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<mods:title id="8886F74FAE7911B7A10A7762E1C43F93">A new Epipleoneura Williamson, 1915 (Zygoptera, Coenagrionidae) from northern Brazil and notes on E. venezuelensis Rácenis, 1955</mods:title>
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<mods:affiliation id="17E3E7360C3F2559738A29E129331889">Facultad de Ciencias Naturales y Ciencias de la Salud, Universidad Nacional de la Patagonia “ San Juan Bosco ”, Chubut, Argentina. Programa de Pós-Graduação em Ecologia, Instituto de Ciências Biológicas (ICB), Laboratório de Ecologia e Conservação (LA- BECO), Universidade Federal do Pará (UFPA), Belém, Brazil. &amp; danianjos _ santos @ comahue-conicet. gob. ar; https: // orcid. org / 0000 - 0002 - 2889 - 5964</mods:affiliation>
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<emphasis id="B917EAB3FFF4FFC435D6F9EDFE7B2877" bold="true" box="[151,429,1554,1581]" italics="true" pageId="1" pageNumber="154">Epipleoneura lencionii</emphasis>
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Pessacq &amp; Anjos-Santos
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(
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, 
<figureCitation id="13582A24FFF4FFC4341DF9C9FEBA280B" box="[348,364,1590,1617]" captionStart="FIGURE 5" captionStartId="8.[152,255,1962,1986]" captionTargetBox="[419,1173,893,1939]" captionTargetId="figure-337@8.[415,1173,885,1939]" captionTargetPageId="8" captionText="FIGURE 5. Distribution map of Epipleoneura lencionii and E. venezuelensis. Indicated the hydrological basins:Amazon basin, Tocantins-Plata basin and Plata-Paraná-Paraguay basin." figureDoi="http://doi.org/10.5281/zenodo.7413442" httpUri="https://zenodo.org/record/7413442/files/figure.png" pageId="1" pageNumber="154">5</figureCitation>
, 
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)
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<emphasis id="B917EAB3FFF4FFC435D6F984FE6428C9" box="[151,434,1659,1683]" italics="true" pageId="1" pageNumber="154">Epipleoneura venezuelensis</emphasis>
: 
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, figs. 113a–b (illustration of male S10 and caudal appendages in lateral view, male epiproct in posterior view).
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AC-01, segment 4 (
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<specimenCount id="9D65FD28FFF4FFC4317CF8FBFBBE2945" box="[1085,1128,1796,1823]" pageId="1" pageNumber="154" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED72AE64FFF4FFC43136F8FBFB3B2944" box="[1143,1261,1796,1822]" pageId="1" pageNumber="154">LABECO</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B0B3CFCFFF4FFC43040F8FAFA4E291E" ID-GBIF-Occurrence="3988016307" collectionCode="LABECO" country="Brazil" latitude="-10.071639" location="Fazenda Experimental Catuaba" longLatPrecision="1" longitude="-67.61503" municipality="Ponto" pageId="1" pageNumber="154" specimenCount="1" specimenCount-male="1" stateProvince="Acre" typeStatus="paratype">
<collectingMunicipality id="6BB8ACDBFFF4FFC43040F8FAFA942945" box="[1281,1346,1797,1823]" pageId="1" pageNumber="154">Ponto</collectingMunicipality>
AC-01, segment 8; 
<specimenCount id="9D65FD28FFF4FFC43454F8D7FEEB2919" box="[277,317,1832,1859]" pageId="1" pageNumber="154" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED72AE64FFF4FFC4340BF8D7FE162918" box="[330,448,1832,1858]" pageId="1" pageNumber="154">LABECO</collectionCode>
), 
<location id="8EBC607AFFF4FFC43493F8D7FCE62918" LSID="urn:lsid:plazi:treatment:03CA87B7FFF4FFC035D6F9EDFA9529FD:8EBC607AFFF4FFC43493F8D7FCE62918" box="[466,816,1832,1858]" country="Brazil" latitude="-10.071639" longLatPrecision="1" longitude="-67.61503" municipality="Ponto" name="Fazenda Experimental Catuaba" pageId="1" pageNumber="154" stateProvince="Acre">Fazenda Experimental Catuaba</location>
, 
<location id="8EBC607AFFF4FFC43678F8D6FCAC2919" LSID="urn:lsid:plazi:treatment:03CA87B7FFF4FFC035D6F9EDFA9529FD:8EBC607AFFF4FFC43678F8D6FCAC2919" box="[825,890,1833,1859]" country="Brazil" latitude="-10.071639" longLatPrecision="1" longitude="-67.61503" municipality="Ponto" name="Ponto" pageId="1" pageNumber="154" stateProvince="Acre">Ponto</location>
AC-04, segment 1 (
<geoCoordinate id="EE575066FFF4FFC43115F8D6FB342918" box="[1108,1250,1832,1860]" degrees="10" direction="south" minutes="4" orientation="latitude" pageId="1" pageNumber="154" precision="1" seconds="17.9" value="-10.071639">10º4′17.9″ S</geoCoordinate>
; 
<geoCoordinate id="EE575066FFF4FFC431ACF8D6FA46291E" box="[1261,1424,1833,1860]" degrees="67" direction="west" minutes="36" orientation="longitude" pageId="1" pageNumber="154" precision="1" seconds="54.1" value="-67.61503">67º36′54.1″W</geoCoordinate>
)
</materialsCitation>
; 
<materialsCitation id="3B0B3CFCFFF4FFC435D6F8B3FDCC293C" ID-GBIF-Occurrence="3988016310" box="[151,538,1868,1895]" collectionCode="LABECO" country="Brazil" pageId="1" pageNumber="154" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Acre" typeStatus="paratype">
<specimenCount id="9D65FD28FFF4FFC435D6F8B3FF13293D" box="[151,197,1868,1895]" pageId="1" pageNumber="154" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D65FD28FFF4FFC43592F8B3FF29293D" box="[211,255,1868,1895]" pageId="1" pageNumber="154" type="female">1 ♀</specimenCount>
(in tandem) (
<collectionCode id="ED72AE64FFF4FFC434DDF8B3FDC4293C" box="[412,530,1868,1894]" pageId="1" pageNumber="154">LABECO</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B0B3CFCFFF4FFC43767F8B2FEA229F5" ID-GBIF-Occurrence="3988016302" collectingDate="2017-09-20" country="Brazil" county="Assis County" latitude="-10.069" location="Ponto" longLatPrecision="15" longitude="-67.604996" pageId="1" pageNumber="154" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Acre" typeStatus="paratype">
same as previous; 
<specimenCount id="9D65FD28FFF4FFC437BBF8B3FCFE293D" box="[762,808,1868,1895]" pageId="1" pageNumber="154" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D65FD28FFF4FFC43677F8B3FCB7293D" box="[822,865,1868,1895]" pageId="1" pageNumber="154" type="female">1 ♀</specimenCount>
(
<emphasis id="B917EAB3FFF4FFC43635F8B2FC5C293C" box="[884,906,1869,1894]" italics="true" pageId="1" pageNumber="154">in</emphasis>
tandem), same as previous except: Ponto AC-09, segment 5 (
<geoCoordinate id="EE575066FFF4FFC43404F88EFE1629D6" box="[325,448,1905,1932]" degrees="10" direction="south" minutes="4" orientation="latitude" pageId="1" pageNumber="154" precision="1" seconds="8.4" value="-10.069">10º4′8.4″S</geoCoordinate>
; 
<geoCoordinate id="EE575066FFF4FFC4348DF88EFD8F29D6" box="[460,601,1905,1932]" degrees="67" direction="west" minutes="36" orientation="longitude" pageId="1" pageNumber="154" precision="15" seconds="18" value="-67.604996">67º36′18″W</geoCoordinate>
), 
<date id="FFDD1061FFF4FFC4372CF88FFD3E29D0" box="[621,744,1904,1930]" pageId="1" pageNumber="154" value="2017-09-20">
<collectingDate id="EF99E989FFF4FFC4372CF88FFD3E29D0" box="[621,744,1904,1930]" pageId="1" pageNumber="154" value="2017-09-20">20.ix.2017</collectingDate>
</date>
; 
<specimenCount id="9D65FD28FFF4FFC437B5F88FFCCE29D0" box="[756,792,1904,1930]" pageId="1" pageNumber="154" type="male">1♂</specimenCount>
, 
<specimenCount id="9D65FD28FFF4FFC43667F88FFC9F29D0" box="[806,841,1904,1930]" pageId="1" pageNumber="154" type="female">1♀</specimenCount>
(in tandem), same as previous except: 
<collectingCounty id="62BD4E2DFFF4FFC43041F88FFA4F29D0" box="[1280,1433,1904,1930]" pageId="1" pageNumber="154">Assis County</collectingCounty>
, small stream, 
<location id="8EBC607AFFF4FFC43472F86AFEA229F5" LSID="urn:lsid:plazi:treatment:03CA87B7FFF4FFC035D6F9EDFA9529FD:8EBC607AFFF4FFC43472F86AFEA229F5" box="[307,372,1941,1967]" country="Brazil" county="Assis County" latitude="-10.069" longLatPrecision="15" longitude="-67.604996" name="Ponto" pageId="1" pageNumber="154" stateProvince="Acre">Ponto</location>
</materialsCitation>
<materialsCitation id="3B0B3CFCFFF4FFC4343DF86BFA9229F5" ID-GBIF-Occurrence="3988016306" box="[380,1348,1940,1968]" collectingDate="2022-08-21" collectionCode="PCPE" country="Brazil" latitude="-10.903055" location="Ponto" longLatPrecision="21" longitude="-69.52112" pageId="1" pageNumber="154" specimenCode="PCPE5" specimenCount="7" specimenCount-male="7" stateProvince="Acre" typeStatus="paratype">
<specimenCode id="DBC59EDAFFF4FFC4343DF86BFE1B29F5" box="[380,461,1940,1967]" collectionCode="PCPE" pageId="1" pageNumber="154">PCPE5</specimenCode>
(
<geoCoordinate id="EE575066FFF4FFC4349DF86AFD8B29EA" box="[476,605,1941,1968]" degrees="10" direction="south" minutes="54" orientation="latitude" pageId="1" pageNumber="154" precision="15" seconds="11" value="-10.903055">10º54′11″S</geoCoordinate>
; 
<geoCoordinate id="EE575066FFF4FFC43728F86AFD2329EA" box="[617,757,1941,1968]" degrees="69" direction="west" minutes="31" orientation="longitude" pageId="1" pageNumber="154" precision="15" seconds="16" value="-69.52112">69º31′16″W</geoCoordinate>
), 
<date id="FFDD1061FFF4FFC43648F86BFC4229F4" box="[777,916,1940,1966]" pageId="1" pageNumber="154" value="2022-08-21">
<collectingDate id="EF99E989FFF4FFC43648F86BFC4229F4" box="[777,916,1940,1966]" pageId="1" pageNumber="154" value="2022-08-21">21.viii.2022</collectingDate>
</date>
; 
<specimenCount id="9D65FD28FFF4FFC436DEF86AFC0829F5" box="[927,990,1941,1967]" pageId="1" pageNumber="154" type="male">7 ♂♂</specimenCount>
same as previous except: 
<location id="8EBC607AFFF4FFC43042F86AFA9229F5" LSID="urn:lsid:plazi:treatment:03CA87B7FFF4FFC035D6F9EDFA9529FD:8EBC607AFFF4FFC43042F86AFA9229F5" box="[1283,1348,1941,1967]" country="Brazil" latitude="-10.903055" longLatPrecision="21" longitude="-69.52112" name="Ponto" pageId="1" pageNumber="154" stateProvince="Acre">Ponto</location>
</materialsCitation>
<materialsCitation id="3B0B3CFCFFF4FFC4300AF86BFC9D2988" ID-GBIF-Occurrence="3988016304" collectingDate="2022-08-16" collectionCode="PCPE" collectorName="C. C. Mendoza" country="Brazil" latitude="-10.889861" longLatPrecision="15" longitude="-69.61083" pageId="1" pageNumber="154" specimenCode="PCPE9" specimenCount="1" stateProvince="Acre" typeStatus="paratype">
<specimenCode id="DBC59EDAFFF4FFC4300AF86BFA4A29F4" box="[1355,1436,1940,1966]" collectionCode="PCPE" pageId="1" pageNumber="154">PCPE9</specimenCode>
(
<geoCoordinate id="EE575066FFF4FFC435DEF846FEE1298E" box="[159,311,1977,2004]" degrees="10" direction="south" minutes="53" orientation="latitude" pageId="1" pageNumber="154" precision="1" seconds="23.5" value="-10.889861">
10º53′23.5″ 
<collectionCode id="ED72AE64FFF4FFC43468F846FEE1298E" box="[297,311,1977,2004]" pageId="1" pageNumber="154">S</collectionCode>
</geoCoordinate>
; 
<geoCoordinate id="EE575066FFF4FFC43403F846FE06298E" box="[322,464,1977,2004]" degrees="69" direction="west" minutes="36" orientation="longitude" pageId="1" pageNumber="154" precision="15" seconds="39" value="-69.61083">
69º36′39″ 
<collectionCode id="ED72AE64FFF4FFC434F2F846FE06298E" box="[435,464,1977,2004]" pageId="1" pageNumber="154">W</collectionCode>
</geoCoordinate>
), 
<date id="FFDD1061FFF4FFC434A2F847FDBA2988" box="[483,620,1976,2002]" pageId="1" pageNumber="154" value="2022-08-16">
<collectingDate id="EF99E989FFF4FFC434A2F847FDBA2988" box="[483,620,1976,2002]" pageId="1" pageNumber="154" value="2022-08-16">16.viii.2022</collectingDate>
</date>
, 
<collectorName id="26965377FFF4FFC43739F847FCCC2988" box="[632,794,1976,2002]" pageId="1" pageNumber="154">C.C. Mendoza</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586FF69FD832F1B" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586FF69FDED2EEA" bold="true" box="[199,571,150,176]" pageId="2" pageNumber="155">Additional material examined.</emphasis>
(subsequently lost in 
<collectionCode id="ED72AE64FFF7FFC7367FFF68FC5F2EEB" box="[830,905,151,177]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="155">MNRJ</collectionCode>
fire). 
