" pageNumber="37">B-Gill</collectorName>
.
<quantity id="4C9B9B01FFDAFFE5FE374096BE9D388D" box="[415,495,669,695]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="36" pageNumber="37" unit="m" value="400.0">
<elevation id="004ED1D7FFDAFFE5FE374096BE9D388D" box="[415,495,669,695]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="36" pageNumber="37" unit="m" value="400.0">400 m</elevation>
</quantity>
”;
<collectionCode id="ED72AE21FFDAFFE5FDA04096BD2E388D" box="[520,604,669,695]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FDD04096BA11388D" ID-GBIF-Occurrence="3093980316" box="[632,1379,669,695]" collectingDate="1982-06-06" collectingDateMax="1982-06-24" collectingDateMin="1982-06-06" collectionCode="CNMC" collectorName="B-Gill" country="Panama" elevation="400" location="Chepo-Carti Rd." pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FDD04096BDDB388D" box="[632,681,669,695]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FD694096BCFA388D" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FD694096BCFA388D" box="[705,904,669,695]" country="Panama" name="Chepo-Carti Rd." pageId="36" pageNumber="37" stateProvince="Panama">Chepo-Carti Rd.</location>
<date id="FFDD1024FFDAFFE5FC384096BB49388D" box="[912,1083,669,695]" pageId="36" pageNumber="37" value="1982-06-06" valueMax="1982-06-24" valueMin="1982-06-06">
<collectingDate id="EF99E9CCFFDAFFE5FC384096BB49388D" box="[912,1083,669,695]" pageId="36" pageNumber="37" value="1982-06-06" valueMax="1982-06-24" valueMin="1982-06-06">6–24-VI-1982</collectingDate>
</date>
.
<collectorName id="26965332FFDAFFE5FBE04096BBE5388D" box="[1096,1175,669,695]" pageId="36" pageNumber="37">B-Gill</collectorName>
.
<quantity id="4C9B9B01FFDAFFE5FB0B4096BB81388D" box="[1187,1267,669,695]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="36" pageNumber="37" unit="m" value="400.0">
<elevation id="004ED1D7FFDAFFE5FB0B4096BB81388D" box="[1187,1267,669,695]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="36" pageNumber="37" unit="m" value="400.0">400 m</elevation>
</quantity>
”;
<collectionCode id="ED72AE21FFDAFFE5FAA54096BA11388D" box="[1293,1379,669,695]" collectionName="USA, Colorado, Fruita, Colorado National Monument" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" lsid="urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="36" pageNumber="37">CNMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FF1540C9BD8438E6" ID-GBIF-Occurrence="3093980566" box="[189,758,706,732]" collectingDate="1982-06-06" collectingDateMax="1982-06-24" collectingDateMin="1982-06-06" collectionCode="IEXA" collectorName="B-Gill" country="Panama" elevation="400" location="Chepo-Carti Rd." pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FF1540C9BF9838E6" box="[189,234,706,732]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFDAFFE5FD1840C9BD8438E6" box="[688,758,706,732]" pageId="36" pageNumber="37">IEXA</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FCA240C9BA1438E6" ID-GBIF-Occurrence="3093980521" box="[778,1382,706,732]" collectingDate="1982-06-06" collectingDateMax="1982-06-24" collectingDateMin="1982-06-06" collectionCode="TAMU" collectorName="B-Gill" country="Panama" elevation="400" location="Chepo-Carti Rd." pageId="36" pageNumber="37" specimenCount="2" specimenCount-female="2" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FCA240C9BC3E38E6" box="[778,844,706,732]" pageId="36" pageNumber="37" type="female">2 ♀♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED72AE21FFDAFFE5FABA40C9BA1438E6" box="[1298,1382,706,732]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FF1540EDBA58393A" ID-GBIF-Occurrence="3093980434" box="[189,1322,742,769]" collectingDate="8 Sept" collectingMethod="Manure trap" collectionCode="VMC" collectorName="H. Stockwell." country="Panama" latitude="9.166667" location="9 km SE Bayano Bridge." longLatPrecision="1300" longitude="-78.76667" pageId="36" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FF1540EDBF98393B" box="[189,234,742,769]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; “
<locationDeviation id="9A6E2FC6FFDAFFE5FEAC40EDBD54393A" box="[260,550,742,769]" pageId="36" pageNumber="37">
<quantity id="4C9B9B01FFDAFFE5FEAC40EDBE32393A" box="[260,320,742,768]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="36" pageNumber="37" unit="km" value="9.0">9 km</quantity>
SE Bayano Bridge
</locationDeviation>
.
