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<mods:title id="1371763A8F9AAC6F09491B5F675E9B99">An annotated catalogue and bibliography of the taxonomy, synonymy and distribution of the Recent Vetigastropoda of South Africa (Mollusca)</mods:title>
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<treatment id="03CA87EF70718956F99FFCE24B156C64" ID-DOI="http://doi.org/10.5281/zenodo.5630984" ID-GBIF-Taxon="119646303" ID-Zenodo-Dep="5630984" LSID="urn:lsid:plazi:treatment:03CA87EF70718956F99FFCE24B156C64" httpUri="http://treatment.plazi.org/id/03CA87EF70718956F99FFCE24B156C64" lastPageNumber="84" pageId="83" pageNumber="84" scope_order="VETIGASTROPODA" scope_phylum="Mollusca" scope_subClass="VETIGASTROPODA">
<subSubSection id="C379657270718956F99FFCE24DD56A59" box="[151,655,854,879]" pageId="83" pageNumber="84" type="nomenclature">
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<taxonomicName id="4C634D7A70718956F99FFCE24DD56A59" authority="H. Adams, 1873" authorityName="H. Adams" authorityYear="1873" box="[151,655,854,879]" class="Gastropoda" family="Chilodontidae" genus="Pagodatrochus" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="variabilis">
<emphasis id="B917EAEB70718956F99FFCE24DD56A59" bold="true" box="[151,655,854,879]" pageId="83" pageNumber="84">
<emphasis id="B917EAEB70718956F99FFCE24EED6A59" bold="true" box="[151,439,854,879]" italics="true" pageId="83" pageNumber="84">Pagodatrochus variabilis</emphasis>
(
<bibRefCitation id="EFF24B0870718956F8CFFCE24DDC6A59" author="Adams" box="[455,646,854,879]" pageId="83" pageNumber="84" refString="Adams, H. (1873) Description of seventeen new species of land and marine shells. Proceedings of the Zoological Society of London, 41 (1873), 205 - 209." type="journal article" year="1873">H. Adams, 1873</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C379657270718956F99FFCCE4C836ADA" pageId="83" pageNumber="84" type="reference_group">
<paragraph id="8BDC36F970718956F99FFCCE4D746A99" blockId="83.[151,1436,854,1005]" pageId="83" pageNumber="84">
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<taxonomicName id="4C634D7A70718956F99FFCCE4D6D6AA6" authority="H. Adams, 1873: 207" authorityName="H. Adams" authorityPageNumber="207" authorityYear="1873" box="[151,567,890,912]" class="Gastropoda" family="Solariellidae" genus="Minolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="variabilis">
<emphasis id="B917EAEB70718956F99FFCCE4E0A6AA6" box="[151,336,890,912]" italics="true" pageId="83" pageNumber="84">Minolia variabilis</emphasis>
<treatmentCitation id="0AC210E870718956F85FFCCF4D6D6AA6" author="Adams" box="[343,567,890,912]" page="207" pageId="83" pageNumber="84" year="1873">
<bibRefCitation id="EFF24B0870718956F85FFCCF4D596AA6" author="Adams" box="[343,515,890,912]" pageId="83" pageNumber="84" refString="Adams, H. (1873) Description of seventeen new species of land and marine shells. Proceedings of the Zoological Society of London, 41 (1873), 205 - 209." type="journal article" year="1873">H. Adams, 1873</bibRefCitation>
: 207
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</taxonomicName>
, pl. 23, fig. 10.
<typeStatus id="54D8885B70718956FBD5FCCF4C556AA6" box="[733,783,891,912]" pageId="83" pageNumber="84">Type</typeStatus>
loc.: Persian Gulf;
<typeStatus id="54D8885B70718956FAD2FCCE4B696AA6" box="[986,1075,890,912]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
in NHM (NHMUK 1878.1.28.96), figured by
<bibRefCitation id="EFF24B0870718956F83EFC2D4E8F6A99" author="Herbert" box="[310,469,921,943]" pageId="83" pageNumber="84" refString="Herbert, D. G. (1989 b) Pagodatrochus, a new genus for Minolia variabilis H. Adams, 1873 (Gastropoda: Trochidae). Journal of Molluscan Studies, 55, 365 - 372. http: // dx. doi. org / 10.1093 / mollus / 55.3.365" type="journal article" year="1989" yearSuffix="b">Herbert (1989b</bibRefCitation>
: fig. 1e).