<materialsCitation id="3B0B3CFCFFF7FFC73694FF68FB622EA3" ID-GBIF-Occurrence="3988016312" collectingDate="1982-07-14" collectorName="Ruis" country="Brazil" latitude="-19.688055" location="Nova Uniao" longLatPrecision="21" longitude="-43.583332" municipality="Guburana" pageId="2" pageNumber="155" specimenCount="7" specimenCount-male="7" stateProvince="Minas Gerais">
<specimenCount id="9D65FD28FFF7FFC73694FF68FBCF2EEB" box="[981,1049,151,177]" pageId="2" pageNumber="155" type="male">7 ♂♂</specimenCount>
, 
<emphasis id="B917EAB3FFF7FFC73168FF69FBA42EEA" bold="true" box="[1065,1138,150,176]" pageId="2" pageNumber="155">
<collectingCountry id="F3747631FFF7FFC73168FF69FBA42EEA" box="[1065,1138,150,176]" name="Brazil" pageId="2" pageNumber="155">Brazil</collectingCountry>
</emphasis>
, 
<collectingRegion id="49A7F843FFF7FFC731C2FF68FAB42EEB" box="[1155,1378,151,177]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="155">Minas Gerais State</collectingRegion>
, 
<collectingMunicipality id="6BB8ACDBFFF7FFC73030FF68FF302E8F" pageId="2" pageNumber="155">Guburana</collectingMunicipality>
, 
<quantity id="4C9B9B44FFF7FFC735B5FF45FEE62E8F" box="[244,304,186,213]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="155" unit="km" value="5.0">5 km</quantity>
from 
<location id="8EBC607AFFF7FFC73438FF44FDD52E8F" LSID="urn:lsid:plazi:treatment:03CA87B7FFF4FFC035D6F9EDFA9529FD:8EBC607AFFF7FFC73438FF44FDD52E8F" box="[377,515,187,213]" country="Brazil" latitude="-19.688055" longLatPrecision="21" longitude="-43.583332" municipality="Guburana" name="Nova Uniao" pageId="2" pageNumber="155" stateProvince="Minas Gerais">Nova União</location>
(about 
<geoCoordinate id="EE575066FFF7FFC7371FFF43FD092E8D" box="[606,735,188,215]" degrees="19" direction="south" minutes="41" orientation="latitude" pageId="2" pageNumber="155" precision="15" seconds="17" value="-19.688055">19º41′17″S</geoCoordinate>
, 
<geoCoordinate id="EE575066FFF7FFC737ACFF43FCAD2E8D" box="[749,891,188,215]" degrees="43" direction="west" minutes="35" orientation="longitude" pageId="2" pageNumber="155" precision="15" seconds="00" value="-43.583332">43º35′00″W</geoCoordinate>
), 
<date id="FFDD1061FFF7FFC736D1FF44FBCA2E8F" box="[912,1052,187,213]" pageId="2" pageNumber="155" value="1982-07-14">
<collectingDate id="EF99E989FFF7FFC736D1FF44FBCA2E8F" box="[912,1052,187,213]" pageId="2" pageNumber="155" value="1982-07-14">14. vii.1982</collectingDate>
</date>
, 
<collectorName id="26965377FFF7FFC7316BFF44FB8B2E8F" box="[1066,1117,187,213]" pageId="2" pageNumber="155">Ruis</collectorName>
leg. 
<collectorName id="26965377FFF7FFC731D9FF44FB6A2E8F" box="[1176,1212,187,213]" pageId="2" pageNumber="155">All</collectorName>
material labeled as “ 
<taxonomicName id="4C634D22FFF7FFC735E5FF20FEEA2EA3" authorityName="Williamson" authorityYear="1915" box="[164,316,223,249]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAB3FFF7FFC735E5FF20FEEA2EA3" box="[164,316,223,249]" italics="true" pageId="2" pageNumber="155">Epipleoneura</emphasis>
</taxonomicName>
, para estudar” (
<taxonomicName id="4C634D22FFF7FFC734B0FF20FD5F2EA3" authorityName="Williamson" authorityYear="1915" box="[497,649,223,249]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAB3FFF7FFC734B0FF20FD5F2EA3" box="[497,649,223,249]" italics="true" pageId="2" pageNumber="155">Epipleoneura</emphasis>
</taxonomicName>
to study), sentence included by 
<collectorName id="26965377FFF7FFC736B9FF20FB622EA3" box="[1016,1204,223,249]" pageId="2" pageNumber="155">Prof. J.M. Costa</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B0B3CFCFFF7FFC731FEFF20FD982F1B" ID-GBIF-Occurrence="3988016305" collectingDate="1983-12-23" collectorName="L. F. Netto" country="Brazil" county="Rio Ouro Preto" latitude="-19.688055" location="Rio da Anta" longLatPrecision="21" longitude="-43.583332" municipality="Nova Uniao" pageId="2" pageNumber="155" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D65FD28FFF7FFC731FEFF20FB3A2EA3" box="[1215,1260,223,249]" pageId="2" pageNumber="155" type="male">1 ♂</specimenCount>
, same as previous except 
<collectingCounty id="62BD4E2DFFF7FFC73455FEFCFE142F47" box="[276,450,259,285]" pageId="2" pageNumber="155">Rio Ouro Preto</collectingCounty>
, 
<collectingMunicipality id="6BB8ACDBFFF7FFC7348DFEFCFD812F47" box="[460,599,259,285]" pageId="2" pageNumber="155">Nova União</collectingMunicipality>
, 
<location id="8EBC607AFFF7FFC73723FEFCFD302F47" LSID="urn:lsid:plazi:treatment:03CA87B7FFF4FFC035D6F9EDFA9529FD:8EBC607AFFF7FFC73723FEFCFD302F47" box="[610,742,259,285]" country="Brazil" county="Rio Ouro Preto" latitude="-19.688055" longLatPrecision="21" longitude="-43.583332" municipality="Nova Uniao" name="Rio da Anta" pageId="2" pageNumber="155">
Rio 
<collectorName id="26965377FFF7FFC737D0FEFCFD302F47" box="[657,742,259,285]" pageId="2" pageNumber="155">da Anta</collectorName>
</location>
(about 
<geoCoordinate id="EE575066FFF7FFC7367AFEFBFC6A2F45" box="[827,956,260,287]" degrees="19" direction="south" minutes="41" orientation="latitude" pageId="2" pageNumber="155" precision="15" seconds="17" value="-19.688055">
19º41′17″ 
<collectionCode id="ED72AE64FFF7FFC736ECFEFBFC6A2F45" box="[941,956,260,287]" pageId="2" pageNumber="155">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="EE575066FFF7FFC73686FEFBFB832F45" box="[967,1109,260,287]" degrees="43" direction="west" minutes="35" orientation="longitude" pageId="2" pageNumber="155" precision="15" seconds="00" value="-43.583332">
43º35′00″ 
<collectionCode id="ED72AE64FFF7FFC73179FEFBFB832F45" box="[1080,1109,260,287]" pageId="2" pageNumber="155">W</collectionCode>
</geoCoordinate>
), 
<date id="FFDD1061FFF7FFC73126FEFCFB3F2F47" box="[1127,1257,259,285]" pageId="2" pageNumber="155" value="1983-12-23">
<collectingDate id="EF99E989FFF7FFC73126FEFCFB3F2F47" box="[1127,1257,259,285]" pageId="2" pageNumber="155" value="1983-12-23">23.xii.1983</collectingDate>
</date>
, 
<collectorName id="26965377FFF7FFC731B5FEFCFAB32F47" box="[1268,1381,259,285]" pageId="2" pageNumber="155">L.F. Netto</collectorName>
leg., 
<collectorName id="26965377FFF7FFC735D6FED8FF272F1B" box="[151,241,295,321]" pageId="2" pageNumber="155">Labeled</collectorName>
as 
<taxonomicName id="4C634D22FFF7FFC73456FED8FD982F1B" authorityName=": Lencioni" authorityYear="2005" box="[279,590,295,321]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="155" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B917EAB3FFF7FFC73456FED8FD982F1B" box="[279,590,295,321]" italics="true" pageId="2" pageNumber="155">Epipleoneura venezuelensis</emphasis>
</taxonomicName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C379652AFFF7FFC73586FEB4FC3C2FD3" pageId="2" pageNumber="155" type="etymology">
<paragraph id="8BDC36A1FFF7FFC73586FEB4FC3C2FD3" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586FEB4FE9E2F3F" bold="true" box="[199,328,331,357]" pageId="2" pageNumber="155">Etymology</emphasis>
. This species is named after Frederico A. A. Lencioni, a great motivator for the continuity of our studies with 
<taxonomicName id="4C634D22FFF7FFC73464FE90FE522FD3" authorityName="Fabricius" authorityYear="1793" box="[293,388,367,393]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="155" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
and the first to report the identification of this species.