<geoCoordinate id="EE575023FFDAFFE5FD9840EDBDFB393A" box="[560,649,742,768]" degrees="9" direction="north" minutes="10" orientation="latitude" pageId="36" pageNumber="37" precision="925" value="9.166667">9°10’N</geoCoordinate>
,
<geoCoordinate id="EE575023FFDAFFE5FD3D40EDBC70393A" box="[661,770,742,768]" degrees="78" direction="west" minutes="46" orientation="longitude" pageId="36" pageNumber="37" precision="925" value="-78.76667">78°46’W</geoCoordinate>
.
<date id="FFDD1024FFDAFFE5FCA540EDBC25393A" box="[781,855,742,768]" pageId="36" pageNumber="37">
<collectingDate id="EF99E9CCFFDAFFE5FCA540EDBC25393A" box="[781,855,742,768]" pageId="36" pageNumber="37">8 Sept</collectingDate>
</date>
’74.
<collectorName id="26965332FFDAFFE5FC3C40ECBB47393A" box="[916,1077,742,769]" pageId="36" pageNumber="37">H. Stockwell.</collectorName>
<collectingMethod id="52224EF3FFDAFFE5FB9340ECBBA3393A" box="[1083,1233,742,769]" pageId="36" pageNumber="37">Manure trap</collectingMethod>
”;
<collectionCode id="ED72AE21FFDAFFE5FB4F40EDBA58393A" box="[1255,1322,742,768]" pageId="36" pageNumber="37">VMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FAEA40EDBB54391F" ID-GBIF-Occurrence="3093980598" collectingDate="1929-07-01" collectionCode="GHC" collectorName="Collector C. H. Curran" country="Panama" location="Barro Colo Isld. Canal Zone." pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FAEA40EDBA03393B" box="[1346,1393,742,769]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FF6F4100BD5C391C" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FF6F4100BD5C391C" box="[199,558,779,806]" country="Panama" name="Barro Colo Isld. Canal Zone." pageId="36" pageNumber="37" stateProvince="Panama">Barro Colo Isld. Canal Zone.</location>
<date id="FFDD1024FFDAFFE5FD904100BDD0391F" box="[568,674,779,805]" pageId="36" pageNumber="37" value="1929-07-01">
<collectingDate id="EF99E9CCFFDAFFE5FD904100BDD0391F" box="[568,674,779,805]" pageId="36" pageNumber="37" value="1929-07-01">1.7.1929</collectingDate>
</date>
.
<collectorName id="26965332FFDAFFE5FD194100BCBD391F" box="[689,975,779,806]" pageId="36" pageNumber="37">Collector C. H. Curran</collectorName>
”;
<collectionCode id="ED72AE21FFDAFFE5FC404100BB54391F" box="[1000,1062,779,805]" pageId="36" pageNumber="37">GHC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FBEF4100BE843970" ID-GBIF-Occurrence="3093980543" collectionCode="VMC" collectorName="F. S. Blanton Collr." country="Panama" location="Canal Zone." pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FBEF4100BB0B391C" box="[1095,1145,779,806]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FB3C4100BA58391C" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FB3C4100BA58391C" box="[1172,1322,779,806]" country="Panama" name="Canal Zone." pageId="36" pageNumber="37" stateProvince="Panama">Canal Zone.</location>
1951.