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</paragraph>
<paragraph id="8BDC36F970718956F99FFC0C4C836ADA" blockId="83.[151,1436,854,1005]" pageId="83" pageNumber="84">
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<taxonomicName id="4C634D7A70718956F99FFC0C4EC36AF8" box="[151,409,952,974]" class="Gastropoda" family="Chilodontidae" genus="Pagodatrochus" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="variabilis">
<emphasis id="B917EAEB70718956F99FFC0C4EC36AF8" box="[151,409,952,974]" italics="true" pageId="83" pageNumber="84">Pagodatrochus variabilis</emphasis>
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<treatmentCitation id="0AC210E870718956F8BBFC0C4DDF6AF8" author="Herbert" box="[435,645,952,974]" page="366" pageId="83" pageNumber="84" year="1989">
<bibRefCitation id="EFF24B0870718956F8BBFC0C4D0B6AF8" author="Herbert" box="[435,593,952,974]" pageId="83" pageNumber="84" refString="Herbert, D. G. (1989 b) Pagodatrochus, a new genus for Minolia variabilis H. Adams, 1873 (Gastropoda: Trochidae). Journal of Molluscan Studies, 55, 365 - 372. http: // dx. doi. org / 10.1093 / mollus / 55.3.365" type="journal article" year="1989" yearSuffix="b">Herbert, 1989b</bibRefCitation>
: 366
</treatmentCitation>
, figs 1af, 2, 3, 4ac, 5 (detailed synonymy).
<treatmentCitation id="0AC210E870718956FD68FC0C4A1C6AF8" author="Bosch" box="[1120,1350,952,974]" page="37" pageId="83" pageNumber="84" year="1995">
<bibRefCitation id="EFF24B0870718956FD68FC0C4A456AF8" author="Bosch" box="[1120,1311,952,974]" pageId="83" pageNumber="84" refString="Bosch, D. T, Dance, S. P., Moolenbeek, R. G. &amp; Oliver, P. G. (1995) Seashells of Eastern Arabia. Motivate Publishing, Abu Dhabi, Dubai and London, 296 pp." type="book" year="1995">
Bosch
<emphasis id="B917EAEB70718956FDAFFC0D4B8F6AF8" box="[1191,1237,952,974]" italics="true" pageId="83" pageNumber="84">et al</emphasis>
., 1995
</bibRefCitation>
: 37
</treatmentCitation>
, fig. 52.
<treatmentCitation id="0AC210E870718956F9CFFC624EE36ADA" author="Zuschin" box="[199,441,982,1005]" page="99" pageId="83" pageNumber="84" year="2009">
<bibRefCitation id="EFF24B0870718956F9CFFC624ECE6ADA" author="Zuschin" box="[199,404,982,1005]" pageId="83" pageNumber="84" refString="Zuschin, M., Janssen, R. &amp; Baal, C. (2009) Gastropods and their habitats from the northern Red Sea (Egypt: Safaga). Part 1: Patellogastropoda, Vetigastropoda and Cycloneritimorpha. Annalen des Naturhistorischen Museums in Wien, 111 (A), 73 - 158." type="journal article" year="2009">
Zuschin
<emphasis id="B917EAEB70718956F817FC6C4E116ADA" box="[287,331,982,1005]" italics="true" pageId="83" pageNumber="84">et al</emphasis>
., 2009
</bibRefCitation>
: 99
</treatmentCitation>
, pl. 9, figs 14.
<treatmentCitation id="0AC210E870718956FB54FC624C436ADA" author="Bandel" box="[604,793,982,1004]" page="461" pageId="83" pageNumber="84" year="2010">
<bibRefCitation id="EFF24B0870718956FB54FC624DB26ADA" author="Bandel" box="[604,744,982,1004]" pageId="83" pageNumber="84" refString="Bandel, K. (2010) Relationships of the Triassic Eucycloidea Koken, 1897 (Mollusca, Gastropoda) to modern genera such as Pagodatrochus, Calliotropis and Euchelus, based on morphology of the early shell. Bulletin of Geosciences, 85, 435 - 486. http: // dx. doi. org / 10.3140 / bull. geosci. 1208" type="journal article" year="2010">Bandel, 2010</bibRefCitation>
: 461
</treatmentCitation>
, fig. 10.