</paragraph>
</subSubSection>
<subSubSection id="C379652AFFF7FFC63586FE6DFBD82F47" lastPageId="3" lastPageNumber="156" pageId="2" pageNumber="155" type="description">
<paragraph id="8BDC36A1FFF7FFC73586FE6DFDC82FF7" blockId="2.[151,1437,150,2013]" box="[199,542,402,429]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586FE6DFDC82FF7" bold="true" box="[199,542,402,429]" pageId="2" pageNumber="155">
Description of male 
<typeStatus id="54D88803FFF7FFC734F6FE6CFDC82FF7" box="[439,542,403,429]" pageId="2" pageNumber="155" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586FE48FB002C43" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586FE48FED22F8B" box="[199,260,439,465]" italics="true" pageId="2" pageNumber="155">Head</emphasis>
. Dorsally metallic green; antennifer black, with a dorsal carina; frons black, angulate; postclypeus black, anteclypeus yellow; labrum black, with its apex yellow; genae black; rear of head black, ventrally with a yellow band bordering the eyes; labium and maxilla yellow, sides of mandibles black, remaining yellow.
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586FDDCFA4B2D63" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586FDDCFEF62C67" box="[199,288,547,573]" italics="true" pageId="2" pageNumber="155">Thorax.</emphasis>
Prothorax black. Mesepisternum metallic green, mesepimeron and metepisternum black with metallic green reflections, metepimeron black with yellow antero-ventral and posterior metallic green areas; pterothoracic venter yellow; coxae and tibia flexor side brown; trochanters, tibia extensor side and basal femur light yellow, remaining femur black; tarsus yellow with black apex. Wings hyaline, venation black; MP ending 0.5 cell distally from the vein descending from subnodus; IR2 and RP3 separated by a short crossvein one cell posterior to their origin; divergence of RP-RA (arculus) at Ax 2, in lefts Hw, distal to it in remaining wings; RP2 beginning at Px 
<quantity id="4C9B9B44FFF7FFC73033FD29FA4A2CAB" box="[1394,1436,726,753]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="155" unit="in" value="5.0">5 in</quantity>
Fw, between Px 3 and 4 (closer to 4) in Hw; IR1 beginning at Px 
<quantity id="4C9B9B44FFF7FFC7363BFD04FC732D4F" box="[890,933,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="2" pageNumber="155" unit="in" value="8.0">8 in</quantity>
left Fw and Px 
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right Fw, at Px 
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Hw; pt dark brown, about 2/3 underlying cell; 12 Px in Fw, 
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Hw; one additional intercalary between RP1 and RP2.
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586FCBCFC062D07" blockId="2.[151,1437,150,2013]" box="[199,976,835,861]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586FCBCFEEF2D07" box="[199,313,835,861]" italics="true" pageId="2" pageNumber="155">Abdomen.</emphasis>
Black, ventrally dark brown. S1–3 dorsally metallic green.
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586FC98FC022DFF" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586FC98FEBD2DDB" box="[199,363,871,897]" italics="true" pageId="2" pageNumber="155">Genital ligula.</emphasis>
(
<figureCitation id="13582A24FFF7FFC7343BFC98FE272DDB" box="[378,497,871,897]" captionStart="FIGURE 1" captionStartId="3.[152,255,1938,1962]" captionTargetBox="[188,1398,537,1898]" captionTargetId="figure-206@3.[176,1410,522,1911]" captionTargetPageId="3" captionText="FIGURE 1. Epipleoneura lencionii sp. nov. holotype, male, Minas Gerais State, Brazil. (a) caudal appendages, posterior view; (b) caudal appendages, lateral view; (c) genital ligula, ectal view; (d) genital ligula, lateral view. Epipleoneura venezuelensis, male, Parque Nacional Predelta, Entre Ríos Province, Argentina: (e) caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7413434" httpUri="https://zenodo.org/record/7413434/files/figure.png" pageId="2" pageNumber="155">Figs. 1c–d</figureCitation>
) lacking internal fold, with a pair of latero-posterior pedunculate processes, apex of S3 with a wide V cleft and lateral corners triangular and directed laterally.
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586FC50FBEE2A27" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586FC50FE702D93" box="[199,422,943,969]" italics="true" pageId="2" pageNumber="155">Caudal appendages</emphasis>
. Cercus (
<figureCitation id="13582A24FFF7FFC7374EFC50FD532D93" box="[527,645,943,969]" captionStart="FIGURE 1" captionStartId="3.[152,255,1938,1962]" captionTargetBox="[188,1398,537,1898]" captionTargetId="figure-206@3.[176,1410,522,1911]" captionTargetPageId="3" captionText="FIGURE 1. Epipleoneura lencionii sp. nov. holotype, male, Minas Gerais State, Brazil. (a) caudal appendages, posterior view; (b) caudal appendages, lateral view; (c) genital ligula, ectal view; (d) genital ligula, lateral view. Epipleoneura venezuelensis, male, Parque Nacional Predelta, Entre Ríos Province, Argentina: (e) caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7413434" httpUri="https://zenodo.org/record/7413434/files/figure.png" pageId="2" pageNumber="155">Figs. 1a–b</figureCitation>
) is shorter than S10, with a dorsal and a ventral branch, the former well developed, directed postero-dorsally, approximately rounded in cross section and armed with an apical hook. Ventral branch small, developed as a projection of the cercus base. The cercus possesses a well-developed inner-basal branch directed meso-ventrally, and its apex is directed posteriorly (
<figureCitation id="13582A24FFF7FFC736D9FBE4FC3F2A6F" box="[920,1001,1051,1077]" captionStart="FIGURE 1" captionStartId="3.[152,255,1938,1962]" captionTargetBox="[188,1398,537,1898]" captionTargetId="figure-206@3.[176,1410,522,1911]" captionTargetPageId="3" captionText="FIGURE 1. Epipleoneura lencionii sp. nov. holotype, male, Minas Gerais State, Brazil. (a) caudal appendages, posterior view; (b) caudal appendages, lateral view; (c) genital ligula, ectal view; (d) genital ligula, lateral view. Epipleoneura venezuelensis, male, Parque Nacional Predelta, Entre Ríos Province, Argentina: (e) caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7413434" httpUri="https://zenodo.org/record/7413434/files/figure.png" pageId="2" pageNumber="155">Fig. 1a</figureCitation>
). Paraproct conical. Epiproct (
<figureCitation id="13582A24FFF7FFC73002FBE4FA442A6F" box="[1347,1426,1051,1077]" captionStart="FIGURE 1" captionStartId="3.[152,255,1938,1962]" captionTargetBox="[188,1398,537,1898]" captionTargetId="figure-206@3.[176,1410,522,1911]" captionTargetPageId="3" captionText="FIGURE 1. Epipleoneura lencionii sp. nov. holotype, male, Minas Gerais State, Brazil. (a) caudal appendages, posterior view; (b) caudal appendages, lateral view; (c) genital ligula, ectal view; (d) genital ligula, lateral view. Epipleoneura venezuelensis, male, Parque Nacional Predelta, Entre Ríos Province, Argentina: (e) caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7413434" httpUri="https://zenodo.org/record/7413434/files/figure.png" pageId="2" pageNumber="155">Fig. 1a</figureCitation>
) with two short, rounded, apical lobes and two lateral lobes directed outwards, approximately parallel to the epiproct transversal axis and approximately in the same plane as the surface of the epiproct.
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586FB78FC1B2AFB" blockId="2.[151,1437,150,2013]" box="[199,973,1158,1185]" pageId="2" pageNumber="155">Measurements. Total length 35; abdomen length 30; Fw: 19; Hw: 18.</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586FB55FE102A9E" blockId="2.[151,1437,150,2013]" box="[199,454,1194,1220]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586FB55FE102A9E" bold="true" box="[199,454,1194,1220]" pageId="2" pageNumber="155">Description of female</emphasis>
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586FB30FAFE2B6B" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586FB30FED22AB3" box="[199,260,1231,1257]" italics="true" pageId="2" pageNumber="155">Head</emphasis>
. Dorsally metallic green, antennifer dorsum black, front yellow; antefrons yellow with dorsal margin and a spot under each antennifer black; postclypeus black, anteclypeus light yellow; labrum yellow, with a basal median black spot; genae yellow; distal half of head venter yellow, remaining black; labrum and maxilla yellow.
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586FAC4FAC12853" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586FAC4FECE2B0F" box="[199,280,1339,1365]" italics="true" pageId="2" pageNumber="155">Thorax</emphasis>
. Prothorax dorsal surface metallic green, ventrally bordered by a yellow stripe, remaining lateral surface light brown; posterior lobe (
<figureCitation id="13582A24FFF7FFC7374AFAA0FD992B23" box="[523,591,1375,1401]" captionStart="FIGURE 2" captionStartId="4.[152,255,1768,1792]" captionTargetBox="[165,1424,182,1737]" captionTargetId="figure-18@4.[154,1432,181,1743]" captionTargetPageId="4" captionText="FIGURE 2. Epipleoneura lencionii sp. nov. female, Minas Gerais State, Brazil. (a) prothorax posterior lobe, dorsal view, photograph (left), and drawing (right); (b) prothorax posterior lobe, postero-lateral view, photograph, and drawing; (c) prothorax posterior lobe, lateral view, photograph, and drawing. PP: posterior process." figureDoi="http://doi.org/10.5281/zenodo.7413436" httpUri="https://zenodo.org/record/7413436/files/figure.png" pageId="2" pageNumber="155">Fig. 2</figureCitation>
) with an erected medial lobe, its anterior margin with a shallow concavity, its posterior margin straight, not surpassing the level of the lateral lobes; there is a rounded basal posterior process (indicated PP in 
<figureCitation id="13582A24FFF7FFC73410FA58FE452B9B" box="[337,403,1447,1473]" captionStart="FIGURE 2" captionStartId="4.[152,255,1768,1792]" captionTargetBox="[165,1424,182,1737]" captionTargetId="figure-18@4.[154,1432,181,1743]" captionTargetPageId="4" captionText="FIGURE 2. Epipleoneura lencionii sp. nov. female, Minas Gerais State, Brazil. (a) prothorax posterior lobe, dorsal view, photograph (left), and drawing (right); (b) prothorax posterior lobe, postero-lateral view, photograph, and drawing; (c) prothorax posterior lobe, lateral view, photograph, and drawing. PP: posterior process." figureDoi="http://doi.org/10.5281/zenodo.7413436" httpUri="https://zenodo.org/record/7413436/files/figure.png" pageId="2" pageNumber="155">Fig. 2</figureCitation>
) on the posterior margin of the medial lobe. Mesothorax metallic green, except for a yellow stripe below the mesopleural suture; metathorax yellow, except for a brown stripe at the posterior half of the metopleural suture; pterothoracic venter yellow; legs yellow, except for the brown apex of femora and tibia.