<collectorName id="26965332FFDAFFE5FF15413BBEEF3970" box="[189,413,816,842]" pageId="36" pageNumber="37">F.S. Blanton Collr.</collectorName>
”;
<collectionCode id="ED72AE21FFDAFFE5FE1B413BBE843970" box="[435,502,816,842]" pageId="36" pageNumber="37">VMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B0B3CB9FFDAFFE5FDA7413BBA653970" ID-GBIF-Occurrence="3093980562" box="[527,1303,816,843]" collectingDate="1999-02-15" collectingDateMax="1999-02-24" collectingDateMin="1999-02-15" collectionCode="TAMU" collectorName="J. E. Wappes" country="Panama" location="Soberania Nac. PQ." pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<specimenCount id="9D65FD6DFFDAFFE5FDA7413BBD4C3970" box="[527,574,816,842]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; “
<location id="8EBC603FFFDAFFE5FDFD413BBC4D3970" LSID="urn:lsid:plazi:treatment:03CA87F2FFDDFFE7FDC94688BD063B71:8EBC603FFFDAFFE5FDFD413BBC4D3970" box="[597,831,816,842]" country="Panama" name="Soberania Nac. PQ." pageId="36" pageNumber="37" stateProvince="Panama">Soberania Nac. PQ.</location>
<date id="FFDD1024FFDAFFE5FCEE413BBB7A3970" box="[838,1032,816,843]" pageId="36" pageNumber="37" value="1999-02-15" valueMax="1999-02-24" valueMin="1999-02-15">
<collectingDate id="EF99E9CCFFDAFFE5FCEE413BBB7A3970" box="[838,1032,816,843]" pageId="36" pageNumber="37" value="1999-02-15" valueMax="1999-02-24" valueMin="1999-02-15">15–24 Feb 1999</collectingDate>
</date>
.
<collectorName id="26965332FFDAFFE5FBBC413BBBD83970" box="[1044,1194,816,842]" pageId="36" pageNumber="37">J.E. Wappes</collectorName>
”;
<collectionCode id="ED72AE21FFDAFFE5FB17413BBA653970" box="[1215,1303,816,842]" collectionName="USA, Texas, College Station, Texas A &amp; M University" pageId="36" pageNumber="37">TAMU</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36E4FFDAFFE5FF154178BE2F39B6" blockId="36.[189,349,882,909]" box="[189,349,882,909]" pageId="36" pageNumber="37">
<heading id="D0948188FFDAFFE5FF154178BE2F39B6" bold="true" box="[189,349,882,909]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B917EAF6FFDAFFE5FF154178BE2F39B6" bold="true" box="[189,349,882,909]" pageId="36" pageNumber="37">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFDAFFE5FF1541A8BD273987" blockId="36.[189,597,931,957]" box="[189,597,931,957]" pageId="36" pageNumber="37">
<collectingCountry id="F3747674FFDAFFE5FF1541A8BE6F3987" box="[189,285,931,957]" name="Panama" pageId="36" pageNumber="37">Panama</collectingCountry>
,
<collectingCountry id="F3747674FFDAFFE5FE8141A8BEFB3987" box="[297,393,931,957]" name="Panama" pageId="36" pageNumber="37">Panama</collectingCountry>
, Cerro Campana.
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFDAFFE4FF1541EEBB783926" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
<paragraph id="8BDC36E4FFDAFFE5FF1541EEBE3C39C5" blockId="36.[189,334,997,1023]" box="[189,334,997,1023]" pageId="36" pageNumber="37">
<heading id="D0948188FFDAFFE5FF1541EEBE3C39C5" bold="true" box="[189,334,997,1023]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B917EAF6FFDAFFE5FF1541EEBE3C39C5" bold="true" box="[189,334,997,1023]" pageId="36" pageNumber="37">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFDAFFE5FF42461EBE8D3E0B" blockId="36.[189,511,1045,1109]" box="[234,511,1045,1073]" pageId="36" pageNumber="37">
<emphasis id="B917EAF6FFDAFFE5FF42461EBE0F3E0A" bold="true" box="[234,381,1045,1072]" pageId="36" pageNumber="37">Major male</emphasis>
(
<typeStatus id="54D88846FFDAFFE5FE25461DBE853E0B" box="[397,503,1046,1073]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="8BDC36E4FFDAFFE5FF154631BEF53E6F" blockId="36.[189,511,1045,1109]" box="[189,391,1082,1109]" pageId="36" pageNumber="37">
Length
<quantity id="4C9B9B01FFDAFFE5FEBF4630BEF13E6F" box="[279,387,1083,1109]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="36" pageNumber="37" unit="mm" value="19.3">19.3 mm</quantity>
.