<bibRefCitation id="EFF24B0870718956FA7AFC634C8F6ADA" author="Jay" box="[882,981,982,1004]" pageId="83" pageNumber="84" refString="Jay, M. (2014) Mollusques de l'ile de la Reunion. Version 24. Available from: http: // vieoceane. free. fr / mollusques / intro _ mollusque. htm (accessed 15 April 2015)" type="journal article" year="2014">Jay, 2014</bibRefCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C379657270718956F99FFBAD4B156C64" pageId="83" pageNumber="84" type="distribution">
<paragraph id="8BDC36F970718956F99FFBAD4EB56D4C" blockId="83.[151,1436,1049,1362]" pageId="83" pageNumber="84">
<emphasis id="B917EAEB70718956F99FFBAD4E6A6D04" bold="true" box="[151,304,1049,1074]" pageId="83" pageNumber="84">Distribution.</emphasis>
Western Indian Ocean, from the Persian Gulf and Red Sea through East Africa, the
<collectingCountry id="F374766970718956FC0BFBAE4A306D04" box="[1283,1386,1050,1074]" name="Comoros" pageId="83" pageNumber="84">Comoros</collectingCountry>
and Mascarene Islands to the KwaZulu-Natal south coast (off Park Rynie); local material
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, but mostly less than
<quantity id="4C9B9B1C70718956F9C6FBD64E4F6D4F" box="[206,277,1122,1146]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="83" pageNumber="84" unit="m" value="100.0">100 m</quantity>
(living
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).
</paragraph>
<paragraph id="8BDC36F970718956F9CFFB314B156C64" blockId="83.[151,1436,1049,1362]" pageId="83" pageNumber="84">
<emphasis id="B917EAEB70718956F9CFFB314E4B6DA8" bold="true" box="[199,273,1157,1182]" pageId="83" pageNumber="84">Notes.</emphasis>
The phylogenetic relationships of this genus are unclear.
<bibRefCitation id="EFF24B0870718956FAACFB314B016DA8" author="Herbert" box="[932,1115,1157,1182]" pageId="83" pageNumber="84" refString="Herbert, D. G. (1989 b) Pagodatrochus, a new genus for Minolia variabilis H. Adams, 1873 (Gastropoda: Trochidae). Journal of Molluscan Studies, 55, 365 - 372. http: // dx. doi. org / 10.1093 / mollus / 55.3.365" type="journal article" year="1989" yearSuffix="b">Herbert (1989b)</bibRefCitation>
tentatively referred it to the
<taxonomicName id="4C634D7A70718956F99FFB1D4E4E6DF4" box="[151,276,1193,1218]" class="Gastropoda" family="Trochidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="83" pageNumber="84" phylum="Mollusca" rank="subFamily" subFamily="Gibbulinae">Gibbulinae</taxonomicName>
[=
<taxonomicName id="4C634D7A70718956F838FB1D4E8C6DF4" box="[304,470,1193,1218]" class="Gastropoda" family="Trochidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="83" pageNumber="84" phylum="Mollusca" rank="subFamily" subFamily="Cantharidinae">Cantharidinae</taxonomicName>
], but more recently
<bibRefCitation id="EFF24B0870718956FBBCFB1D4C0F6DF4" author="Bandel" box="[692,853,1193,1218]" pageId="83" pageNumber="84" refString="Bandel, K. (2010) Relationships of the Triassic Eucycloidea Koken, 1897 (Mollusca, Gastropoda) to modern genera such as Pagodatrochus, Calliotropis and Euchelus, based on morphology of the early shell. Bulletin of Geosciences, 85, 435 - 486. http: // dx. doi. org / 10.3140 / bull. geosci. 1208" type="journal article" year="2010">Bandel (2010)</bibRefCitation>
suggested that the genus was related to seguenzioid (eucycloid) taxa and created for it a new family, the Pagodatrochidae.
<bibRefCitation id="EFF24B0870718956FAD7FB794BD16DD0" author="Herbert" box="[991,1163,1229,1254]" pageId="83" pageNumber="84" refString="Herbert, D. G. (2012) A revision of the Chilodontidae (Gastropoda: Vetigastropoda: Seguenzioidea) of southern Africa and the south-western Indian Ocean. African Invertebrates, 53, 381 - 502. http: // dx. doi. org / 10.5733 / afin. 053.0209" type="journal article" year="2012">Herbert (2012)</bibRefCitation>
noted that while some characters of the shell and
<taxonomicName id="4C634D7A70718956F8D5FB454D796C3C" box="[477,547,1265,1290]" class="Gastropoda" family="Calliostomatidae" genus="Falsimargarita" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="radula">radula</taxonomicName>
of
<taxonomicName id="4C634D7A70718956FB5CFB454DA46C3C" box="[596,766,1265,1290]" class="Gastropoda" family="Chilodontidae" genus="Pagodatrochus" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="83" pageNumber="84" phylum="Mollusca" rank="genus">
<emphasis id="B917EAEB70718956FB5CFB454DA46C3C" box="[596,766,1265,1290]" italics="true" pageId="83" pageNumber="84">Pagodatrochus</emphasis>
</taxonomicName>
are indeed similar to those of chilodontid seguenzioids, features of the external anatomy are less consistent with such a relationship. In the absence of molecular data I consider it preferable to treat the genus as a vetigastropod taxon of uncertain affinity.
</paragraph>
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