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586F9ECFC09280B" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586F9ECFEEF2877" box="[199,313,1555,1581]" italics="true" pageId="2" pageNumber="155">Abdomen.</emphasis>
Dorsally black with metallic green reflections, remaining yellow; apex of S3–7 with black color extending ventrally, the base of S3–7 with yellow color extending dorsally.
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586F9A5FAA7295F" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="155">
<emphasis id="B917EAB3FFF7FFC73586F9A5FE0F282F" bold="true" box="[199,473,1626,1653]" pageId="2" pageNumber="155">
Variation in 
<typeStatus id="54D88803FFF7FFC7341DF9A4FE02282F" box="[348,468,1627,1653]" pageId="2" pageNumber="155" type="paratype">paratypes</typeStatus>
.
</emphasis>
Divergence of RP-RA (arculus) distal to Ax 
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all specimens. Px: Fw 10 (62.5%), 11 (37.5%); Hw 9 (100%). RP2: at Px 3 (25%), 4 (25%), between 4 and 5 (25%), and 5 (25%) in Fw; at Px 
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Hw in all specimens. IR1: at Px 6 (25%), and 7 (75%) in Fw; at Px 6 (75%) or 7 (25%) in Hw. In 
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collected in Assis County (
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; 
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) in 2022, the lateral lobes of the epiproct are in the same plane as the surface of the epiproct, but they are directed latero-distally at an angle of about 45º with respect to the transversal axis.
</paragraph>
<paragraph id="8BDC36A1FFF7FFC73586F8F0FC3529CF" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="155">Measurements. (males n = 4, females n = 2): Male: total length 24.9–27.32 [mean 30.1; Standard Deviation (SD) 0.45]; abdomen length 24.9–27.32 [mean 30.1; SD 0.45]; Fw 18–19 [mean 18.2; SD 0.45]; Hw 17–17.5 [mean 17.1; SD 0.53]. Female: total length 24.9–27.32 [mean 30.1; SD 0.45]; abdomen length 24.9–27.32 [mean 30.1; SD 0.45]; Fw 18–19 [mean 18.2; SD 0.45]; Hw 17–17.5 [mean 17.1; SD 0.53].</paragraph>
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<emphasis id="B917EAB3FFF7FFC73586F861FDE929E2" bold="true" box="[199,575,1950,1976]" pageId="2" pageNumber="155">Variation in additional material</emphasis>
. Divergence of RP-RA (arculus) distal to Ax 
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all specimens. Venation: Px: Fw 10 (66.6%), 11 (45%), 12 (8,3%); Hw 9 (50%), 10 (50%). RP2: at Px 
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Fw in all specimens except for one in right wing at Px 4; at Px 
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Hw in all specimens except for 
<specimenCount id="9D65FD28FFF6FFC63604FF68FC3A2EEB" box="[837,1004,151,177]" pageId="3" pageNumber="156" type="generic">two specimens</specimenCount>
at Px 
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one wing each. IR1: at Px 6 (25%), 7 (58,33%) or 8 (16,6%) in Fw; at Px 6 (75%) or 7 (25%) in Hw.
</paragraph>
<paragraph id="8BDC36A1FFF6FFC63586FF20FBD82F47" blockId="3.[151,1436,151,501]" pageId="3" pageNumber="156">Measurements (n = 6): Total length 24.9–27.32 [mean 30.1; SD 0.45]; abdomen length 24.9–27.32 [mean 30.1 SD 0.45]; Fw 18–19 [mean 18.2; SD 0.45]; Hw 17–17.5 [mean 17.1; SD 0.53].</paragraph>
</subSubSection>
<subSubSection id="C379652AFFF6FFC03586FED9FDAC2961" lastPageId="5" lastPageNumber="158" pageId="3" pageNumber="156" type="diagnosis">
<paragraph id="8BDC36A1FFF6FFC63586FED9FB692FAF" blockId="3.[151,1436,151,501]" pageId="3" pageNumber="156">
<emphasis id="B917EAB3FFF6FFC63586FED9FE102F1A" bold="true" box="[199,454,294,320]" pageId="3" pageNumber="156">Differential diagnosis</emphasis>
. In the males of 
<taxonomicName id="4C634D22FFF6FFC637C3FED8FCD52F1B" authorityName="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen" authorityYear="2022" box="[642,771,295,321]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="156" phylum="Arthropoda" rank="species" species="lencionii">
<emphasis id="B917EAB3FFF6FFC637C3FED8FCD52F1B" box="[642,771,295,321]" italics="true" pageId="3" pageNumber="156">E. lencionii</emphasis>
</taxonomicName>
, the epiproct (
<figureCitation id="13582A24FFF6FFC636E4FED8FC252F1B" box="[933,1011,295,321]" captionStart="FIGURE 1" captionStartId="3.[152,255,1938,1962]" captionTargetBox="[188,1398,537,1898]" captionTargetId="figure-206@3.[176,1410,522,1911]" captionTargetPageId="3" captionText="FIGURE 1. Epipleoneura lencionii sp. nov. holotype, male, Minas Gerais State, Brazil. (a) caudal appendages, posterior view; (b) caudal appendages, lateral view; (c) genital ligula, ectal view; (d) genital ligula, lateral view. Epipleoneura venezuelensis, male, Parque Nacional Predelta, Entre Ríos Province, Argentina: (e) caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7413434" httpUri="https://zenodo.org/record/7413434/files/figure.png" pageId="3" pageNumber="156">Fig. 1a</figureCitation>
) posses two rounded apical lobes and two lateral lobes directed outwards, approximately parallel with respect to the transversal axis of the epiproct and approximately in the same plane as the surface of the epiproct. In the female, the posterior lobe of the prothorax (
<figureCitation id="13582A24FFF6FFC635DEFE6CFF322FF7" box="[159,228,403,429]" captionStart="FIGURE 2" captionStartId="4.[152,255,1768,1792]" captionTargetBox="[165,1424,182,1737]" captionTargetId="figure-18@4.[154,1432,181,1743]" captionTargetPageId="4" captionText="FIGURE 2. Epipleoneura lencionii sp. nov. female, Minas Gerais State, Brazil. (a) prothorax posterior lobe, dorsal view, photograph (left), and drawing (right); (b) prothorax posterior lobe, postero-lateral view, photograph, and drawing; (c) prothorax posterior lobe, lateral view, photograph, and drawing. PP: posterior process." figureDoi="http://doi.org/10.5281/zenodo.7413436" httpUri="https://zenodo.org/record/7413436/files/figure.png" pageId="3" pageNumber="156">Fig. 2</figureCitation>
) posses an erected medial lobe, with its anterior margin with a shallow concavity, and its posterior margin straight, not surpassing the level of the lateral lobes in dorsal view (
<figureCitation id="13582A24FFF6FFC636CCFE48FC042F8B" box="[909,978,439,465]" captionStart="FIGURE 2" captionStartId="4.[152,255,1768,1792]" captionTargetBox="[165,1424,182,1737]" captionTargetId="figure-18@4.[154,1432,181,1743]" captionTargetPageId="4" captionText="FIGURE 2. Epipleoneura lencionii sp. nov. female, Minas Gerais State, Brazil. (a) prothorax posterior lobe, dorsal view, photograph (left), and drawing (right); (b) prothorax posterior lobe, postero-lateral view, photograph, and drawing; (c) prothorax posterior lobe, lateral view, photograph, and drawing. PP: posterior process." figureDoi="http://doi.org/10.5281/zenodo.7413436" httpUri="https://zenodo.org/record/7413436/files/figure.png" pageId="3" pageNumber="156">Fig. 2</figureCitation>
), additionally, there is a rounded process with a basal position on the posterior side of the posterior lobe (posterior process in 
<figureCitation id="13582A24FFF6FFC63104FE24FB652FAF" box="[1093,1203,475,501]" captionStart="FIGURE 2" captionStartId="4.[152,255,1768,1792]" captionTargetBox="[165,1424,182,1737]" captionTargetId="figure-18@4.[154,1432,181,1743]" captionTargetPageId="4" captionText="FIGURE 2. Epipleoneura lencionii sp. nov. female, Minas Gerais State, Brazil. (a) prothorax posterior lobe, dorsal view, photograph (left), and drawing (right); (b) prothorax posterior lobe, postero-lateral view, photograph, and drawing; (c) prothorax posterior lobe, lateral view, photograph, and drawing. PP: posterior process." figureDoi="http://doi.org/10.5281/zenodo.7413436" httpUri="https://zenodo.org/record/7413436/files/figure.png" pageId="3" pageNumber="156">Fig. 2a–b</figureCitation>
).
</paragraph>
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<paragraph id="8BDC36A1FFF6FFC635D9F86DFB6429A8" blockId="3.[151,1436,1938,2034]" pageId="3" pageNumber="156">
<emphasis id="B917EAB3FFF6FFC635D9F86DFECE29F0" bold="true" box="[152,280,1938,1962]" pageId="3" pageNumber="156">FIGURE 1.</emphasis>
<taxonomicName id="4C634D22FFF6FFC6345CF86DFDD029F0" authority="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen, 2022" authorityName="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen" authorityYear="2022" box="[285,518,1938,1962]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="156" phylum="Arthropoda" rank="species" species="lencionii" status="sp. nov.">
<emphasis id="B917EAB3FFF6FFC6345CF86DFDD029F0" box="[285,518,1938,1962]" italics="true" pageId="3" pageNumber="156">Epipleoneura lencionii</emphasis>
</taxonomicName>
<emphasis id="B917EAB3FFF6FFC6374AF86DFD8D29F0" bold="true" box="[523,603,1938,1962]" pageId="3" pageNumber="156">
<taxonomicNameLabel id="A22457C8FFF6FFC6374AF86DFD8D29F0" box="[523,603,1938,1962]" pageId="3" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype, male, Minas Gerais State, Brazil. (a) caudal appendages, posterior view; (b) caudal appendages, lateral view; (c) genital ligula, ectal view; (d) genital ligula, lateral view. 
<taxonomicName id="4C634D22FFF6FFC6313AF849FA402994" authorityName=": Lencioni" authorityYear="2005" box="[1147,1430,1974,1998]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="156" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B917EAB3FFF6FFC6313AF849FA402994" box="[1147,1430,1974,1998]" italics="true" pageId="3" pageNumber="156">Epipleoneura venezuelensis</emphasis>
</taxonomicName>
, male, Parque Nacional Predelta, Entre Ríos Province, Argentina: (e) caudal appendages, posterior view.