</paragraph>
<paragraph id="8BDC36E4FFDAFFE5FF154675BD593F3C" blockId="36.[189,1401,1149,1286]" pageId="36" pageNumber="37">
HEAD. Clypeus bidentate, black on anterior margin, bright metallic red, with green sheen on posterior portion, roughened sculpture. Genae bright metallic red, with green sheen; roughened sculpture. Front black, bright metallic red on portions adjacent to cephalic horn. Cephalic horn black, curved posteriorly over pronotum (
<figureCitation id="13582A61FFDAFFE5FED246E7BEA93F3C" box="[378,475,1260,1286]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="36" pageNumber="37">Figs 2H</figureCitation>
,
<figureCitation id="13582A61FFDAFFE5FE4046E7BD6C3F3C" box="[488,542,1260,1286]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="36" pageNumber="37">10A</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFDAFFE5FF154724BC203C1F" blockId="36.[189,1399,1326,1573]" pageId="36" pageNumber="37">
PRONOTUM. Uniformly black, with bright metallic red sheen, becoming completely black posteriorly and on lateral margins of posterolateral angles. Keel absent in the middle of anterior pronotal margin. Disc triangular, flat, with two weakly developed, elongate tubercles on anterior portion. Triangle smooth; scabriculous; impunctate. Sides with smooth sculpture; scabriculous; with almost effaced punctures. Lateral lines of pronotal triangle straight. Posterolateral angles weakly developed, widened, slightly projected posteriorly. Lateral fossae distinctly impressed. Basal fossae obtusely oval, distinctly impressed. Posterior margin impunctate (
<figureCitation id="13582A61FFDAFFE5FD084400BC733C1F" box="[672,769,1547,1573]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="36" pageNumber="37">Figs 2H</figureCitation>
,
<figureCitation id="13582A61FFDAFFE5FCA64400BC363C1F" box="[782,836,1547,1573]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="36" pageNumber="37">10A</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFDAFFE5FF154445BC0B3C8B" blockId="36.[189,1398,1614,1713]" pageId="36" pageNumber="37">
ELYTRA. Fine striae, smooth, dull black, with dark metallic blue sheen; impressed basally as distinct fossae, scabriculous, with almost effaced to effaced punctation. Interstriae black, smooth, scabriculous, impunctate. Sutural margin without apical tooth (
<figureCitation id="13582A61FFDAFFE5FD57449CBC1E3C8B" box="[767,876,1687,1713]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="36" pageNumber="37">Fig. 10A</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFDAFFE5FF1544D0BDCA3CCE" blockId="36.[189,696,1754,1781]" box="[189,696,1754,1781]" pageId="36" pageNumber="37">PROTIBIAE. Quadridentate with apical spine.</paragraph>
<paragraph id="8BDC36E4FFDAFFE5FF154515BD9F3D66" blockId="36.[188,1399,1821,1884]" pageId="36" pageNumber="37">TERGITE VIII. Bright metallic red, with green sheen, scabriculous; with rough, almost completely effaced punctures. Basal margin with thick, small setae.</paragraph>
<paragraph id="8BDC36E4FFDAFFE5FF15458EBD193DD2" blockId="36.[189,1398,1925,2024]" pageId="36" pageNumber="37">
GENITALIA. Right lobe of endophallite copulatrix more developed than left lobe. Right lobe obtusely triangular in shape, rounded superiorly, and weakly developed. Left lobe bent posteriorly. Central ridge and column similar in size (