</paragraph>
</caption>
<caption id="DF1C6629FFF1FFC135D9F917FC4C2912" ID-DOI="http://doi.org/10.5281/zenodo.7413436" ID-Zenodo-Dep="7413436" httpUri="https://zenodo.org/record/7413436/files/figure.png" pageId="4" pageNumber="157" startId="4.[152,255,1768,1792]" targetBox="[165,1424,182,1737]" targetPageId="4">
<paragraph id="8BDC36A1FFF1FFC135D9F917FC4C2912" blockId="4.[151,1436,1768,1864]" pageId="4" pageNumber="157">
<emphasis id="B917EAB3FFF1FFC135D9F917FEC4295A" bold="true" box="[152,274,1768,1792]" pageId="4" pageNumber="157">FIGURE 2</emphasis>
. 
<taxonomicName id="4C634D22FFF1FFC1345FF917FDD1295A" authority="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen, 2022" authorityName="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen" authorityYear="2022" box="[286,519,1768,1792]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="lencionii" status="sp. nov.">
<emphasis id="B917EAB3FFF1FFC1345FF917FDD1295A" box="[286,519,1768,1792]" italics="true" pageId="4" pageNumber="157">Epipleoneura lencionii</emphasis>
</taxonomicName>
<emphasis id="B917EAB3FFF1FFC1374CF917FD8B295A" bold="true" box="[525,605,1768,1792]" pageId="4" pageNumber="157">
<taxonomicNameLabel id="A22457C8FFF1FFC1374CF917FD8B295A" box="[525,605,1768,1792]" pageId="4" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female, Minas Gerais State, Brazil. (a) prothorax posterior lobe, dorsal view, photograph (left), and drawing (right); (b) prothorax posterior lobe, postero-lateral view, photograph, and drawing; (c) prothorax posterior lobe, lateral view, photograph, and drawing. PP: posterior process.
</paragraph>
</caption>
<paragraph id="8BDC36A1FFF1FFC03586F889FB4C2F47" blockId="4.[151,1436,1910,2009]" lastBlockId="5.[151,1437,150,1185]" lastPageId="5" lastPageNumber="158" pageId="4" pageNumber="157">
The male cercus and epiproct of 
<taxonomicName id="4C634D22FFF1FFC13705F888FD1E29CA" authorityName="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen" authorityYear="2022" box="[580,712,1910,1936]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="lencionii">
<emphasis id="B917EAB3FFF1FFC13705F888FD1E29CA" box="[580,712,1910,1936]" italics="true" pageId="4" pageNumber="157">E. lencionii</emphasis>
</taxonomicName>
resemble those of 
<taxonomicName id="4C634D22FFF1FFC136E4F888FB8829CA" authorityName=": Lencioni" authorityYear="2005" box="[933,1118,1910,1936]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B917EAB3FFF1FFC136E4F888FB8829CA" box="[933,1118,1910,1936]" italics="true" pageId="4" pageNumber="157">E. venezuelensis</emphasis>
</taxonomicName>
, its sibling species. In both taxa, the epiproct is very similar; the rounded apical lobes and the lateral lobes (
<figureCitation id="13582A24FFF1FFC1314AF865FB9D29EE" box="[1035,1099,1946,1972]" captionStart="FIGURE 1" captionStartId="3.[152,255,1938,1962]" captionTargetBox="[188,1398,537,1898]" captionTargetId="figure-206@3.[176,1410,522,1911]" captionTargetPageId="3" captionText="FIGURE 1. Epipleoneura lencionii sp. nov. holotype, male, Minas Gerais State, Brazil. (a) caudal appendages, posterior view; (b) caudal appendages, lateral view; (c) genital ligula, ectal view; (d) genital ligula, lateral view. Epipleoneura venezuelensis, male, Parque Nacional Predelta, Entre Ríos Province, Argentina: (e) caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7413434" httpUri="https://zenodo.org/record/7413434/files/figure.png" pageId="4" pageNumber="157">Fig. 1</figureCitation>
) separate them from other described species of 
<taxonomicName id="4C634D22FFF1FFC13422F841FE2D2982" authorityName="Williamson" authorityYear="1915" box="[355,507,1982,2008]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAB3FFF1FFC13422F841FE2D2982" box="[355,507,1982,2008]" italics="true" pageId="4" pageNumber="157">Epipleoneura</emphasis>
</taxonomicName>
. However, in 
<taxonomicName id="4C634D22FFF1FFC137D9F840FC982982" authorityName=": Lencioni" authorityYear="2005" box="[664,846,1982,2008]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B917EAB3FFF1FFC137D9F840FC982982" box="[664,846,1982,2008]" italics="true" pageId="4" pageNumber="157">E. venezuelensis</emphasis>
</taxonomicName>
(
<figureCitation id="13582A24FFF1FFC1361CF841FC7D2982" box="[861,939,1982,2008]" captionStart="FIGURE 1" captionStartId="3.[152,255,1938,1962]" captionTargetBox="[188,1398,537,1898]" captionTargetId="figure-206@3.[176,1410,522,1911]" captionTargetPageId="3" captionText="FIGURE 1. Epipleoneura lencionii sp. nov. holotype, male, Minas Gerais State, Brazil. (a) caudal appendages, posterior view; (b) caudal appendages, lateral view; (c) genital ligula, ectal view; (d) genital ligula, lateral view. Epipleoneura venezuelensis, male, Parque Nacional Predelta, Entre Ríos Province, Argentina: (e) caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7413434" httpUri="https://zenodo.org/record/7413434/files/figure.png" pageId="4" pageNumber="157">Fig. 1e</figureCitation>
) the lateral lobes of the epiproct are directed latero-distally at an angle of 45º or more with respect to the transversal axis of the epiproct, lying below the plane corresponding to the surface of the epiproct, bent down and nearly perpendicular to it in some specimens (
<figureCitation id="13582A24FFF0FFC03005FF44FA442E8F" box="[1348,1426,187,213]" captionStart="FIGURE 1" captionStartId="3.[152,255,1938,1962]" captionTargetBox="[188,1398,537,1898]" captionTargetId="figure-206@3.[176,1410,522,1911]" captionTargetPageId="3" captionText="FIGURE 1. Epipleoneura lencionii sp. nov. holotype, male, Minas Gerais State, Brazil. (a) caudal appendages, posterior view; (b) caudal appendages, lateral view; (c) genital ligula, ectal view; (d) genital ligula, lateral view. Epipleoneura venezuelensis, male, Parque Nacional Predelta, Entre Ríos Province, Argentina: (e) caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7413434" httpUri="https://zenodo.org/record/7413434/files/figure.png" pageId="5" pageNumber="158">Fig. 1e</figureCitation>
) while in 
<taxonomicName id="4C634D22FFF0FFC035BAFF20FEAB2EA3" authorityName="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen" authorityYear="2022" box="[251,381,223,249]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="lencionii">
<emphasis id="B917EAB3FFF0FFC035BAFF20FEAB2EA3" box="[251,381,223,249]" italics="true" pageId="5" pageNumber="158">E. lencionii</emphasis>
</taxonomicName>
, as aforementioned, the lateral lobes are approximately parallel to the transversal axis and lie approximately in the same plane (
<figureCitation id="13582A24FFF0FFC03753FEFCFDB42F47" box="[530,610,259,285]" captionStart="FIGURE 1" captionStartId="3.[152,255,1938,1962]" captionTargetBox="[188,1398,537,1898]" captionTargetId="figure-206@3.[176,1410,522,1911]" captionTargetPageId="3" captionText="FIGURE 1. Epipleoneura lencionii sp. nov. holotype, male, Minas Gerais State, Brazil. (a) caudal appendages, posterior view; (b) caudal appendages, lateral view; (c) genital ligula, ectal view; (d) genital ligula, lateral view. Epipleoneura venezuelensis, male, Parque Nacional Predelta, Entre Ríos Province, Argentina: (e) caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7413434" httpUri="https://zenodo.org/record/7413434/files/figure.png" pageId="5" pageNumber="158">Fig. 1a</figureCitation>
). The genital ligula is very similar in both species.
</paragraph>
<paragraph id="8BDC36A1FFF0FFC03586FED8FE2A2CF3" blockId="5.[151,1437,150,1185]" pageId="5" pageNumber="158">
In the females, the posterior lobe of the prothorax differs between the two species. In 
<taxonomicName id="4C634D22FFF0FFC031C2FED8FAEF2F1B" authorityName=": Lencioni" authorityYear="2005" box="[1155,1337,295,321]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B917EAB3FFF0FFC031C2FED8FAEF2F1B" box="[1155,1337,295,321]" italics="true" pageId="5" pageNumber="158">E. venezuelensis</emphasis>
</taxonomicName>
, the medial lobe forms a deep scoop-like process, with its posterior margin rounded and extending posteriorly, exceeding the level of the lateral lobes in dorsal view (
<figureCitation id="13582A24FFF0FFC037C1FE90FD072FD3" box="[640,721,367,393]" captionStart="FIGURE 3" captionStartId="6.[152,255,1926,1950]" captionTargetBox="[214,1370,189,1888]" captionTargetId="figure-18@6.[213,1374,181,1901]" captionTargetPageId="6" captionText="FIGURE 3. Epipleoneura venezuelensis female, Parque Nacional Predelta, Entre Ríos Province, Argentina. (a) prothorax posterior lobe, dorsal view, photograph, and drawing; (b) prothorax posterior lobe, postero-lateral view, photograph, and drawing; (c) prothorax posterior lobe, lateral view, photograph, and drawing. PP: posterior process." figureDoi="http://doi.org/10.5281/zenodo.7413438" httpUri="https://zenodo.org/record/7413438/files/figure.png" pageId="5" pageNumber="158">Fig. 3a</figureCitation>
), while in 
<taxonomicName id="4C634D22FFF0FFC03606FE90FC112FD3" authorityName="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen" authorityYear="2022" box="[839,967,367,393]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="lencionii">
<emphasis id="B917EAB3FFF0FFC03606FE90FC112FD3" box="[839,967,367,393]" italics="true" pageId="5" pageNumber="158">E. lencionii</emphasis>
</taxonomicName>
there is a shallow concavity in the medial lobe, and its posterior margin is straight and does not surpass the level of the lateral lobes (
<figureCitation id="13582A24FFF0FFC031C8FE6CFB0F2FF7" box="[1161,1241,403,429]" captionStart="FIGURE 1" captionStartId="3.[152,255,1938,1962]" captionTargetBox="[188,1398,537,1898]" captionTargetId="figure-206@3.[176,1410,522,1911]" captionTargetPageId="3" captionText="FIGURE 1. Epipleoneura lencionii sp. nov. holotype, male, Minas Gerais State, Brazil. (a) caudal appendages, posterior view; (b) caudal appendages, lateral view; (c) genital ligula, ectal view; (d) genital ligula, lateral view. Epipleoneura venezuelensis, male, Parque Nacional Predelta, Entre Ríos Province, Argentina: (e) caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7413434" httpUri="https://zenodo.org/record/7413434/files/figure.png" pageId="5" pageNumber="158">Fig. 1a</figureCitation>
). Additionally, in 
<taxonomicName id="4C634D22FFF0FFC035D6FE48FECE2F8B" authorityName="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen" authorityYear="2022" box="[151,280,439,465]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="lencionii">
<emphasis id="B917EAB3FFF0FFC035D6FE48FECE2F8B" box="[151,280,439,465]" italics="true" pageId="5" pageNumber="158">E. lencionii</emphasis>
</taxonomicName>
there is a rounded posterior process on the medial lobe (
<figureCitation id="13582A24FFF0FFC036D3FE48FBDF2F8B" box="[914,1033,439,465]" captionStart="FIGURE 2" captionStartId="4.[152,255,1768,1792]" captionTargetBox="[165,1424,182,1737]" captionTargetId="figure-18@4.[154,1432,181,1743]" captionTargetPageId="4" captionText="FIGURE 2. Epipleoneura lencionii sp. nov. female, Minas Gerais State, Brazil. (a) prothorax posterior lobe, dorsal view, photograph (left), and drawing (right); (b) prothorax posterior lobe, postero-lateral view, photograph, and drawing; (c) prothorax posterior lobe, lateral view, photograph, and drawing. PP: posterior process." figureDoi="http://doi.org/10.5281/zenodo.7413436" httpUri="https://zenodo.org/record/7413436/files/figure.png" pageId="5" pageNumber="158">Figs. 2a–b</figureCitation>
) that does not extend dorsally. In 
<taxonomicName id="4C634D22FFF0FFC030C5FE48FEF92FAF" authorityName=": Lencioni" authorityYear="2005" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B917EAB3FFF0FFC030C5FE48FEF92FAF" italics="true" pageId="5" pageNumber="158">E. venezuelensis</emphasis>
</taxonomicName>
this process is elongated dorso-ventrally, contacting the dorsal margin of the medial lobe (
<figureCitation id="13582A24FFF0FFC03056FE24FA462FAF" box="[1303,1424,475,501]" captionStart="FIGURE 3" captionStartId="6.[152,255,1926,1950]" captionTargetBox="[214,1370,189,1888]" captionTargetId="figure-18@6.[213,1374,181,1901]" captionTargetPageId="6" captionText="FIGURE 3. Epipleoneura venezuelensis female, Parque Nacional Predelta, Entre Ríos Province, Argentina. (a) prothorax posterior lobe, dorsal view, photograph, and drawing; (b) prothorax posterior lobe, postero-lateral view, photograph, and drawing; (c) prothorax posterior lobe, lateral view, photograph, and drawing. PP: posterior process." figureDoi="http://doi.org/10.5281/zenodo.7413438" httpUri="https://zenodo.org/record/7413438/files/figure.png" pageId="5" pageNumber="158">Figs. 3b–c</figureCitation>
). In some specimens of 
<taxonomicName id="4C634D22FFF0FFC034D9FE00FD872C43" authorityName=": Lencioni" authorityYear="2005" box="[408,593,511,537]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B917EAB3FFF0FFC034D9FE00FD872C43" box="[408,593,511,537]" italics="true" pageId="5" pageNumber="158">E. venezuelensis</emphasis>
</taxonomicName>
, this structure is rounded (
<figureCitation id="13582A24FFF0FFC036C4FE00FC0F2C43" box="[901,985,511,537]" captionStart="FIGURE 3" captionStartId="6.[152,255,1926,1950]" captionTargetBox="[214,1370,189,1888]" captionTargetId="figure-18@6.[213,1374,181,1901]" captionTargetPageId="6" captionText="FIGURE 3. Epipleoneura venezuelensis female, Parque Nacional Predelta, Entre Ríos Province, Argentina. (a) prothorax posterior lobe, dorsal view, photograph, and drawing; (b) prothorax posterior lobe, postero-lateral view, photograph, and drawing; (c) prothorax posterior lobe, lateral view, photograph, and drawing. PP: posterior process." figureDoi="http://doi.org/10.5281/zenodo.7413438" httpUri="https://zenodo.org/record/7413438/files/figure.png" pageId="5" pageNumber="158">Fig. 3b</figureCitation>
), while in others its posterior margin is more or less straight. We studied 
<specimenCount id="9D65FD28FFF0FFC03749FDDCFD542C67" box="[520,642,547,573]" pageId="5" pageNumber="158" type="female">23 females</specimenCount>
of 
<taxonomicName id="4C634D22FFF0FFC037E7FDDCFC8A2C67" authorityName=": Lencioni" authorityYear="2005" box="[678,860,547,573]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B917EAB3FFF0FFC037E7FDDCFC8A2C67" box="[678,860,547,573]" italics="true" pageId="5" pageNumber="158">E. venezuelensis</emphasis>
</taxonomicName>
from eight populations, and the posterior margin of the pronotum of these specimens is like that described for the female specimens of 
<taxonomicName id="4C634D22FFF0FFC03171FDB8FB302C3B" authorityName=": Lencioni" authorityYear="2005" box="[1072,1254,583,609]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B917EAB3FFF0FFC03171FDB8FB302C3B" box="[1072,1254,583,609]" italics="true" pageId="5" pageNumber="158">E. venezuelensis</emphasis>
</taxonomicName>
from 
<collectingCountry id="F3747631FFF0FFC03068FDB8FA4A2C3B" box="[1321,1436,583,609]" name="Venezuela" pageId="5" pageNumber="158">Venezuela</collectingCountry>
(
<bibRefCitation id="EFF24B50FFF0FFC035DEFD94FEFD2CDF" author="Santos, N. D. dos" box="[159,299,618,645]" pageId="5" pageNumber="158" pagination="187 - 190" refId="ref7495" refString="Santos, N. D. dos (1957) Descricao do alotipo femea de &quot; Epipleoneura venezuelensis &quot; Racenis, 1955 e notas sobre o macho (Protoneuridae: Odonata). Revista Brasileira de Biologia, 17 (2), 187 - 190." type="journal article" year="1957">Santos 1957</bibRefCitation>
) (
<figureCitation id="13582A24FFF0FFC03403FD94FE502CDF" box="[322,390,619,645]" captionStart="FIGURE 4" captionStartId="7.[152,255,1191,1215]" captionTargetBox="[151,1410,181,1167]" captionTargetId="figure-17@7.[151,1436,181,1167]" captionTargetPageId="7" captionText="FIGURE 4. Epipleoneura venezuelensis female, Venezuela. (a) prothorax posterior lobe, dorsal view. Picture provided by Dr. R.W. Garrison." figureDoi="http://doi.org/10.5281/zenodo.7413440" httpUri="https://zenodo.org/record/7413440/files/figure.png" pageId="5" pageNumber="158">Fig. 4</figureCitation>
), close to the 
<typeStatus id="54D88803FFF0FFC03760FD94FD872CDF" box="[545,593,619,645]" pageId="5" pageNumber="158">type</typeStatus>
locality, the drawings presented in 
<bibRefCitation id="EFF24B50FFF0FFC0369DFD94FB5F2CDF" author="Pessacq, P." box="[988,1161,619,645]" pageId="5" pageNumber="158" pagination="201 - 234" refId="ref7268" refString="Pessacq, P. (2014) Synopsis of Epipleoneura (Zygoptera, Coenagrionidae, &quot; Protoneuridae &quot;), with emphasis on its Brazilian species. Zootaxa, 3872 (3), 201 - 234." type="journal article" year="2014">Pessacq (2014)</bibRefCitation>
, and the photograph of a female from 
<collectingCountry id="F3747631FFF0FFC03466FD70FE4C2CF3" box="[295,410,655,681]" name="Venezuela" pageId="5" pageNumber="158">Venezuela</collectingCountry>
(
<figureCitation id="13582A24FFF0FFC034EBFD70FE392CF3" box="[426,495,655,681]" captionStart="FIGURE 4" captionStartId="7.[152,255,1191,1215]" captionTargetBox="[151,1410,181,1167]" captionTargetId="figure-17@7.[151,1436,181,1167]" captionTargetPageId="7" captionText="FIGURE 4. Epipleoneura venezuelensis female, Venezuela. (a) prothorax posterior lobe, dorsal view. Picture provided by Dr. R.W. Garrison." figureDoi="http://doi.org/10.5281/zenodo.7413440" httpUri="https://zenodo.org/record/7413440/files/figure.png" pageId="5" pageNumber="158">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BDC36A1FFF0FFC03586FD4CFE692D07" blockId="5.[151,1437,150,1185]" pageId="5" pageNumber="158">
Studying the photographs of 
<taxonomicName id="4C634D22FFF0FFC03752FD4CFD7D2C97" authorityName="Williamson" authorityYear="1915" box="[531,683,691,717]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAB3FFF0FFC03752FD4CFD7D2C97" box="[531,683,691,717]" italics="true" pageId="5" pageNumber="158">Epipleoneura</emphasis>
</taxonomicName>
females provided by R.W. Garrison, we observed specimens with a prothorax morphology very similar or the same as 
<emphasis id="B917EAB3FFF0FFC037A7FD28FCB82CAB" box="[742,878,727,753]" italics="true" pageId="5" pageNumber="158">
<taxonomicName id="4C634D22FFF0FFC037A7FD28FCBC2CAB" authorityName="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen" authorityYear="2022" box="[742,874,727,753]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="lencionii">E. lencionii</taxonomicName>
,
</emphasis>
with a shallow concavity in the prothorax medial lobe (material from 
<collectingCountry id="F3747631FFF0FFC03436FD04FE1F2D4F" box="[375,457,763,789]" name="Bolivia" pageId="5" pageNumber="158">Bolivia</collectingCountry>
and 
<collectingCountry id="F3747631FFF0FFC03741FD04FD922D4F" box="[512,580,763,789]" name="Brazil" pageId="5" pageNumber="158">Brazil</collectingCountry>
in Amazonas state) and specimens with a morphology closer to 
<taxonomicName id="4C634D22FFF0FFC03055FD04FF192D62" authorityName=": Lencioni" authorityYear="2005" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B917EAB3FFF0FFC03055FD04FF192D62" italics="true" pageId="5" pageNumber="158">E. venezuelensis</emphasis>
</taxonomicName>
(material from 
<collectingCountry id="F3747631FFF0FFC034C0FCE0FE362D63" box="[385,480,799,825]" name="Ecuador" pageId="5" pageNumber="158">Ecuador</collectingCountry>
, 
<collectingCountry id="F3747631FFF0FFC034ADFCE0FDE62D63" box="[492,560,799,825]" name="Brazil" pageId="5" pageNumber="158">Brazil</collectingCountry>
[
<collectingRegion id="49A7F843FFF0FFC0377FFCE0FDA32D63" box="[574,629,799,825]" country="Brazil" name="Para" pageId="5" pageNumber="158">Pará</collectingRegion>
, and 
<collectingRegion id="49A7F843FFF0FFC037F0FCE0FCC82D63" box="[689,798,799,825]" country="Brazil" name="Rondonia" pageId="5" pageNumber="158">Rondônia</collectingRegion>
states] and 
<collectingCountry id="F3747631FFF0FFC036E4FCE0FBC42D63" box="[933,1042,799,825]" name="Paraguay" pageId="5" pageNumber="158">Paraguay</collectingCountry>
), with a deep scoop-like process in the prothorax medial lobe.