<figureCitation id="13582A61FFDAFFE5FDAB45C5BD2F3DD2" box="[515,605,1998,2024]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta &amp; Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya &amp; Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta &amp; Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma &amp; Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="36" pageNumber="37">Fig. 1L</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFDBFFE4FF424303BE0F3B19" blockId="37.[189,1401,264,394]" box="[234,381,264,291]" pageId="37" pageNumber="38">
<emphasis id="B917EAF6FFDBFFE4FF424303BE0F3B19" bold="true" box="[234,381,264,291]" pageId="37" pageNumber="38">Minor male</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFDBFFE4FF154320BD8D3BB3" blockId="37.[189,1401,264,394]" pageId="37" pageNumber="38">Like the major male, except for the reduction of the secondary sexual characters (i.e., cephalic horn, pronotal triangle and tubercles, and pronotal posterolateral angles). Occasionally, the pronotal disc is completely bright metallic red, with green sheen.</paragraph>
<paragraph id="8BDC36E4FFDBFFE4FF4243BBBE373BF0" blockId="37.[188,1400,432,698]" box="[234,325,432,458]" pageId="37" pageNumber="38">
<emphasis id="B917EAF6FFDBFFE4FF4243BBBE373BF0" bold="true" box="[234,325,432,458]" pageId="37" pageNumber="38">Female</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFDBFFE4FF1543D9BB5C3880" blockId="37.[188,1400,432,698]" pageId="37" pageNumber="38">
Similar to the male, except for the head showing a cephalic trituberculate carina; with weakly developed tubercles; carinate middle tubercle, slightly more frontally projected than lateral tubercles; frons with distinctly impressed punctures; pronotal sculpture completely smooth, with superficially impressed punctures; pronotum almost completely dull black in the central portion, laterally and posteriorly bright metallic red with metallic green sheen; pronotal process trituberculate, lacking concavities; pronotal tubercles rounded, nearly aligned, well-spaced; with middle tubercle more developed than lateral tubercles; posterior pronotal midline almost completely effaced (
<figureCitation id="13582A61FFDBFFE4FC1D40ABBB523880" box="[949,1056,672,698]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="37" pageNumber="38">Fig. 10B</figureCitation>
).
</paragraph>
<paragraph id="8BDC36E4FFDBFFE4FF4240EBBE1238C0" blockId="37.[189,1034,736,797]" box="[234,352,736,762]" pageId="37" pageNumber="38">
<emphasis id="B917EAF6FFDBFFE4FF4240EBBE1238C0" bold="true" box="[234,352,736,762]" pageId="37" pageNumber="38">Variation</emphasis>
</paragraph>
<paragraph id="8BDC36E4FFDBFFE4FF154108BB783926" blockId="37.[189,1034,736,797]" box="[189,1034,770,797]" pageId="37" pageNumber="38">
Mean length
<quantity id="4C9B9B01FFDBFFE4FEF04108BEB23927" box="[344,448,771,797]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="37" pageNumber="38" unit="mm" value="17.8">17.8 mm</quantity>
(
<quantity id="4C9B9B01FFDBFFE4FE784109BDF33927" box="[464,641,770,797]" metricMagnitude="-2" metricUnit="m" metricValue="1.785" metricValueMax="2.11" metricValueMin="1.46" pageId="37" pageNumber="38" unit="mm" value="17.85" valueMax="21.1" valueMin="14.6">14.6–21.1 mm</quantity>
). Colour variation was not found.