</paragraph>
<paragraph id="8BDC36A1FFF0FFC03586FC98FD422A6F" blockId="5.[151,1437,150,1185]" pageId="5" pageNumber="158">
Some male specimens of 
<taxonomicName id="4C634D22FFF0FFC034A9FC98FDBF2DDB" authorityName="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen" authorityYear="2022" box="[488,617,871,897]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="lencionii">
<emphasis id="B917EAB3FFF0FFC034A9FC98FDBF2DDB" box="[488,617,871,897]" italics="true" pageId="5" pageNumber="158">E. lencionii</emphasis>
</taxonomicName>
collected in Assis County show similar morphology to 
<taxonomicName id="4C634D22FFF0FFC0319CFC98FA432DDB" authorityName=": Lencioni" authorityYear="2005" box="[1245,1429,871,897]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B917EAB3FFF0FFC0319CFC98FA432DDB" box="[1245,1429,871,897]" italics="true" pageId="5" pageNumber="158">E. venezuelensis</emphasis>
</taxonomicName>
, that is, the lateral lobes of the epiproct are directed latero-distally, but with an angle less than 45º in relation to the transverse axis of the epiproct. However, the associated female possesses the typical morphology of the species. The morphological difference observed is more evident in females than in males, in the case of intermediate male morphology (i.e., lateral lobes of the epiproct not parallel to the transverse axis but not 45º to it), the male identification depends on the association with females.
</paragraph>
<paragraph id="8BDC36A1FFF0FFC03586FBC1FC792AFB" blockId="5.[151,1437,150,1185]" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC03586FBC1FDB22A02" bold="true" box="[199,612,1086,1113]" pageId="5" pageNumber="158">Habitat and reproductive behavior</emphasis>
. Adult 
<taxonomicName id="4C634D22FFF0FFC037F4FBC0FCE32A03" authorityName="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen" authorityYear="2022" box="[693,821,1087,1113]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="lencionii">
<emphasis id="B917EAB3FFF0FFC037F4FBC0FCE32A03" box="[693,821,1087,1113]" italics="true" pageId="5" pageNumber="158">E. lencionii</emphasis>
</taxonomicName>
individuals were collected between 11:00 AM and 2:00 PM near small streams, in areas dominated by disturbed pastures as well as areas with greater forest cover (
<tableCitation id="C6E1031AFFF0FFC03079FB9CFA5D2A27" box="[1336,1419,1123,1149]" captionStart="TABLE 1" captionStartId="5.[152,244,1232,1256]" captionTargetPageId="5" captionText="TABLE 1. Environmental variables at the collection points of E. lencionii sp. nov. HII: Habitat Integrity Index, for more details see Nessimian et al. (2017)" httpUri="http://table.plazi.org/id/DF1C6629FFF0FFC035D9FB2FFDD02B57" pageId="5" pageNumber="158" tableUuid="DF1C6629FFF0FFC035D9FB2FFDD02B57">Table 1</tableCitation>
). These streams drained a matrix of agriculture and regenerating forests.
</paragraph>
<caption id="DF1C6629FFF0FFC035D9FB2FFDD02B57" ID-Table-UUID="DF1C6629FFF0FFC035D9FB2FFDD02B57" httpUri="http://table.plazi.org/id/DF1C6629FFF0FFC035D9FB2FFDD02B57" pageId="5" pageNumber="158" startId="5.[152,244,1232,1256]" targetBox="[166,1417,1308,1525]" targetIsTable="true" targetPageId="5">
<paragraph id="8BDC36A1FFF0FFC035D9FB2FFDD02B57" blockId="5.[151,1436,1232,1293]" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC035D9FB2FFED82AB2" bold="true" box="[152,270,1232,1256]" pageId="5" pageNumber="158">TABLE 1.</emphasis>
Environmental variables at the collection points of 
<taxonomicName id="4C634D22FFF0FFC03673FB2EFC7A2AB2" authority="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen, 2022" authorityName="Pessacq &amp; Anjos-Santos &amp; Carvalho &amp; Calvão &amp; Mendoza-Penagos &amp; Juen" authorityYear="2022" box="[818,940,1232,1256]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="lencionii" status="sp. nov.">
<emphasis id="B917EAB3FFF0FFC03673FB2EFC7A2AB2" box="[818,940,1232,1256]" italics="true" pageId="5" pageNumber="158">E. lencionii</emphasis>
</taxonomicName>
<emphasis id="B917EAB3FFF0FFC036F2FB2EFBDE2AB3" bold="true" box="[947,1032,1233,1257]" pageId="5" pageNumber="158">
<taxonomicNameLabel id="A22457C8FFF0FFC036F2FB2EFBDE2AB3" box="[947,1032,1233,1257]" pageId="5" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
HII: Habitat Integrity Index, for more details see Nessimian 
<emphasis id="B917EAB3FFF0FFC034C2FB0AFE6F2B56" box="[387,441,1268,1292]" italics="true" pageId="5" pageNumber="158">et al.</emphasis>
(2017)
</paragraph>
</caption>
<paragraph id="8BDC36A1FFF0FFC035E7FAE3FA5F2BAF" pageId="5" pageNumber="158">
<table id="F963C401FFF0003A35E7FAE3FA5F2BAF" box="[166,1417,1308,1525]" gridcols="11" gridrows="4" pageId="5" pageNumber="158">
<tr id="355334E3FFF0003A35E7FAE3FA5F2B26" box="[166,1417,1308,1404]" gridrow="0" pageId="5" pageNumber="158">
<th id="76825D9FFFF0003A35E7FAE3FF222B26" box="[166,244,1308,1404]" gridcol="0" gridrow="0" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC035E7FAE3FF222B6E" bold="true" box="[166,244,1308,1332]" pageId="5" pageNumber="158">Stream</emphasis>
</th>
<th id="76825D9FFFF0003A3457FAE3FEB72B26" box="[278,353,1308,1404]" gridcol="1" gridrow="0" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC03457FAE3FEEB2B02" bold="true" box="[278,336,1308,1368]" pageId="5" pageNumber="158">Land Use</emphasis>
</th>
<th id="76825D9FFFF0003A34C4FAE3FE682B26" box="[389,446,1308,1404]" gridcol="2" gridrow="0" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC034C4FAE3FE7A2B6E" bold="true" box="[389,428,1308,1332]" pageId="5" pageNumber="158">HII</emphasis>
</th>
<th id="76825D9FFFF0003A34A2FAE3FD9D2B26" box="[483,587,1308,1404]" gridcol="3" gridrow="0" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC034A2FAE3FDD02B02" bold="true" pageId="5" pageNumber="158">Air temp. (C)</emphasis>
</th>
<th id="76825D9FFFF0003A3728FAE3FD262B26" box="[617,752,1308,1404]" gridcol="4" gridrow="0" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC03728FAE3FDAA2B02" bold="true" pageId="5" pageNumber="158">Water temp. ©</emphasis>
</th>
<th id="76825D9FFFF0003A3655FAE3FCA12B26" box="[788,887,1308,1404]" gridcol="5" gridrow="0" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC03655FAE3FCE92B26" bold="true" box="[788,887,1308,1404]" pageId="5" pageNumber="158">Relative humidity (%)</emphasis>
</th>
<th id="76825D9FFFF0003A36EBFAE3FC262B26" box="[938,1008,1308,1404]" gridcol="6" gridrow="0" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC036EBFAE3FC0D2B02" bold="true" box="[938,1004,1308,1368]" pageId="5" pageNumber="158">Depth (cm)</emphasis>
</th>
<th id="76825D9FFFF0003A3159FAE3FB8B2B26" box="[1048,1117,1308,1404]" gridcol="7" gridrow="0" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC03159FAE3FBE82B02" bold="true" box="[1048,1117,1308,1368]" pageId="5" pageNumber="158">Width (m)</emphasis>
</th>
<th id="76825D9FFFF0003A313BFAE3FB652B26" box="[1146,1203,1308,1404]" gridcol="8" gridrow="0" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC0313BFAE3FB792B6E" bold="true" box="[1146,1199,1308,1332]" pageId="5" pageNumber="158">Flow</emphasis>
</th>
<th id="76825D9FFFF0003A319AFAE3FAF92B26" box="[1243,1327,1308,1404]" gridcol="9" gridrow="0" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC0319AFAE3FB222B02" bold="true" box="[1243,1327,1308,1368]" pageId="5" pageNumber="158">Canopy %</emphasis>
</th>
<th id="76825D9FFFF0003A3011FAE3FA5F2B26" box="[1360,1417,1308,1404]" gridcol="10" gridrow="0" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC03011FAE3FAA42B6E" bold="true" box="[1360,1394,1308,1332]" pageId="5" pageNumber="158">pH</emphasis>
</th>
</tr>
<tr id="355334E3FFF0003A35E7FA73FA5F2BFF" box="[166,1417,1420,1445]" gridrow="1" pageId="5" pageNumber="158">
<th id="76825D9FFFF0003A35E7FA73FF222BFF" box="[166,244,1420,1445]" gridcol="0" gridrow="1" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC035E7FA73FF322BFE" bold="true" box="[166,228,1420,1444]" pageId="5" pageNumber="158">AC01</emphasis>
</th>
<td id="76825D9FFFF0003A3457FA73FEB72BFF" box="[278,353,1420,1445]" gridcol="1" gridrow="1" pageId="5" pageNumber="158">Pasture</td>
<td id="76825D9FFFF0003A34C4FA73FE682BFF" box="[389,446,1420,1445]" gridcol="2" gridrow="1" pageId="5" pageNumber="158">0.417</td>
<td id="76825D9FFFF0003A34A2FA73FD9D2BFF" box="[483,587,1420,1445]" gridcol="3" gridrow="1" pageId="5" pageNumber="158">28.700</td>
<td id="76825D9FFFF0003A3728FA73FD262BFF" box="[617,752,1420,1445]" gridcol="4" gridrow="1" pageId="5" pageNumber="158">28.700</td>
<td id="76825D9FFFF0003A3655FA73FCA12BFF" box="[788,887,1420,1445]" gridcol="5" gridrow="1" pageId="5" pageNumber="158">74.000</td>
<td id="76825D9FFFF0003A36EBFA73FC262BFF" box="[938,1008,1420,1445]" gridcol="6" gridrow="1" pageId="5" pageNumber="158">14.800</td>
<td id="76825D9FFFF0003A3159FA73FB8B2BFF" box="[1048,1117,1420,1445]" gridcol="7" gridrow="1" pageId="5" pageNumber="158">1.800</td>
<td id="76825D9FFFF0003A313BFA73FB652BFF" box="[1146,1203,1420,1445]" gridcol="8" gridrow="1" pageId="5" pageNumber="158">0.050</td>
<td id="76825D9FFFF0003A319AFA73FAF92BFF" box="[1243,1327,1420,1445]" gridcol="9" gridrow="1" pageId="5" pageNumber="158">85.313</td>
<td id="76825D9FFFF0003A3011FA73FA5F2BFF" box="[1360,1417,1420,1445]" gridcol="10" gridrow="1" pageId="5" pageNumber="158">6.235</td>
</tr>
<tr id="355334E3FFF0003A35E7FA4BFA5F2B97" box="[166,1417,1460,1485]" gridrow="2" pageId="5" pageNumber="158">
<th id="76825D9FFFF0003A35E7FA4BFF222B97" box="[166,244,1460,1485]" gridcol="0" gridrow="2" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC035E7FA4BFF352B96" bold="true" box="[166,227,1460,1484]" pageId="5" pageNumber="158">AC04</emphasis>
</th>
<td id="76825D9FFFF0003A3457FA4BFEB72B97" box="[278,353,1460,1485]" gridcol="1" gridrow="2" pageId="5" pageNumber="158">Forest</td>
<td id="76825D9FFFF0003A34C4FA4BFE682B97" box="[389,446,1460,1485]" gridcol="2" gridrow="2" pageId="5" pageNumber="158">0.970</td>
<td id="76825D9FFFF0003A34A2FA4BFD9D2B97" box="[483,587,1460,1485]" gridcol="3" gridrow="2" pageId="5" pageNumber="158">30.500</td>
<td id="76825D9FFFF0003A3728FA4BFD262B97" box="[617,752,1460,1485]" gridcol="4" gridrow="2" pageId="5" pageNumber="158">26.700</td>
<td id="76825D9FFFF0003A3655FA4BFCA12B97" box="[788,887,1460,1485]" gridcol="5" gridrow="2" pageId="5" pageNumber="158">81.000</td>
<td id="76825D9FFFF0003A36EBFA4BFC262B97" box="[938,1008,1460,1485]" gridcol="6" gridrow="2" pageId="5" pageNumber="158">28.800</td>
<td id="76825D9FFFF0003A3159FA4BFB8B2B97" box="[1048,1117,1460,1485]" gridcol="7" gridrow="2" pageId="5" pageNumber="158">1.800</td>
<td id="76825D9FFFF0003A313BFA4BFB652B97" box="[1146,1203,1460,1485]" gridcol="8" gridrow="2" pageId="5" pageNumber="158">0.111</td>
<td id="76825D9FFFF0003A319AFA4BFAF92B97" box="[1243,1327,1460,1485]" gridcol="9" gridrow="2" pageId="5" pageNumber="158">89.555</td>
<td id="76825D9FFFF0003A3011FA4BFA5F2B97" box="[1360,1417,1460,1485]" gridcol="10" gridrow="2" pageId="5" pageNumber="158">6.350</td>
</tr>
<tr id="355334E3FFF0003A35E7FA23FA5F2BAF" box="[166,1417,1500,1525]" gridrow="3" pageId="5" pageNumber="158">
<th id="76825D9FFFF0003A35E7FA23FF222BAF" box="[166,244,1500,1525]" gridcol="0" gridrow="3" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC035E7FA23FF352BAE" bold="true" box="[166,227,1500,1524]" pageId="5" pageNumber="158">AC09</emphasis>
</th>
<td id="76825D9FFFF0003A3457FA23FEB72BAF" box="[278,353,1500,1525]" gridcol="1" gridrow="3" pageId="5" pageNumber="158">Forest</td>
<td id="76825D9FFFF0003A34C4FA23FE682BAF" box="[389,446,1500,1525]" gridcol="2" gridrow="3" pageId="5" pageNumber="158">0.870</td>
<td id="76825D9FFFF0003A34A2FA23FD9D2BAF" box="[483,587,1500,1525]" gridcol="3" gridrow="3" pageId="5" pageNumber="158">31.300</td>
<td id="76825D9FFFF0003A3728FA23FD262BAF" box="[617,752,1500,1525]" gridcol="4" gridrow="3" pageId="5" pageNumber="158">25.050</td>
<td id="76825D9FFFF0003A3655FA23FCA12BAF" box="[788,887,1500,1525]" gridcol="5" gridrow="3" pageId="5" pageNumber="158">79.000</td>
<td id="76825D9FFFF0003A36EBFA23FC262BAF" box="[938,1008,1500,1525]" gridcol="6" gridrow="3" pageId="5" pageNumber="158">12.000</td>
<td id="76825D9FFFF0003A3159FA23FB8B2BAF" box="[1048,1117,1500,1525]" gridcol="7" gridrow="3" pageId="5" pageNumber="158">0.900</td>
<td id="76825D9FFFF0003A313BFA23FB652BAF" box="[1146,1203,1500,1525]" gridcol="8" gridrow="3" pageId="5" pageNumber="158">0.051</td>
<td id="76825D9FFFF0003A319AFA23FAF92BAF" box="[1243,1327,1500,1525]" gridcol="9" gridrow="3" pageId="5" pageNumber="158">91.987</td>
<td id="76825D9FFFF0003A3011FA23FA5F2BAF" box="[1360,1417,1500,1525]" gridcol="10" gridrow="3" pageId="5" pageNumber="158">6.250</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDC36A1FFF0FFC03586F9DBFDAC2961" blockId="5.[151,1436,1572,1961]" pageId="5" pageNumber="158">
Regarding the species’ reproductive behavior (
<figureCitation id="13582A24FFF0FFC0379CF9DBFCF22865" box="[733,804,1572,1599]" captionStart="FIGURE 7" captionStartId="9.[152,255,1991,2015]" captionTargetBox="[202,1385,1166,1967]" captionTargetId="figure-32@9.[202,1385,1166,1967]" captionTargetPageId="9" captionText="FIGURE 7. Epipleoneura lencionii, photographic sequence of male perching and male and female in tandem and oviposition." figureDoi="http://doi.org/10.5281/zenodo.7413446" httpUri="https://zenodo.org/record/7413446/files/figure.png" pageId="5" pageNumber="158">Fig. 7</figureCitation>
), males were found perched on dry branches or on the tips of leaves approximately 
<quantity id="4C9B9B44FFF0FFC034A2F9B6FDEC2839" box="[483,570,1609,1635]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="158" unit="m" value="1.0">1 meter</quantity>
above the water surface. Disputes between males for territory control were observed. Tandem and copulation occur on the branches and leaves found on the stream bank. After finishing copulation, both descend in tandem towards the stream channel in search of a suitable place for oviposition (where the current is slow). The male holds the female for approximately 4–6 seconds, while she submerges her abdomen to oviposit, after which she is released. The female remains with her abdomen submerged in the water for 10 to 15 seconds. The substrate chosen for oviposition are the fine roots of aquatic plants. These are the first observations of the reproductive behavior in 
<taxonomicName id="4C634D22FFF0FFC0349AF8DFFDA52960" authorityName="Williamson" authorityYear="1915" box="[475,627,1824,1850]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAB3FFF0FFC0349AF8DFFDA52960" box="[475,627,1824,1850]" italics="true" pageId="5" pageNumber="158">Epipleoneura</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C379652AFFF0FFC03586F8BBFA9529FD" pageId="5" pageNumber="158" type="distribution">
<paragraph id="8BDC36A1FFF0FFC03586F8BBFA9529FD" blockId="5.[151,1436,1572,1961]" pageId="5" pageNumber="158">
<emphasis id="B917EAB3FFF0FFC03586F8BBFE8F2904" bold="true" box="[199,345,1860,1886]" pageId="5" pageNumber="158">Distribution</emphasis>
. Central and west 
<collectingCountry id="F3747631FFF0FFC03769F8BBFDB92904" box="[552,623,1860,1886]" name="Brazil" pageId="5" pageNumber="158">Brazil</collectingCountry>
, in 
<collectingRegion id="49A7F843FFF0FFC037D5F8BAFD1C2905" box="[660,714,1861,1887]" country="Brazil" name="Acre" pageId="5" pageNumber="158">Acre</collectingRegion>
and 
<collectingRegion id="49A7F843FFF0FFC037BEF8BBFC422904" box="[767,916,1860,1886]" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="158">Minas Gerais</collectingRegion>
States (
<figureCitation id="13582A24FFF0FFC036ADF8BBFBE62905" box="[1004,1072,1860,1887]" captionStart="FIGURE 5" captionStartId="8.[152,255,1962,1986]" captionTargetBox="[419,1173,893,1939]" captionTargetId="figure-337@8.[415,1173,885,1939]" captionTargetPageId="8" captionText="FIGURE 5. Distribution map of Epipleoneura lencionii and E. venezuelensis. Indicated the hydrological basins:Amazon basin, Tocantins-Plata basin and Plata-Paraná-Paraguay basin." figureDoi="http://doi.org/10.5281/zenodo.7413442" httpUri="https://zenodo.org/record/7413442/files/figure.png" pageId="5" pageNumber="158">Fig. 5</figureCitation>
). All 
<typeStatus id="54D88803FFF0FFC0312CF8BAFB4B2905" box="[1133,1181,1861,1887]" pageId="5" pageNumber="158">type</typeStatus>
material was collected in 
<collectingRegion id="49A7F843FFF0FFC035F3F896FF3D29D9" box="[178,235,1897,1923]" country="Brazil" name="Acre" pageId="5" pageNumber="158">Acre</collectingRegion>
, the 
<typeStatus id="54D88803FFF0FFC0345FF896FE9829D9" box="[286,334,1897,1923]" pageId="5" pageNumber="158">type</typeStatus>
locality is in Rio Branco, at 
<geoCoordinate id="EE575066FFF0FFC037CDF895FCCC29D9" box="[652,794,1897,1925]" degrees="10" direction="south" minutes="7" orientation="latitude" pageId="5" pageNumber="158" precision="1" seconds="38.5" value="-10.127361">10º7′38.5″ S</geoCoordinate>
; 
<geoCoordinate id="EE575066FFF0FFC03667F895FC6B29D9" box="[806,957,1897,1925]" degrees="67" direction="west" minutes="38" orientation="longitude" pageId="5" pageNumber="158" precision="1" seconds="6.7" value="-67.63519">67º38′6.7″ W</geoCoordinate>
. The additional material studied, lost in the MNRJ fire, was collected close to Nova União City, 
<collectingRegion id="49A7F843FFF0FFC037A4F873FC6C29FD" box="[741,954,1932,1959]" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="158">Minas Gerais State</collectingRegion>
(about 
<geoCoordinate id="EE575066FFF0FFC03151F871FB4029FD" box="[1040,1174,1933,1961]" degrees="19" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="158" precision="15" seconds="17" value="-19.688055">19º41′17″ S</geoCoordinate>
, 
<geoCoordinate id="EE575066FFF0FFC031E2F871FAE129FD" box="[1187,1335,1933,1961]" degrees="43" direction="west" minutes="35" orientation="longitude" pageId="5" pageNumber="158" precision="15" seconds="00" value="-43.583332">43º35′00″ W</geoCoordinate>
).
</paragraph>
</subSubSection>
</treatment>
</document>