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFDBFFE4FF154148BA5B3997" pageId="37" pageNumber="38" type="distribution">
<paragraph id="8BDC36E4FFDBFFE4FF154148BE243967" blockId="37.[189,342,835,861]" box="[189,342,835,861]" pageId="37" pageNumber="38">
<heading id="D0948188FFDBFFE4FF154148BE243967" bold="true" box="[189,342,835,861]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B917EAF6FFDBFFE4FF154148BE243967" bold="true" box="[189,342,835,861]" pageId="37" pageNumber="38">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFDBFFE4FF15417ABA5B3997" blockId="37.[189,1398,880,941]" pageId="37" pageNumber="38">
<collectingCountry id="F3747674FFDBFFE4FF15417ABE6839B1" box="[189,282,881,907]" name="Panama" pageId="37" pageNumber="38">Panama</collectingCountry>
and north-Caribbean
<collectingCountry id="F3747674FFDBFFE4FDB2417ABDEB39B0" box="[538,665,880,907]" name="Costa Rica" pageId="37" pageNumber="38">Costa Rica</collectingCountry>
(
<figureCitation id="13582A61FFDBFFE4FD01417BBC7339B1" box="[681,769,880,907]" captionStart="Fig" captionStartId="51.[189,232,1233,1259]" captionTargetBox="[193,1395,270,1197]" captionTargetId="figure-241@51.[189,1398,265,1199]" captionTargetPageId="51" captionText="Fig. 16. Distribution of the Phanaeus endymion species group in Central America (in part)." figureDoi="http://doi.org/10.5281/zenodo.4836798" httpUri="https://zenodo.org/record/4836798/files/figure.png" pageId="37" pageNumber="38">Fig. 16</figureCitation>
). The distributions of
<taxonomicName id="4C634D67FFDBFFE4FBAA417ABBD839B0" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[1026,1194,881,906]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B917EAF6FFDBFFE4FBAA417ABBD839B0" box="[1026,1194,881,906]" italics="true" pageId="37" pageNumber="38">P. panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224578DFFDBFFE4FB18417ABA7F39B1" box="[1200,1293,881,907]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C634D67FFDBFFE4FAB0417ABA0439B0" authorityName="Arnaud" authorityYear="2002" box="[1304,1398,880,906]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B917EAF6FFDBFFE4FAB0417ABA0439B0" box="[1304,1398,880,906]" italics="true" pageId="37" pageNumber="38">P. malyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C634D67FFDBFFE4FF47419FBE213997" baseAuthorityName="GBIF Secretariat" baseAuthorityYear="2019" box="[239,339,916,941]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pyrois">
<emphasis id="B917EAF6FFDBFFE4FF47419FBE213997" box="[239,339,916,941]" italics="true" pageId="37" pageNumber="38">P. pyrois</emphasis>
</taxonomicName>
show large areas of sympatry. Nevertheless, all these species are easily recognized.
</paragraph>
</subSubSection>
<subSubSection id="C379656FFFDBFFE7FF1541D8BD063B71" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BDC36E4FFDBFFE4FF1541D8BE5D39D7" blockId="37.[189,303,979,1005]" box="[189,303,979,1005]" pageId="37" pageNumber="38">
<heading id="D0948188FFDBFFE4FF1541D8BE5D39D7" bold="true" box="[189,303,979,1005]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B917EAF6FFDBFFE4FF1541D8BE5D39D7" bold="true" box="[189,303,979,1005]" pageId="37" pageNumber="38">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36E4FFDBFFE4FF15460ABA043E5A" blockId="37.[189,1398,1025,1120]" pageId="37" pageNumber="38">
<taxonomicName id="4C634D67FFDBFFE4FF15460ABEB53E21" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[189,455,1025,1051]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B917EAF6FFDBFFE4FF15460ABEB53E21" box="[189,455,1025,1051]" italics="true" pageId="37" pageNumber="38">Phanaeus panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224578DFFDBFFE4FE7A460ABD423E21" box="[466,560,1025,1051]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
has frequently been confused with
<taxonomicName id="4C634D67FFDBFFE4FC414609BB0E3E21" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[1001,1148,1025,1051]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B917EAF6FFDBFFE4FC414609BB0E3E21" box="[1001,1148,1025,1051]" italics="true" pageId="37" pageNumber="38">P. olsoufieffi</emphasis>
</taxonomicName>
by previous authors (
<bibRefCitation id="EFF24B15FFDBFFE4FF6D4628BE0F3E07" author="Edmonds W. D." box="[197,381,1059,1085]" pageId="37" pageNumber="38" pagination="1 - 105" refId="ref37487" refString="Edmonds W. D. 1994. Revision of Phanaeus Macleay, a New World genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Contributions in Science, Natural History Museum of Los Angeles County 443: 1 - 105." type="journal article" year="1994">Edmonds 1994</bibRefCitation>
;
<bibRefCitation id="EFF24B15FFDBFFE4FE234628BD483E07" author="Arnaud P." box="[395,570,1059,1085]" pageId="37" pageNumber="38" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b</bibRefCitation>
;
<bibRefCitation id="EFF24B15FFDBFFE4FDE04628BC1C3E07" author="Edmonds W. D. &amp; Zidek J." box="[584,878,1059,1085]" pageId="37" pageNumber="38" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. &amp; Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds &amp; Zídek 2012</bibRefCitation>
,
<bibRefCitation id="EFF24B15FFDBFFE4FCD44628BBD73E07" author="Solis A. &amp; Kohlmann B." box="[892,1189,1059,1085]" pageId="37" pageNumber="38" pagination="1 - 32" refId="ref42322" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica. Zootaxa 3482: 1 - 32." type="journal article" year="2012">Solís &amp; Kohlmann 2012</bibRefCitation>
;
<bibRefCitation id="EFF24B15FFDBFFE4FB1B4628BF8F3E5A" author="Kohlmann B. &amp; Arriaga-Jimenez A. &amp; Ros M." pageId="37" pageNumber="38" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. &amp; Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
Kohlmann
<emphasis id="B917EAF6FFDBFFE4FA91462FBA1D3E07" box="[1337,1391,1059,1085]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
. 2018
</bibRefCitation>
). Minor males and females of both species are strongly mimetic. More specimens from western
</paragraph>
<caption id="DF1C666CFFDBFFE4FF1745C7BB1D3DDC" ID-DOI="http://doi.org/10.5281/zenodo.4836786" ID-Zenodo-Dep="4836786" box="[191,1135,1996,2023]" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="37" pageNumber="38" startId="37.[191,234,1996,2022]" targetBox="[210,1374,1195,1954]" targetPageId="37">
<paragraph id="8BDC36E4FFDBFFE4FF1745C7BB1D3DDC" blockId="37.[191,1135,1996,2023]" box="[191,1135,1996,2023]" pageId="37" pageNumber="38">
<emphasis id="B917EAF6FFDBFFE4FF1745C7BE693DDC" bold="true" box="[191,283,1996,2022]" pageId="37" pageNumber="38">Fig. 10.</emphasis>
<taxonomicName id="4C634D67FFDBFFE4FE8845C7BD543DDC" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[288,550,1996,2022]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B917EAF6FFDBFFE4FE8845C7BD543DDC" box="[288,550,1996,2022]" italics="true" pageId="37" pageNumber="38">Phanaeus panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224578DFFDBFFE4FD8545C6BDFA3DDD" box="[557,648,1997,2023]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B917EAF6FFDBFFE4FD2545C7BDD03DDC" bold="true" box="[653,674,1996,2022]" pageId="37" pageNumber="38">A</emphasis>
. Holotype, ♂ (TAMU).
<emphasis id="B917EAF6FFDBFFE4FC6845C7BCA63DDC" bold="true" box="[960,980,1996,2022]" pageId="37" pageNumber="38">B</emphasis>
. ♀ (TAMU).
</paragraph>
</caption>
<paragraph id="8BDC36E4FFD8FFE7FF154301BD063B71" blockId="38.[189,1398,265,331]" pageId="38" pageNumber="39">
<collectingCountry id="F3747674FFD8FFE7FF154301BE683B1E" box="[189,282,266,292]" name="Panama" pageId="38" pageNumber="39">Panama</collectingCountry>
and northern
<collectingCountry id="F3747674FFD8FFE7FE624302BD4C3B19" box="[458,574,265,291]" name="Colombia" pageId="38" pageNumber="39">Colombia</collectingCountry>
are needed to be revised in order to determine if
<taxonomicName id="4C634D67FFD8FFE7FB054301BA323B19" authorityName="Balthasar" authorityYear="1939" baseAuthorityName="Balthasar" baseAuthorityYear="1939" box="[1197,1344,265,291]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B917EAF6FFD8FFE7FB054301BA323B19" box="[1197,1344,265,291]" italics="true" pageId="38" pageNumber="39">P. olsoufieffi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C634D67FFD8FFE7FF154339BE173B71" authority="Moctezuma &amp; Halffter, 2021" authorityName="Moctezuma &amp; Halffter" authorityYear="2021" box="[189,357,306,331]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B917EAF6FFD8FFE7FF154339BE173B71" box="[189,357,306,331]" italics="true" pageId="38" pageNumber="39">P. panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224578DFFD8FFE7FEC4433ABEB43B71" box="[364,454,305,331]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
are sympatric.
</paragraph>
</subSubSection>
</treatment>
</document>
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